List of Chthoniidae species
Encyclopedia
This is a list of the described species of the pseudoscorpion
Pseudoscorpion
A pseudoscorpion, , is an arachnid belonging to the order Pseudoscorpionida, also known as Pseudoscorpiones or Chelonethida....

 family Chthoniidae
Chthoniidae
Chthoniidae is a family of pseudoscorpions within the superfamily Chthonioidea. The family contains more than 600 species in about 30 genera. Three fossil species are known from Baltic and Dominican amber.-Genera:...

. The data is taken from Joel Hallan's Biology Catalog.
  • Afrochthonius Beier, 1930
  • Afrochthonius brincki Beier, 1955Lesotho
    Lesotho
    Lesotho , officially the Kingdom of Lesotho, is a landlocked country and enclave, surrounded by the Republic of South Africa. It is just over in size with a population of approximately 2,067,000. Its capital and largest city is Maseru. Lesotho is a member of the Commonwealth of Nations. The name...

  • Afrochthonius ceylonicus Beier, 1973Sri Lanka
    Sri Lanka
    Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka is a country off the southern coast of the Indian subcontinent. Known until 1972 as Ceylon , Sri Lanka is an island surrounded by the Indian Ocean, the Gulf of Mannar and the Palk Strait, and lies in the vicinity of India and the...

  • Afrochthonius godfreyi (Ellingsen, 1912)Southern Africa
    Southern Africa
    Southern Africa is the southernmost region of the African continent, variably defined by geography or geopolitics. Within the region are numerous territories, including the Republic of South Africa ; nowadays, the simpler term South Africa is generally reserved for the country in English.-UN...

  • Afrochthonius inaequalis Beier, 1958 — Southern Africa
  • Afrochthonius natalensis Beier, 1931 — Southern Africa
  • Afrochthonius reductus Beier, 1973 — Sri Lanka
  • Afrochthonius similis Beier, 1930Namibia
    Namibia
    Namibia, officially the Republic of Namibia , is a country in southern Africa whose western border is the Atlantic Ocean. It shares land borders with Angola and Zambia to the north, Botswana to the east and South Africa to the south and east. It gained independence from South Africa on 21 March...


  • Allochthonius J.C. Chamberlin, 1929
  • Allochthonius borealis Sato, 1984Japan
    Japan
    Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

  • Allochthonius buanensis W.K.Lee, 1982Korea
    Korea
    Korea ) is an East Asian geographic region that is currently divided into two separate sovereign states — North Korea and South Korea. Located on the Korean Peninsula, Korea is bordered by the People's Republic of China to the northwest, Russia to the northeast, and is separated from Japan to the...

  • Allochthonius kinkaiensis Sakayori, 2002 — Japan
  • Allochthonius montanus Sakayori, 2000 — Honshu, Tochigi, Mount Kohshin, Japan
  • Allochthonius opticus (Ellingsen, 1907) — Japan
  • Allochthonius opticus opticus (Ellingsen, 1907) — Japan
  • Allochthonius opticus coreanus Morikawa, 1970 — Korea
  • Allochthonius opticus troglophilus Morikawa, 1956 — Japan
    • Allochthonius shintoisticus J.C. Chamberlin, 1929 — Japan
    • Allochthonius tamurai Sakayori, 1999 — Japan
    • Allochthonius biocularis (Morikawa, 1956) — Japan
    • Allochthonius ishikawai (Morikawa, 1954) — Japan
  • Allochthonius ishikawai ishikawai (Morikawa, 1954) — Japan
  • Allochthonius ishikawai deciclavatus (Morikawa, 1956) — Japan
  • Allochthonius ishikawai kyushuensis (Morikawa, 1960) — Japan
  • Allochthonius ishikawai shiragatakiensis (Morikawa, 1954) — Japan
  • Allochthonius ishikawai uenoi (Morikawa, 1956) — Japan
    • Allochthonius uyamadensis (Morikawa, 1954) — Japan

  • Aphrastochthonius J.C. Chamberlin, 1962
  • Aphrastochthonius alteriae Muchmore, 1977Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

  • Aphrastochthonius cubanus Dumitresco & Orghidan, 1977Cuba
    Cuba
    The Republic of Cuba is an island nation in the Caribbean. The nation of Cuba consists of the main island of Cuba, the Isla de la Juventud, and several archipelagos. Havana is the largest city in Cuba and the country's capital. Santiago de Cuba is the second largest city...

  • Aphrastochthonius grubbsi Muchmore, 1984California
    California
    California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

  • Aphrastochthonius major Muchmore, 1973 — Mexico
  • Aphrastochthonius pachysetus Muchmore, 1976New Mexico
    New Mexico
    New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...

  • Aphrastochthonius palmitensis Muchmore, 1986 — Mexico
  • Aphrastochthonius parvus Muchmore, 1972 — Mexico
  • Aphrastochthonius patei Muchmore, 1982 — Mexico
  • Aphrastochthonius pecki Muchmore, 1968Alabama
    Alabama
    Alabama is a state located in the southeastern region of the United States. It is bordered by Tennessee to the north, Georgia to the east, Florida and the Gulf of Mexico to the south, and Mississippi to the west. Alabama ranks 30th in total land area and ranks second in the size of its inland...

  • Aphrastochthonius russelli Muchmore, 1972 — Mexico
  • Aphrastochthonius similis Muchmore, 1984 — California
  • Aphrastochthonius tenax J.C. Chamberlin, 1962
  • Aphrastochthonius verapazanus Muchmore, 1972Guatemala
    Guatemala
    Guatemala is a country in Central America bordered by Mexico to the north and west, the Pacific Ocean to the southwest, Belize to the northeast, the Caribbean to the east, and Honduras and El Salvador to the southeast...


  • Apochthonius J.C. Chamberlin, 1929
  • Apochthonius coecus (Packard, 1884)Virginia
    Virginia
    The Commonwealth of Virginia , is a U.S. state on the Atlantic Coast of the Southern United States. Virginia is nicknamed the "Old Dominion" and sometimes the "Mother of Presidents" after the eight U.S. presidents born there...

  • Apochthonius colecampi Muchmore, 1967Missouri
    Missouri
    Missouri is a US state located in the Midwestern United States, bordered by Iowa, Illinois, Kentucky, Tennessee, Arkansas, Oklahoma, Kansas and Nebraska. With a 2010 population of 5,988,927, Missouri is the 18th most populous state in the nation and the fifth most populous in the Midwest. It...

  • Apochthonius diabolus Muchmore, 1967Arkansas
    Arkansas
    Arkansas is a state located in the southern region of the United States. Its name is an Algonquian name of the Quapaw Indians. Arkansas shares borders with six states , and its eastern border is largely defined by the Mississippi River...

  • Apochthonius forbesi Benedict, 1979Oregon
    Oregon
    Oregon is a state in the Pacific Northwest region of the United States. It is located on the Pacific coast, with Washington to the north, California to the south, Nevada on the southeast and Idaho to the east. The Columbia and Snake rivers delineate much of Oregon's northern and eastern...

  • Apochthonius grubbsi Muchmore, 1980 — California
  • Apochthonius hobbsi Muchmore, 1994Ohio
    Ohio
    Ohio is a Midwestern state in the United States. The 34th largest state by area in the U.S.,it is the 7th‑most populous with over 11.5 million residents, containing several major American cities and seven metropolitan areas with populations of 500,000 or more.The state's capital is Columbus...

  • Apochthonius holsingeri Muchmore, 1967Virginia
    Virginia
    The Commonwealth of Virginia , is a U.S. state on the Atlantic Coast of the Southern United States. Virginia is nicknamed the "Old Dominion" and sometimes the "Mother of Presidents" after the eight U.S. presidents born there...

  • Apochthonius hypogeus Muchmore, 1976 — Virginia
  • Apochthonius indianensis Muchmore, 1967Indiana
    Indiana
    Indiana is a US state, admitted to the United States as the 19th on December 11, 1816. It is located in the Midwestern United States and Great Lakes Region. With 6,483,802 residents, the state is ranked 15th in population and 16th in population density. Indiana is ranked 38th in land area and is...

  • Apochthonius intermedius J.C. Chamberlin, 1929 — Washington
  • Apochthonius irwini R.O. Schuster, 1966 — California
  • Apochthonius knowltoni Muchmore, 1980Wyoming
    Wyoming
    Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...

  • Apochthonius magnanimus Hoff, 1956 — New Mexico
  • Apochthonius malheuri Benedict & Malcolm, 1973 — Oregon
  • Apochthonius maximus R.O. Schuster, 1966 — California
  • Apochthonius minimus R.O. Schuster, 1966British Columbia
    British Columbia
    British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

    , Oregon, Washington
  • Apochthonius minor Muchmore, 1976Georgia
    Georgia (U.S. state)
    Georgia is a state located in the southeastern United States. It was established in 1732, the last of the original Thirteen Colonies. The state is named after King George II of Great Britain. Georgia was the fourth state to ratify the United States Constitution, on January 2, 1788...

  • Apochthonius moestus (Banks, 1891) — Eastern U.S.
  • Apochthonius mysterius Muchmore, 1976 — Missouri
  • Apochthonius occidentalis J.C. Chamberlin, 1929 — Oregon
  • Apochthonius paucispinosus Muchmore, 1967 — West Virginia
  • Apochthonius russelli Muchmore, 1976 — Alabama
  • Apochthonius titanicus Muchmore, 1976 — Arkansas
  • Apochthonius typhlus Muchmore, 1967Missouri
    Missouri
    Missouri is a US state located in the Midwestern United States, bordered by Iowa, Illinois, Kentucky, Tennessee, Arkansas, Oklahoma, Kansas and Nebraska. With a 2010 population of 5,988,927, Missouri is the 18th most populous state in the nation and the fifth most populous in the Midwest. It...


  • Austrochthonius J.C. Chamberlin, 1929
  • Austrochthonius argentinae Hoff, 1950Argentina
    Argentina
    Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

    , Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • Austrochthonius australis Hoff, 1951Australia
    Australia
    Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

  • Austrochthonius bolivianus Beier, 1930Bolivia
    Bolivia
    Bolivia officially known as Plurinational State of Bolivia , is a landlocked country in central South America. It is the poorest country in South America...

  • Austrochthonius cavicola Beier, 1968 — Australia
  • Austrochthonius chilensis (J.C. Chamberlin, 1923)
  • Austrochthonius chilensis chilensis (J.C. Chamberlin, 1923)Chile
    Chile
    Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

    , Argentina
  • Austrochthonius chilensis magalhanicus Beier, 1964 — Chile
  • Austrochthonius chilensis transversus Beier, 1964 — Argentina
    • Austrochthonius easti Harvey, 1991 — western Australia
    • Austrochthonius iguazuensis Vitali-di Castri, 1975 — Brazil
    • Austrochthonius insularis Vitali-di Castri, 1968Crozet Islands
      Crozet Islands
      The Crozet Islands are a sub-antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarctic Lands.-Geography:...

    • Austrochthonius mordax Beier, 1967New Zealand
      New Zealand
      New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

    • Austrochthonius paraguayensis Vitali-di Castri, 1975Paraguay
      Paraguay
      Paraguay , officially the Republic of Paraguay , is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to the east and northeast, and Bolivia to the northwest. Paraguay lies on both banks of the Paraguay River, which runs through the center of the...

    • Austrochthonius parvus (Mello-Leitão, 1939) — Argentina
    • Austrochthonius persimilis Beier, 1930 — Chile
    • Austrochthonius rapax Beier, 1976 — New Zealand
    • Austrochthonius semiserratus Beier, 1930 — Chile
    • Austrochthonius tullgerni (Beier, 1931)Southern Africa
      Southern Africa
      Southern Africa is the southernmost region of the African continent, variably defined by geography or geopolitics. Within the region are numerous territories, including the Republic of South Africa ; nowadays, the simpler term South Africa is generally reserved for the country in English.-UN...

    • Austrochthonius zealandicus Beier, 1966 — New Zealand
  • Austrochthonius zealandicus zealandicus Beier, 1966 — New Zealand
  • Austrochthonius zealandicus obscurus Beier, 1966 — New Zealand

  • Caribchthonius Muchmore, 1976
  • Caribchthonius butleri Muchmore, 1976Virgin Islands
    Virgin Islands
    The Virgin Islands are the western island group of the Leeward Islands, which are the northern part of the Lesser Antilles, which form the border between the Caribbean Sea and the Atlantic Ocean...

  • Caribchthonius orthodentatus Muchmore, 1976Belize
    Belize
    Belize is a constitutional monarchy and the northernmost country in Central America. Belize has a diverse society, comprising many cultures and languages. Even though Kriol and Spanish are spoken among the population, Belize is the only country in Central America where English is the official...


  • Centrochthonius Beier, 1931
  • Centrochthonius koslovi (Redikorzev, 1918)Tibet
    Tibet
    Tibet is a plateau region in Asia, north-east of the Himalayas. It is the traditional homeland of the Tibetan people as well as some other ethnic groups such as Monpas, Qiang, and Lhobas, and is now also inhabited by considerable numbers of Han and Hui people...

    , India
    India
    India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

    , Nepal
    Nepal
    Nepal , officially the Federal Democratic Republic of Nepal, is a landlocked sovereign state located in South Asia. It is located in the Himalayas and bordered to the north by the People's Republic of China, and to the south, east, and west by the Republic of India...

  • Centrochthonius schnitnikovi (Redikorzev, 1934)Russia
    Russia
    Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

  • Centrochthonius sichuanensis Schawaller, 1995Sichuan
    Sichuan
    ' , known formerly in the West by its postal map spellings of Szechwan or Szechuan is a province in Southwest China with its capital in Chengdu...


  • Chiliochthonius Vitali-di Castri, 1975
  • Chiliochthonius centralis Vitali-di Castri, 1975Chile
    Chile
    Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

  • Chiliochthonius montanus Vitali-di Castri, 1975 — Chile
  • Chiliochthonius usuriensis Beier, 1979 — Russia (?)

  • Chthonius C.L. Koch, 1843
  • Chthonius absoloni Beier, 1938Yugoslavia
    Yugoslavia
    Yugoslavia refers to three political entities that existed successively on the western part of the Balkans during most of the 20th century....

  • Chthonius agazzii Beier, 1966Italy
    Italy
    Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

  • Chthonius alpicola Beier, 1951Austria
    Austria
    Austria , officially the Republic of Austria , is a landlocked country of roughly 8.4 million people in Central Europe. It is bordered by the Czech Republic and Germany to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the...

    , Italy
  • Chthonius apollinis Mahnert, 1978Greece
    Greece
    Greece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....

  • Chthonius azerbaidzhanus Schawaller & Dashdamirov, 1988Azerbaijan
    Azerbaijan
    Azerbaijan , officially the Republic of Azerbaijan is the largest country in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded by the Caspian Sea to the east, Russia to the north, Georgia to the northwest, Armenia to the west, and Iran to...

  • Chthonius baccettii Callaini, 1980 — Italy
  • Chthonius balazuci Vachon, 1963France
    France
    The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

  • Chthonius bogovinae Curcic, 1972 — Yugoslavia
  • Chthonius bogovinae bogovinae Curcic, 1972 — Yugoslavia
  • Chthonius bogovinae latidentatus Curcic, 1972 — Yugoslavia
    • Chthonius brandmayri Callaini, 1986 — Yugoslavia
    • Chthonius caprai Gardini, 1977 — Italy
    • Chthonius caoduroi Callaini, 1987 — Italy
    • Chthonius cavernarum Ellingsen, 1909Romania
      Romania
      Romania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...

      , Yugoslavia
    • Chthonius cavophilus Hadzi, 1939Bulgaria
      Bulgaria
      Bulgaria , officially the Republic of Bulgaria , is a parliamentary democracy within a unitary constitutional republic in Southeast Europe. The country borders Romania to the north, Serbia and Macedonia to the west, Greece and Turkey to the south, as well as the Black Sea to the east...

    • Chthonius cebenicus Leclerc, 1981 — France
    • Chthonius cephalotes (Simon, 1875) — France
    • Chthonius chamberlini (Leclerc, 1983)Algeria
      Algeria
      Algeria , officially the People's Democratic Republic of Algeria , also formally referred to as the Democratic and Popular Republic of Algeria, is a country in the Maghreb region of Northwest Africa with Algiers as its capital.In terms of land area, it is the largest country in Africa and the Arab...

      , France
    • Chthonius comottii Inzaghi, 1987 — Italy
    • Chthonius cryptus J.C. Chamberlin, 1962 — Greece
    • Chthonius dacnodes Navás, 1918Spain
      Spain
      Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

      , Azores
      Azores
      The Archipelago of the Azores is composed of nine volcanic islands situated in the middle of the North Atlantic Ocean, and is located about west from Lisbon and about east from the east coast of North America. The islands, and their economic exclusion zone, form the Autonomous Region of the...

      , Italy
    • Chthonius dalmatinus Hadzi, 1930 — Yugoslavia
    • Chthonius densedentatus Beier, 1938 — Italy, Yugoslavia
    • Chthonius diophthalmus Daday, 1888 — eastern Europe
    • Chthonius doderoi Beier, 1930 — France, Italy
  • Chthonius doderoi doderoi Beier, 1930 — France, Italy
  • Chthonius doderoi horridus Beier, 1934 — Italy
    • Chthonius ellingseni Beier, 1928 — Italy, Yugoslavia
    • Chthonius elongatus Lazzeroni, 1970 — Italy
    • Chthonius euganeus Gardini, 1991 — Italy
    • Chthonius exarmatus Beier, 1939 — Yugoslavia
    • Chthonius graecus Beier, 1963 — Greece
    • Chthonius guglielmii Callaini, 1986 — Italy
    • Chthonius halberti Kew, 1916 — France, Great Britain
      Great Britain
      Great Britain or Britain is an island situated to the northwest of Continental Europe. It is the ninth largest island in the world, and the largest European island, as well as the largest of the British Isles...

    • Chthonius herbarii Mahnert, 1980 — Crete
    • Chthonius heterodactylus Tomosvary, 1882central Europe
      Central Europe
      Central Europe or alternatively Middle Europe is a region of the European continent lying between the variously defined areas of Eastern and Western Europe...

    • Chthonius heutaultae Leclerc, 1981 — France
    • Chthonius hungaricus Mahnert, 1981Hungary
      Hungary
      Hungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...

    • Chthonius ilvensis Beier, 1963 — Italy
    • Chthonius imperator Mahnert, 1978 — Greece
    • Chthonius ischnocheles (Hermann, 1804)Europe
      Europe
      Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

      , U.S.
  • Chthonius ischnocheles ischnocheles (Hermann, 1804) — Europe, U.S.
  • Chthonius ischnocheles reductus Beier, 1939southern Europe
    Southern Europe
    The term Southern Europe, at its most general definition, is used to mean "all countries in the south of Europe". However, the concept, at different times, has had different meanings, providing additional political, linguistic and cultural context to the definition in addition to the typical...

  • Chthonius ischnocheles ruffoi Caporiacco, 1951 — Italy
  • Chthonius ischnocheles stammeri Beier, 1942 — Italy
    • Chthonius ischnocheloides Beier, 1974 — Italy
    • Chthonius italicus Beier, 1930 — Italy
    • Chthonius iugoslavicus Curcic, 1972 — Yugoslavia
    • Chthonius jalzici Curcic, 1988 — Yugoslavia
    • Chthonius jonicus Beier, 1931 — southern Europe
    • Chthonius jugorum Beier, 1952 — Italy
    • Chthonius karamanianus Hadzi, 1937 — Yugoslavia
    • Chthonius lanzai Caporiacco, 1947 — Italy
  • Chthonius lanzai lanzai Caporiacco, 1947 — Italy
  • Chthonius lanzai vannii Callaini, 1986 — Italy
    • Chthonius leoi (Callaini, 1988)Sardinia
      Sardinia
      Sardinia is the second-largest island in the Mediterranean Sea . It is an autonomous region of Italy, and the nearest land masses are the French island of Corsica, the Italian Peninsula, Sicily, Tunisia and the Spanish Balearic Islands.The name Sardinia is from the pre-Roman noun *sard[],...

    • Chthonius leruthi Beier, 1939 — Romania
    • Chthonius lesnik Curcic, 1994 — Yugoslavia
    • Chthonius lessiniensis Schawaller, 1982 — Italy
    • Chthonius ligusticus Beier, 1930 — Italy
    • Chthonius lindbergi Beier, 1956 — Crete
    • Chthonius litoralis Hadzi, 1933 — Italy, Yugoslavia
    • Chthonius lucifugus Mahnert, 1977 — France, Spain
    • Chthonius macedonicus Curcic, 1972 — Yugoslavia
    • Chthonius magnificus Beier, 1938 — Yugoslavia
    • Chthonius malatestai Callaini, 1980 — Italy
    • Chthonius mauritanicus (Callaini, 1988)Morocco
      Morocco
      Morocco , officially the Kingdom of Morocco , is a country located in North Africa. It has a population of more than 32 million and an area of 710,850 km², and also primarily administers the disputed region of the Western Sahara...

    • Chthonius mayi Heurtault-Rossi, 1968 — France
    • Chthonius mazaurici Leclerc, 1981 — France
  • Chthonius mazaurici mazaurici Leclerc, 1981 — France
  • Chthonius mazaurici coironi Leclerc, 1981 — France
    • Chthonius mengei Beier, 1937fossil
      Fossil
      Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

      : Baltic amber
      Baltic amber
      The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite, with about 80% of the world's known amber found there. It dates from 44 million years ago...

    • Chthonius microphthalmus Simon, 1879 — France, Spain
    • Chthonius mingazzinii Callaini, 1991 — Italy
    • Chthonius minotaurus Henderickx, 1997 — Kournas Cave, Crete
      Crete
      Crete is the largest and most populous of the Greek islands, the fifth largest island in the Mediterranean Sea, and one of the thirteen administrative regions of Greece. It forms a significant part of the economy and cultural heritage of Greece while retaining its own local cultural traits...

    • Chthonius monicae Boghean, 1989 — Romania
    • Chthonius motasi Dumitresco & Orghidan, 1964 — Romania
    • Chthonius multidentatus Beier, 1963Sicily
      Sicily
      Sicily is a region of Italy, and is the largest island in the Mediterranean Sea. Along with the surrounding minor islands, it constitutes an autonomous region of Italy, the Regione Autonoma Siciliana Sicily has a rich and unique culture, especially with regard to the arts, music, literature,...

    • Chthonius occultus Beier, 1939 — Yugoslavia
    • Chthonius orthodactyloides Beier, 1967Turkey
      Turkey
      Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

    • Chthonius orthodactylus (Leach, 1817) — Europe
    • Chthonius pacificus Muchmore, 1968California
      California
      California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

    • Chthonius paganus (Hoff, 1961)Colorado
      Colorado
      Colorado is a U.S. state that encompasses much of the Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains...

    • Chthonius paludis (J.C. Chamberlin, 1929) — southern U.S.
    • Chthonius paolettii Beier, 1974 — Italy
    • Chthonius persimilis Beier, 1939 — Yugoslavia
    • Chthonius petrochilosi Heurtault, 1972 — Greece
    • Chthonius pivai Gardini, 1991 — Italy
    • Chthonius ponticoides Mahnert, 1975 — Greece
    • Chthonius ponticus Beier, 1965 — Turkey
    • Chthonius porevidi Curcic, Makarov & Lucic, 1998 — Krivosije, Knezlaz Pecina Cave, Montenegro
      Montenegro
      Montenegro Montenegrin: Crna Gora Црна Гора , meaning "Black Mountain") is a country located in Southeastern Europe. It has a coast on the Adriatic Sea to the south-west and is bordered by Croatia to the west, Bosnia and Herzegovina to the northwest, Serbia to the northeast and Albania to the...

    • Chthonius pristinus Schawaller, 1978 — fossil: Baltic amber
    • Chthonius prove Curcic, Dimitrijevic & Makarov, 1997 — Gornji Morinj, Serbia
      Serbia
      Serbia , officially the Republic of Serbia , is a landlocked country located at the crossroads of Central and Southeast Europe, covering the southern part of the Carpathian basin and the central part of the Balkans...

      , Montenegro
    • Chthonius pusillus Beier, 1947 — Austria
    • Chthonius pygmaeus Beier, 1934 — central Europe
  • Chthonius pygmaeus pygmaeus Beier, 1934 — central Europe
  • Chthonius pygmaeus carinthiacus Beier, 1951 — central Europe
    • Chthonius radjai Curcic, 1988 — Yugoslavia
    • Chthonius raridentatus Hadzi, 1930 — Yugoslavia
    • Chthonius ressli Beier, 1956 — Austria, France, Italy
    • Chthonius satapliaensis Schawaller & Dashdamirov, 1988Russia
      Russia
      Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

    • Chthonius sestasi Mahnert, 1980 — Greece
    • Chthonius shelkovnikovi Redikorzev, 1930 — western Asia
      Asia
      Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

    • Chthonius shulovi Beier, 1963 — Israel
    • Chthonius stevanovici Curcic, 1986 — Yugoslavia
    • Chthonius strinatii Mahnert, 1975 — Greece
    • Chthonius submontanus Beier, 1963 — central Europe
    • Chthonius subterraneus Beier, 1931 — Hungary, Yugoslavia
  • Chthonius subterraneus subterraneus Beier, 1931 — Hungary, Yugoslavia
  • Chthonius subterraneus meuseli Beier, 1939 — Yugoslavia
    • Chthonius tadzhikistanicus Dashdamirov & Schawaller, 1992Tajikistan
      Tajikistan
      Tajikistan , officially the Republic of Tajikistan , is a mountainous landlocked country in Central Asia. Afghanistan borders it to the south, Uzbekistan to the west, Kyrgyzstan to the north, and China to the east....

    • Chthonius tenuis L. Koch, 1873 — Europe, Algeria
    • Chthonius thessalus Mahnert, 1980 — Greece
    • Chthonius trebinjensis Beier, 1938 — Yugoslavia
    • Chthonius troglobius Hadzi, 1937 — Yugoslavia
    • Chthonius troglodites Redikorzev, 1928 — Bulgaria
    • Chthonius tzanoudakisi Mahnert, 1975 — Greece
    • Chthonius aegatensis (Callaini, 1989) — Sicilly
    • Chthonius aguileraorum Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2000 — Cadiz, Villaluenga del Rosario, Cueva de la Hiedra, Spain
    • Chthonius amatei Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
    • Chthonius anatolicus (Beier, 1969)Iran
      Iran
      Iran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia...

      , Turkey
    • Chthonius anophthalmus (Ellingsen, 1908) — Algeria
    • Chthonius apulicus (Beier, 1958) — Italy
    • Chthonius asturiensis (Beier, 1955) — Spain
    • Chthonius atlantis (Mahnert, 1980) — Morocco
    • Chthonius balearicus (Mahnert, 1977)Balearic Islands
      Balearic Islands
      The Balearic Islands are an archipelago of Spain in the western Mediterranean Sea, near the eastern coast of the Iberian Peninsula.The four largest islands are: Majorca, Minorca, Ibiza and Formentera. The archipelago forms an autonomous community and a province of Spain with Palma as the capital...

    • Chthonius bartolii (Gardini, 1976) — Italy
    • Chthonius bauneensis (Callaini, 1983) — Sardinia
    • Chthonius beieri (Lazzeroni, 1966) — Italy
    • Chthonius bellesi (Mahnert, 1989) — Balearic Islands
    • Chthonius berninii (Callaini, 1983) — Sardinia
    • Chthonius bidentatus (Beier, 1938) — Yugoslavia
    • Chthonius boldorii (Beier, 1934) — Italy, Yugoslavia
    • Chthonius bolivari (Beier, 1930) — Spain
    • Chthonius cabreriensis Mahnert, 1993 — Balearic Islands
    • Chthonius cassolai (Beier, 1974) — Sardinia
    • Chthonius catalonicus (Beier, 1939) — Spain
    • Chthonius cavicola Gardini, 1990 — Italy
    • Chthonius chius Schawaller, 1990 — Greece
    • Chthonius concii (Beier, 1953) — Italy
    • Chthonius corcyraeus (Mahnert, 1976) — Greece
    • Chthonius corsicus (Callaini, 1981) — Corsica
    • Chthonius creticus (Mahnert, 1980) — Crete
    • Chthonius daedaleus (Mahnert, 1980) — Crete
    • Chthonius distinguendus (Beier, 1930) — Spain
    • Chthonius dubius Mahnert, 1993 — Canary Islands
    • Chthonius elbanus (Beier, 1963) — Italy, Morocco
    • Chthonius fuscimanus (Simon, 1900) — central Europe, Turkey
    • Chthonius gasparoi (Gardini, 1989) — Greece
    • Chthonius genuensis Gardini, 1990 — Italy
    • Chthonius gestroi (Simon, 1989) — Italy
    • Chthonius gibbus (Beier, 1953) — Spain, Italy
    • Chthonius girgentiensis (Mahnert, 1982)Malta
      Malta
      Malta , officially known as the Republic of Malta , is a Southern European country consisting of an archipelago situated in the centre of the Mediterranean, south of Sicily, east of Tunisia and north of Libya, with Gibraltar to the west and Alexandria to the east.Malta covers just over in...

    • Chthonius giustii (Callaini, 1981) — Corsica
    • Chthonius gracilimanus Mahnert, 1997 — La Palma, Mazo, Salto de Tigalate, Canary Islands
      Canary Islands
      The Canary Islands , also known as the Canaries , is a Spanish archipelago located just off the northwest coast of mainland Africa, 100 km west of the border between Morocco and the Western Sahara. The Canaries are a Spanish autonomous community and an outermost region of the European Union...

    • Chthonius grafittii (Gardini, 1981) — Sardinia
    • Chthonius hiberus (Beier, 1930) — Spain
    • Chthonius hispanus (Beier, 1930) — Spain, Portugal
    • Chthonius insularis (Beier, 1938) — Yugoslavia
    • Chthonius iranicus (Beier, 1971) — Iran
    • Chthonius kabylicus (Callaini, 1983) — Algeria
    • Chthonius kemza Curcic, Lee & Makarov, 1993 — Yugoslavia
    • Chthonius kewi (Gabbutt, 1966) — Great Britain
    • Chthonius longesetosus (Mahnert, 1976) — Morocco
    • Chthonius lucanus (Callaini, 1984) — Italy
    • Chthonius machadoi (Vachon, 1940) — Spain, Morocco
  • Chthonius machadoi machadoi (Vachon, 1940) — Spain, Morocco
  • Chthonius machadoi canariensis (Beier, 1965) — Canary Islands
    • Chthonius mahnerti (Zaragoza, 1984) — Spain
    • Chthonius maltensis (Mahnert, 1975) — Malta
    • Chthonius mariolae Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
    • Chthonius maroccanus (Mahnert, 1980) — Morocco
    • Chthonius mayorali Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
    • Chthonius microtuberculatus (Hadzi, 1937) — Bulgaria, Yugoslavia
    • Chthonius minous (Mahnert, 1980) — Crete
  • Chthonius minous minous (Mahnert, 1980) — Crete
  • Chthonius minous peramae (Mahnert, 1980) — Crete
    • Chthonius minutus (Vachon, 1940)Portugal
      Portugal
      Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...

    • Chthonius nanus (Beier, 1953) — Italy
    • Chthonius nerjaensis Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
    • Chthonius nidicola (Mahnert, 1979) — Switzerland
    • Chthonius parmensis (Beier, 1963) — Italy
    • Chthonius pieltaini (Beier, 1930) — Italy
    • Chthonius pinai (Zaragoza, 1985) — Spain
    • Chthonius platakisi (Mahnert, 1980) — Crete
    • Chthonius poeninus (Mahnert, 1979) — Switzerland
    • Chthonius ponsi Mahnert, 1993 — Balearic Islands
    • Chthonius poseidonis Gardini, 1993 — Sardinia
    • Chthonius pyrenaicus (Beier, 1934) — France, Spain
    • Chthonius remyi (Heurtault, 1975) — Corsica
    • Chthonius rimicola Mahnert, 1993 — Canary Islands
    • Chthonius romanicus (Beier, 1935) — Greece, Iran
    • Chthonius ruizporteroi Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
    • Chthonius sacer (Beier, 1963)Israel
      Israel
      The State of Israel is a parliamentary republic located in the Middle East, along the eastern shore of the Mediterranean Sea...

    • Chthonius samius (Mahnert, 1982) — Greece
    • Chthonius schmalfussi Schawaller, 1990 — Greece
    • Chthonius scythicus Georgescu & Capuse, 1994 — Romania
    • Chthonius sendrai (Zaragoza, 1985) — Spain
    • Chthonius serbicus (Hadzi, 1937) — Bulgaria, Yugoslavia
    • Chthonius setosus Mahnert, 1993 — Canary Islands
    • Chthonius siculus (Beier, 1961) — Italy, Spain
    • Chthonius siscoensis (Heurtault, 1975) — Corsica
    • Chthonius tetrachelatus (Preyssler, 1790)Cosmopolitan
      Cosmopolitan distribution
      In biogeography, a taxon is said to have a cosmopolitan distribution if its range extends across all or most of the world in appropriate habitats. For instance, the killer whale has a cosmopolitan distribution, extending over most of the world's oceans. Other examples include humans, the lichen...

    • Chthonius troglophilus (Beier, 1920) — Italy
    • Chthonius tuberculatus (Hadzi, 1937) — central Europe, Greece
    • Chthonius vachoni (Heurtault-Rossi, 1963) — France
    • Chthonius ventalloi (Beier, 1939) — Spain
  • Chthonius ventalloi ventalloi (Beier, 1939) — Spain
  • Chthonius ventalloi cazorlensis (Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001) — Spain
    • Chthonius verai (Zaragoza, 1987) — Spain
    • Chthonius virginicus (J.C. Chamberlin, 1929) — Eastern U.S.
    • Chthonius zoiai Gardini, 1990 — Italy
    • Chthonius abnormis (Beier, 1939) — Yugoslavia
    • Chthonius caligatus (Beier, 1938) — Yugoslavia
    • Chthonius cavernicola (Beier, 1938) — Yugoslavia
    • Chthonius cerberus (Beier, 1938) — Yugoslavia
    • Chthonius globifer (Simon, 1879) — France, Switzerland
    • Chthonius pancici (Curbic, 1972) — Yugoslavia
    • Chthonius polychaetus (Hadzi, 1937) — Yugoslavia
    • Chthonius purgo Curcic, Lee & Makarov, 1993 — Yugoslavia
    • Chthonius simplex (Beier, 1939) — Yugoslavia
    • Chthonius spelaeophilus (Hadzi, 1930) — Yugoslavia
  • Chthonius spelaeophilus spelaeophilus (Hadzi, 1930) — Yugoslavia
  • Chthonius spelaeophilus histricus (Beier, 1931) — Yugoslavia, Italy
    • Chthonius vandeli (Dumitresco & Orghidan, 1964) — Romania
    • Chthonius californicus (J.C. Chamberlin, 1929) — California
    • Chthonius oregonicus (Muchmore, 1968) — Oregon
    • Chthonius spingolus (R.O. Schuster, 1962) — California
    • Chthonius karamani (Hadzi, 1933) — Yugoslavia
    • Chthonius rogatus (Beier, 1938) — Yugoslavia

  • Congochthonius Beier, 1959
  • Congochthonius nanus Beier, 1959 — Democratic Republic of Congo

  • Drepanochthonius Beier, 1964
  • Drepanochthonius horridus Beier, 1964Chile
    Chile
    Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...


  • Francochthonius Vitali-di Castri, 1975
  • Francochthonius Vitali-di Castri, 1975 — Chile

  • Kleptochthonius J.C. Chamberlin, 1949
  • Kleptochthonius crosbyi (J.C. Chamberlin, 1929)Kentucky
    Kentucky
    The Commonwealth of Kentucky is a state located in the East Central United States of America. As classified by the United States Census Bureau, Kentucky is a Southern state, more specifically in the East South Central region. Kentucky is one of four U.S. states constituted as a commonwealth...

    , North Carolina
    North Carolina
    North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

  • Kleptochthonius geophilus Malcolm & J.C. Chamberlin, 1961Oregon
    Oregon
    Oregon is a state in the Pacific Northwest region of the United States. It is located on the Pacific coast, with Washington to the north, California to the south, Nevada on the southeast and Idaho to the east. The Columbia and Snake rivers delineate much of Oregon's northern and eastern...

  • Kleptochthonius griseomanus Muchmore, 2000 — Hoosier National Forest, Indian Cave, Indiana
    Indiana
    Indiana is a US state, admitted to the United States as the 19th on December 11, 1816. It is located in the Midwestern United States and Great Lakes Region. With 6,483,802 residents, the state is ranked 15th in population and 16th in population density. Indiana is ranked 38th in land area and is...

  • Kleptochthonius inusitatus Muchmore, 1994Ohio
    Ohio
    Ohio is a Midwestern state in the United States. The 34th largest state by area in the U.S.,it is the 7th‑most populous with over 11.5 million residents, containing several major American cities and seven metropolitan areas with populations of 500,000 or more.The state's capital is Columbus...

  • Kleptochthonius lewisorum Muchmore, 2000 — Harrison County, Baelz Cave, Indiana
  • Kleptochthonius magnus Muchmore, 1966Tennessee
    Tennessee
    Tennessee is a U.S. state located in the Southeastern United States. It has a population of 6,346,105, making it the nation's 17th-largest state by population, and covers , making it the 36th-largest by total land area...

  • Kleptochthonius multispinosus (Hoff, 1945) — Eastern U.S.
  • Kleptochthonius oregonus Malcolm & J.C. Chamberlin, 1961 — Oregon
  • Kleptochthonius polychaetus Muchmore, 1994Virginia
    Virginia
    The Commonwealth of Virginia , is a U.S. state on the Atlantic Coast of the Southern United States. Virginia is nicknamed the "Old Dominion" and sometimes the "Mother of Presidents" after the eight U.S. presidents born there...

  • Kleptochthonius sheari Muchmore, 1994 — West Virginia
  • Kleptochthonius affinis (Muchmore, 1976) — Tennessee
  • Kleptochthonius anophthalmus (Muchmore, 1970) — Virginia
  • Kleptochthonius attenuatus (Malcolm & J.C. Chamberlin, 1961) — Kentucky
  • Kleptochthonius barri (Muchmore, 1965) — Tennessee
  • Kleptochthonius binoculatus (Muchmore, 1974) — Virginia
  • Kleptochthonius cerberus (Malcolm & J.C. Chamberlin, 1961) — Kentucky
  • Kleptochthonius charon (Muchmore, 1965) — Tennessee
  • Kleptochthonius daemonius (Muchmore, 1965) — Tennessee
  • Kleptochthonius erebicus (Muchmore, 1965) — Kentucky
  • Kleptochthonius gertschi (Malcolm & J.C. Chamberlin, 1961) — Virginia
  • Kleptochthonius hageni (Muchmore, 1963) — Kentucky
  • Kleptochthonius henroti (Vachon, 1952) West Virginia
  • Kleptochthonius hetricki (Muchmore, 1974) — West Virginia
  • Kleptochthonius hubrichti (Muchmore, 1965) — Kentucky
  • Kleptochthonius infernalis (Malcolm & J.C. Chamberlin, 1961) — Tennessee
  • Kleptochthonius krekeleri (Muchmore, 1965) — Kentucky
  • Kleptochthonius lutzi (Malcolm & J.C. Chamberlin, 1961); — Tennessee
  • Kleptochthonius microphthalmus (Malcolm & J.C. Chamberlin, 1961) — Kentucky
  • Kleptochthonius myoplus (Malcolm & J.C. Chamberlin, 1961) — Tennessee
  • Kleptochthonius orpheus (Muchmore, 1965) — West Virginia
  • Kleptochthonius packardi (Hagen, 1879)
  • Kleptochthonius pluto (Muchmore, 1965) — Tennessee
  • Kleptochthonius proserpinae (Muchmore, 1965) — West Virginia
  • Kleptochthonius proximosetus (Muchmore, 1976) — Virginia
  • Kleptochthonius regulus (Muchmore, 1970) — Virginia
  • Kleptochthonius rex (Malcolm & J.C. Chamberlin, 1961) — Tennessee
  • Kleptochthonius similis (Muchmore, 1976) — Virginia
  • Kleptochthonius stygius (Muchmore, 1965) — Tennessee
  • Kleptochthonius tantalus (Muchmore, 1966) — Tennessee

  • Lagynochthonius Beier, 1951
  • Lagynochthonius annamensis (Beier, 1951)Vietnam
    Vietnam
    Vietnam – sometimes spelled Viet Nam , officially the Socialist Republic of Vietnam – is the easternmost country on the Indochina Peninsula in Southeast Asia. It is bordered by China to the north, Laos to the northwest, Cambodia to the southwest, and the South China Sea –...

  • Lagynochthonius arctus (Beier, 1967)New Guinea
    New Guinea
    New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

  • Lagynochthonius australicus (Beier, 1966)Australia
    Australia
    Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

  • Lagynochthonius bakeri (J.C. Chamberlin, 1929)Philippines
    Philippines
    The Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...

  • Lagynochthonius brincki (Beier, 1973)Sri Lanka
    Sri Lanka
    Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka is a country off the southern coast of the Indian subcontinent. Known until 1972 as Ceylon , Sri Lanka is an island surrounded by the Indian Ocean, the Gulf of Mannar and the Palk Strait, and lies in the vicinity of India and the...

  • Lagynochthonius curvidigitatus Mahnert, 1997Tenerife
    Tenerife
    Tenerife is the largest and most populous island of the seven Canary Islands, it is also the most populated island of Spain, with a land area of 2,034.38 km² and 906,854 inhabitants, 43% of the total population of the Canary Islands. About five million tourists visit Tenerife each year, the...

    , Cueva Felipe Reventon, Canary Islands
    Canary Islands
    The Canary Islands , also known as the Canaries , is a Spanish archipelago located just off the northwest coast of mainland Africa, 100 km west of the border between Morocco and the Western Sahara. The Canaries are a Spanish autonomous community and an outermost region of the European Union...

  • Lagynochthonius dybasi (Beier, 1957)Palau
    Palau
    Palau , officially the Republic of Palau , is an island nation in the Pacific Ocean, east of the Philippines and south of Tokyo. In 1978, after three decades as being part of the United Nations trusteeship, Palau chose independence instead of becoming part of the Federated States of Micronesia, a...

  • Lagynochthonius exiguus (Beier, 1952)Malaysia
  • Lagynochthonius flavus (Mahnert, 1986)Kenya
    Kenya
    Kenya , officially known as the Republic of Kenya, is a country in East Africa that lies on the equator, with the Indian Ocean to its south-east...

  • Lagynochthonius guasirih (Mahnert, 1988)Sarawak
    Sarawak
    Sarawak is one of two Malaysian states on the island of Borneo. Known as Bumi Kenyalang , Sarawak is situated on the north-west of the island. It is the largest state in Malaysia followed by Sabah, the second largest state located to the North- East.The administrative capital is Kuching, which...

  • Lagynochthonius hamatus Harvey, 1988Sumatra
    Sumatra
    Sumatra is an island in western Indonesia, westernmost of the Sunda Islands. It is the largest island entirely in Indonesia , and the sixth largest island in the world at 473,481 km2 with a population of 50,365,538...

  • Lagynochthonius himalayensis (Morikawa, 1968)Nepal
    Nepal
    Nepal , officially the Federal Democratic Republic of Nepal, is a landlocked sovereign state located in South Asia. It is located in the Himalayas and bordered to the north by the People's Republic of China, and to the south, east, and west by the Republic of India...

  • Lagynochthonius hygricus Murthy & Ananthakrishnan, 1977India
    India
    India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

  • Lagynochthonius indicus (Murthy & Ananthakrishnan, 1977) — India
  • Lagynochthonius johni (Redikorzev, 1922)Indonesia
    Indonesia
    Indonesia , officially the Republic of Indonesia , is a country in Southeast Asia and Oceania. Indonesia is an archipelago comprising approximately 13,000 islands. It has 33 provinces with over 238 million people, and is the world's fourth most populous country. Indonesia is a republic, with an...

    , Philippines
  • Lagynochthonius kapi Harvey, 1988 — Indonesia
  • Lagynochthonius kenyensis (Mahnert, 1986) — Kenya
  • Lagynochthonius mordor Harvey, 1989 — Australia
  • Lagynochthonius nagaminei (Sato, 1983)Japan
    Japan
    Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

  • Lagynochthonius novaeguineae (Beier, 1965) — New Guinea
  • Lagynochthonius paucedentatus (Beier, 1955) — Malaysia
  • Lagynochthonius ponapensis (Beier, 1957)Caroline Islands
    Caroline Islands
    The Caroline Islands are a widely scattered archipelago of tiny islands in the western Pacific Ocean, to the north of New Guinea. Politically they are divided between the Federated States of Micronesia in the eastern part of the group, and Palau at the extreme western end...

  • Lagynochthonius roeweri (J.C. Chamberlin, 1930)Java
    Java
    Java is an island of Indonesia. With a population of 135 million , it is the world's most populous island, and one of the most densely populated regions in the world. It is home to 60% of Indonesia's population. The Indonesian capital city, Jakarta, is in west Java...

  • Lagynochthonius salomonensis (Beier, 1966)Solomon Islands
    Solomon Islands
    Solomon Islands is a sovereign state in Oceania, east of Papua New Guinea, consisting of nearly one thousand islands. It covers a land mass of . The capital, Honiara, is located on the island of Guadalcanal...

  • Lagynochthonius sinensis (Beier, 1967)China
    China
    Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

  • Lagynochthonius thorntoni Harvey, 1988 — Java
  • Lagynochthonius tonkinensis (Beier, 1951) — Vietnam

  • Malcolmochthonius Benedict, 1978
  • Malcolmochthonius malcolmi Benedict, 1978 — Oregon
  • Malcolmochthonius oregonus Benedict, 1978 — Oregon
  • Malcolmochthonius perplexus Benedict, 1978 — California

  • Maorichthonius J.C. Chamberlin, 1925
  • Maorichthonius mortenseni J.C. Chamberlin, 1925 — New Zealand

  • Mexichthonius Muchmore, 1975
  • Mexichthonius exoticus Muchmore, 1996Texas
    Texas
    Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

  • Mexichthonius pacal Muchmore, 1978Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

  • Mexichthonius unicus Muchmore, 1975 — Mexico

  • Mundochthonius J.C. Chamberlin, 1929
  • Mundochthonius alpinus Beier, 1947Austria
    Austria
    Austria , officially the Republic of Austria , is a landlocked country of roughly 8.4 million people in Central Europe. It is bordered by the Czech Republic and Germany to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the...

  • Mundochthonius basarukini Schawaller, 1989Russia
    Russia
    Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

  • Mundochthonius bifurcatus Kim & Hong, 1994South Korea
    South Korea
    The Republic of Korea , , is a sovereign state in East Asia, located on the southern portion of the Korean Peninsula. It is neighbored by the People's Republic of China to the west, Japan to the east, North Korea to the north, and the East China Sea and Republic of China to the south...

  • Mundochthonius carpaticus Rafalski, 1948Poland
    Poland
    Poland , officially the Republic of Poland , is a country in Central Europe bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian exclave, to the north...

    , Ukraine
    Ukraine
    Ukraine is a country in Eastern Europe. It has an area of 603,628 km², making it the second largest contiguous country on the European continent, after Russia...

  • Mundochthonius cavernicola Muchmore, 1968Illinois
    Illinois
    Illinois is the fifth-most populous state of the United States of America, and is often noted for being a microcosm of the entire country. With Chicago in the northeast, small industrial cities and great agricultural productivity in central and northern Illinois, and natural resources like coal,...

  • Mundochthonius decoui Dumitresco & Orghidan, 1970Romania
    Romania
    Romania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...

  • Mundochthonius dominicanus Muchmore, 1996Dominican Republic
    Dominican Republic
    The Dominican Republic is a nation on the island of La Hispaniola, part of the Greater Antilles archipelago in the Caribbean region. The western third of the island is occupied by the nation of Haiti, making Hispaniola one of two Caribbean islands that are shared by two countries...

  • Mundochthonius erosidens J.C. Chamberlin, 1929California
    California
    California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

  • Mundochthonius holsingeri Benedict & Malcolm, 1974Virginia
    Virginia
    The Commonwealth of Virginia , is a U.S. state on the Atlantic Coast of the Southern United States. Virginia is nicknamed the "Old Dominion" and sometimes the "Mother of Presidents" after the eight U.S. presidents born there...

  • Mundochthonius japonicus J.C. Chamberlin, 1929Japan
    Japan
    Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

  • Mundochthonius japonicus japonicus J.C. Chamberlin, 1929 — Japan
  • Mundochthonius japonicus imadatei Morikawa, 1956 — Japan
  • Mundochthonius japonicus scolytidis Morikawa, 1954 — Japan
  • Mundochthonius japonicus tripartitus Morikawa, 1956 — Japan
    • Mundochthonius kiyoshii Sakayori, 2002 — Japan
    • Mundochthonius magnus J.C. Chamberlin, 1929 — California
    • Mundochthonius mexicanus Muchmore, 1973Mexico
      Mexico
      The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

    • Mundochthonius minusculus Kim & Hong, 1994 — South Korea
    • Mundochthonius montanus J.C. Chamberlin, 1929Colorado
      Colorado
      Colorado is a U.S. state that encompasses much of the Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains...

      , New Mexico
      New Mexico
      New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...

    • Mundochthonius pacificus (Banks, 1893) — western U.S.
    • Mundochthonius rossi Hoff, 1949 — central U.S.
    • Mundochthonius sandersoni Hoff, 1949 — Illinois
    • Mundochthonius singularis Muchmore, 2001 — Fremont County, Canon City, Fly Cave, Colorado
    • Mundochthonius styriacus Beier, 1971 — Austria
    • Mundochthonius ussuricus Beier, 1979 — Russia

  • Neochthonius J.C. Chamberlin, 1929
  • Neochthonius amplus (R.O. Schuster, 1962)California
    California
    California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

  • Neochthonius decoui (Georgescu & Capuse, 1994)Romania
    Romania
    Romania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...

  • Neochthonius imperialis Muchmore, 1996 — California
  • Neochthonius stanfordianus J.C. Chamberlin, 1929 — California
  • Neochthonius troglodytes Muchmore, 1969 — California

  • Paraliochthonius Beier, 1956
  • Paraliochthonius azanius Mahnert, 1986Kenya
    Kenya
    Kenya , officially known as the Republic of Kenya, is a country in East Africa that lies on the equator, with the Indian Ocean to its south-east...

  • Paraliochthonius canariensis Vachon, 1961Canary Islands
    Canary Islands
    The Canary Islands , also known as the Canaries , is a Spanish archipelago located just off the northwest coast of mainland Africa, 100 km west of the border between Morocco and the Western Sahara. The Canaries are a Spanish autonomous community and an outermost region of the European Union...

  • Paraliochthonius carpenteri Muchmore, 1984 — Bahamas
  • Paraliochthonius hoestlandti Vachon, 1960 — Madeira Islands
  • Paraliochthonius hoestlandti hoestlandti Vachon, 1960 — Madeira Islands
    • Paraliochthonius insulae Hoff, 1963 — Jamaica
    • Paraliochthonius johnstoni (J.C. Chamberlin, 1923) — Mexico
    • Paraliochthonius martini Mahnert, 1989 — Canary Islands
    • Paraliochthonius mirus Mahnert, 2002 — Canary Islands
    • Paraliochthonius puertoricensis Muchmore, 1967Puerto Rico
      Puerto Rico
      Puerto Rico , officially the Commonwealth of Puerto Rico , is an unincorporated territory of the United States, located in the northeastern Caribbean, east of the Dominican Republic and west of both the United States Virgin Islands and the British Virgin Islands.Puerto Rico comprises an...

    • Paraliochthonius singularis (Menozzi, 1924)Europe
      Europe
      Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

    • Paraliochthonius takashimai (Morikawa, 1958) — Japan
    • Paraliochthonius tenebrarum Mahnert, 1989 — Canary Islands
    • Paraliochthonius weygoldti Muchmore, 1967Florida
      Florida
      Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...


  • Pseudochthonius Balzan, 1892
  • Pseudochthonius arubensis Wagenaar-Hummelinck, 1948Aruba
    Aruba
    Aruba is a 33 km-long island of the Lesser Antilles in the southern Caribbean Sea, located 27 km north of the coast of Venezuela and 130 km east of Guajira Peninsula...

  • Pseudochthonius beieri Mahnert, 1978 — Republic of Congo
  • Pseudochthonius billae Vachon, 1941 — Ivory Coast
  • Pseudochthonius biseriatus Mahnert, 2001 — Minas Gerais, Itacarambi, Gruta Olhos dAgua, Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • Pseudochthonius brasiliensis Beier, 1970 — Brazil
  • Pseudochthonius clarus Hoff, 1963Jamaica
    Jamaica
    Jamaica is an island nation of the Greater Antilles, in length, up to in width and 10,990 square kilometres in area. It is situated in the Caribbean Sea, about south of Cuba, and west of Hispaniola, the island harbouring the nation-states Haiti and the Dominican Republic...

  • Pseudochthonius congicus Beier, 1959 — Democratic Republic of Congo
  • Pseudochthonius doctus Hoff, 1963 — Jamaica
  • Pseudochthonius falcatus Muchmore, 1977Belize
    Belize
    Belize is a constitutional monarchy and the northernmost country in Central America. Belize has a diverse society, comprising many cultures and languages. Even though Kriol and Spanish are spoken among the population, Belize is the only country in Central America where English is the official...

  • Pseudochthonius galapagensis Beier, 1977 — Galapagos
  • Pseudochthonius gracilmanus Mahnert, 2001 — Bahia, Gruta Azul, Iraquara, Brazil
  • Pseudochthonius heterodentatus Hoff, 1946Trinidad
    Trinidad
    Trinidad is the larger and more populous of the two major islands and numerous landforms which make up the island nation of Trinidad and Tobago. It is the southernmost island in the Caribbean and lies just off the northeastern coast of Venezuela. With an area of it is also the fifth largest in...

  • Pseudochthonius homodentatus J.C. Chamberlin, 1929Venezuela
    Venezuela
    Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...

  • Pseudochthonius insularis J.C. Chamberlin, 1929St. Vincent
    Saint Vincent (island)
    Saint Vincent is a volcanic island in the Caribbean. It is the largest island of the chain called Saint Vincent and the Grenadines. It is located in the Caribbean Sea, between Saint Lucia and Grenada. It is composed of partially submerged volcanic mountains...

  • Pseudochthonius leleupi Beier, 1959 — Democratic Republic of Congo
  • Pseudochthonius moralesi Muchmore, 1977Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

  • Pseudochthonius mundanus Hoff, 1963 — Jamaica
  • Pseudochthonius naranjitensis (Ellingsen, 1902)Ecuador
    Ecuador
    Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

  • Pseudochthonius orthodactylus Muchmore, 1970 — Brazil
  • Pseudochthonius perreti Mahnert, 1986 — Kenya
  • Pseudochthonius pulchellus (Ellingsen, 1902) — Ecuador
  • Pseudochthonius ricardoi Mahnert, 2001 — Sao Paulo, Vale do Betari, Caverna Agua Suja, Brazil
  • Pseudochthonius simoni (Balzan, 1892) — Venezuela
  • Pseudochthonius squamosus Schawaller, 1980 — fossil: Dominican amber
    Dominican amber
    Dominican amber is amber from the Dominican Republic. Resin from the extinct species Hymenaea protera is the source of Dominican amber and probably of most amber found in the tropics....

  • Pseudochthonius strinatii Beier, 1969 — Brazil
  • Pseudochthonius thibaudi Vitali-di Castro, 1984Guadeloupe
    Guadeloupe
    Guadeloupe is an archipelago located in the Leeward Islands, in the Lesser Antilles, with a land area of 1,628 square kilometres and a population of 400,000. It is the first overseas region of France, consisting of a single overseas department. As with the other overseas departments, Guadeloupe...

  • Pseudochthonius troglobius Muchmore, 1986 — Mexico
  • Pseudochthonius tuxeni Mahnert, 1979 — Brazil
  • Pseudochthonius yucatanus Muchmore, 1977 — Mexico

  • Pseudotyrannochthonius Beier, 1930
  • Pseudotyrannochthonius australiensis Beier, 1966Australia
    Australia
    Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

  • Pseudotyrannochthonius bornemisszai Beier, 1966 — Australia
  • Pseudotyrannochthonius dentifer (Morikawa, 1970)Korea
    Korea
    Korea ) is an East Asian geographic region that is currently divided into two separate sovereign states — North Korea and South Korea. Located on the Korean Peninsula, Korea is bordered by the People's Republic of China to the northwest, Russia to the northeast, and is separated from Japan to the...

  • Pseudotyrannochthonius giganteus Beier, 1971 — Australia
  • Pseudotyrannochthonius gigas Beier, 1979 — Australia
  • Pseudotyrannochthonius gracilis Benedict & Malcolm, 1970California
    California
    California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

    , Washington
  • Pseudotyrannochthonius hamiltonsmithi Beier, 1968 — Australia
  • Pseudotyrannochthonius incognitus (R.O. Schuster, 1966) — western U.S.
  • Pseudotyrannochthonius jonesi (J.C. Chamberlin, 1962) — Australia
  • Pseudotyrannochthonius kobayashii (Morikawa, 1956)Japan
    Japan
    Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

  • Pseudotyrannochthonius kobayashii kobayashii (Morikawa, 1956) — Japan
  • Pseudotyrannochthonius kobayashii akiyoshiensis (Morikawa, 1956) — Japan
  • Pseudotyrannochthonius kobayashii dorogawaensis (Morikawa, 1960) — Japan
    • Pseudotyrannochthonius kubotai (Morkawa, 1954) — Japan
    • Pseudotyrannochthonius octospinosus Beier, 1930Chile
      Chile
      Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

    • Pseudotyrannochthonius queenslandicus Beier, 1969 — Australia
    • Pseudotyrannochthonius rossi Beier, 1964 — Chile
    • Pseudotyrannochthonius silvestrii (Ellingsen, 1905) — Chile
    • Pseudotyrannochthonius solitarius (Hoff, 1951) — Australia
    • Pseudotyrannochthonius tasmanicus Dartnall, 1970Tasmania
      Tasmania
      Tasmania is an Australian island and state. It is south of the continent, separated by Bass Strait. The state includes the island of Tasmania—the 26th largest island in the world—and the surrounding islands. The state has a population of 507,626 , of whom almost half reside in the greater Hobart...

    • Pseudotyrannochthonius typhlus Dartnall, 1970 — Tasmania
    • Pseudotyrannochthonius undecimclavatus (Morikawa, 1956) — Japan
  • Pseudotyrannochthonius undecimclavatus undecimclavatus (Morikawa, 1956) — Japan
  • Pseudotyrannochthonius undecimclavatus kishidai (Morikawa, 1960) — Japan
    • Pseudotyrannochthonius utahensis Muchmore, 1967Utah
      Utah
      Utah is a state in the Western United States. It was the 45th state to join the Union, on January 4, 1896. Approximately 80% of Utah's 2,763,885 people live along the Wasatch Front, centering on Salt Lake City. This leaves vast expanses of the state nearly uninhabited, making the population the...


  • Sathrochthoniella Beier, 1967
  • Sathrochthoniella zealandica Beier, 1967 — New Zealand

  • Sathrochthonius J.C. Chamberlin, 1962
  • Sathrochthonius crassidens Beier, 1966Australia
    Australia
    Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

  • Sathrochthonius insulanus Beier, 1976Lord Howe Island
    Lord Howe Island
    Lord Howe Island is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, directly east of mainland Port Macquarie, and about from Norfolk Island. The island is about 11 km long and between 2.8 km and 0.6 km wide with an area of...

  • Sathrochthonius kaltenbachi Beier, 1966New Caledonia
    New Caledonia
    New Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...

  • Sathrochthonius maoricus Beier, 1976 — New Zealand
  • Sathrochthonius pefauri Vitali-di Castro, 1974 — Chile
  • Sathrochthonius tuena J. C. Chamberlin, 1962 — Australia
  • Sathrochthonius tullgreni J. C. Chamberlin, 1962 — Australia
  • Sathrochthonius venezuelanus Muchmore, 1989Venezuela
    Venezuela
    Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...

  • Sathrochthonius webbi Muchmore, 1982 — Australia

  • Selachochthonius J. C. Chamberlin, 1929
  • Selachochthonius cavernicola Lawrence, 1935 — Southern Africa
  • Selachochthonius heterodentata Beier, 1955 — Southern Africa
  • Selachochthonius serratidentaus (Ellingsen, 1912) — Southern Africa

  • Spelyngochthonius Beier, 1955
  • Spelyngochthonius beieri Gardini, 1994Sardinia
    Sardinia
    Sardinia is the second-largest island in the Mediterranean Sea . It is an autonomous region of Italy, and the nearest land masses are the French island of Corsica, the Italian Peninsula, Sicily, Tunisia and the Spanish Balearic Islands.The name Sardinia is from the pre-Roman noun *sard[],...

  • Spelyngochthonius grafittii Gardini, 1994 — Sardinia
  • Spelyngochthonius heurtaultae Vachon, 1967Spain
    Spain
    Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

    , Sardinia
  • Spelyngochthonius provincialis Vachon & Heurtault-Rossi, 1964 — France
  • Spelyngochthonius sardous Beier, 1955 — Sardinia

  • Stygiochthonius Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001
  • Stygiochthonius barrancoi Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain

  • Troglochthonius Beier, 1939
  • Troglochthonius doratodactylus Helversen, 1968 — Italy
  • Troglochthonius mirabilis Beier, 1939 — Yugoslavia

  • Tyrannochthoniella Beier, 1966
  • Tyrannochthoniella zealandica Beier, 1966 — New Zealand
  • Tyrannochthoniella zealandica zealandica Beier, 1966 — New Zealand
  • Tyrannochthoniella zealandica foveauxana Beier, 1967 — New Zealand

  • Tyrannochthonius J.C. Chamberlin, 1929
  • Tyrannochthonius alabamensis Muchmore, 1996 — Alabama
  • Tyrannochthonius aladdinensis Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
  • Tyrannochthonius albidus (Beier, 1977) — Galapagos
  • Tyrannochthonius amazonicus Mahnert, 1979 — Brazil
  • Tyrannochthonius aralu Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
  • Tyrannochthonius archeri Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
  • Tyrannochthonius assimilis Hong & Tae Heung Ki, 1993 — South Korea
  • Tyrannochthonius attenuatus Muchmore, 1996 — Alabama
  • Tyrannochthonius avernicolus Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
  • Tyrannochthonius bagus Harvey, 1988 — Sumatra
  • Tyrannochthonius bahamensis Muchmore, 1984 — Bahamas
  • Tyrannochthonius barri Muchmore, 1996 — Alabama
  • Tyrannochthonius beieri Morikawa, 1963 — New Guinea, Solomons
  • Tyrannochthonius binoculatus Muchmore, 1996 — Alabama
  • Tyrannochthonius bispinosus (Beier, 1974) — India
  • Tyrannochthonius brasiliensis Mahnert, 1979 — Brazil
  • Tyrannochthonius brevimanus Beier, 1935 — central Africa
  • Tyrannochthonius brooksi Harvey, 1991 — western Australia
  • Tyrannochthonius butleri Harvey, 1991 — western Australia
  • Tyrannochthonius caecatus (Beier, 1976) — New Zealand
  • Tyrannochthonius callidus Hoff, 1959 — Jamaica
  • Tyrannochthonius cavernicola (Beier, 1976) — Lord Howe Island
  • Tyrannochthonius cavicola (Beier, 1967) — Australia
  • Tyrannochthonius centralis Beier, 1931 — Costa Rica, Ecuador
  • Tyrannochthonius chamarro J.C. Chamberlin, 1947 — Caroline Islands
  • Tyrannochthonius chamberlini Muchmore, 1996 — Alabama
  • Tyrannochthonius charon Muchmore, 1996 — Alabama
  • Tyrannochthonius chelatus Murthy & Ananthakrishnan, 1977 — India
  • Tyrannochthonius confusus Mahnert, 1986 — Kenya
  • Tyrannochthonius contractus (Tullgren, 1907) — Africa
  • Tyrannochthonius convivus Beier, 1974 — India
  • Tyrannochthonius curazavius Wagenaar-Hummelinck, 1948 — Curacao
  • Tyrannochthonius densedentatus (Beier, 1967) — New Zealand
  • Tyrannochthonius diabolus Muchmore, 1996 — Alabama
  • Tyrannochthonius ecuadoricus (Beier, 1977) — Ecuador, Peru
  • Tyrannochthonius elegans Beier, 1944 — central Africa
  • Tyrannochthonius erebicus Muchmore, 1996 — Alabama
  • Tyrannochthonius felix Muchmore, 1996 — Alabama
  • Tyrannochthonius ferox Mahnert, 1978 — Republic of Congo
  • Tyrannochthonius fiskei Muchmore, 1996 — Tennessee
  • Tyrannochthonius floridensis Malcolm & Muchmore, 1985 — Florida
  • Tyrannochthonius gezei Vachon, 1941 — Cameroon
  • Tyrannochthonius gigas Beier, 1954 — Venezuela
  • Tyrannochthonius gnomus Muchmore, 1996 — Alabama
  • Tyrannochthonius gomyi Mahnert, 1975 — Reunion
  • Tyrannochthonius grimmeti J.C. Chamberlin, 1929 — New Zealand
  • Tyrannochthonius guadeloupensis Vitali-di Castri, 1984 — Guadeloupe
  • Tyrannochthonius halopotamus Muchmore, 1996 — Alabama
  • Tyrannochthonius helenae (Beier, 1977) — St. Helena
  • Tyrannochthonius heterodentatus Beier, 1930 — India
  • Tyrannochthonius horridus (Beier, 1976) — New Zealand
  • Tyrannochthonius hypogeus Muchmore, 1996 — Kentucky
  • Tyrannochthonius imitatus Hoff, 1959 — Jamaica, Dominican Republic
  • Tyrannochthonius infernalis Muchmore, 1996 — Alabama
  • Tyrannochthonius innominatus Muchmore, 1996 — Florida
  • Tyrannochthonius innoxius Hoff, 1959 — Jamaica
  • Tyrannochthonius insulae Hoff, 1946 — Trinidad
  • Tyrannochthonius intermedius Muchmore, 1986 — Mexico
  • Tyrannochthonius irmleri Mahnert, 1979 — Brazil
  • Tyrannochthonius japonicus (Ellingsen, 1907) — Japan, Taiwan
  • Tyrannochthonius japonicus japonicus (Ellingsen, 1907) — Japan, Taiwan
  • Tyrannochthonius japonicus dogoensis Morikawa, 1954 — Japan
    • Tyrannochthonius jonesi Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
    • Tyrannochthonius kermadecensis (Beier, 1976) — New Zealand
    • Tyrannochthonius krakatau Harvey, 1988 — Indonesia
    • Tyrannochthonius laevis Beier, 1966 — Australia
    • Tyrannochthonius luxtoni (Beier, 1967) — New Zealand
    • Tyrannochthonius mahunkai Mahnert, 1978 — Republic of Congo
    • Tyrannochthonius meneghettii (Caporiacco, 1949) — central Africa
    • Tyrannochthonius meruensis Beier, 1962 — Eastern Africa
    • Tyrannochthonius migrans Mahnert, 1979 — Brazil
    • Tyrannochthonius minor Mahnert, 1979 — Brazil
    • Tyrannochthonius monodi Vachon, 1941 — Cameroon
    • Tyrannochthonius nanus (Beier, 1966) — New Guinea
    • Tyrannochthonius nergal Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
    • Tyrannochthonius noaensis Moyle, 1989 — New Zealand
    • Tyrannochthonius norfolkensis (Beier, 1976) — Norfolk Islands
    • Tyrannochthonius oahuanus Muchmore, 2000 — Hawaii, Oahu, Koolau Range, Puu Konahuanui, Pacific Islands
    • Tyrannochthonius orpheus Muchmore, 1996 — Alabama
    • Tyrannochthonius osiris Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
    • Tyrannochthonius ovatus Vitali-di Castro, 1984 — Martinique
    • Tyrannochthonius pachythorax Redikorzev, 1938 — Vietnam
    • Tyrannochthonius palauanus Beier, 1957 — Caroline Islands
    • Tyrannochthonius pallidus Muchmore, 1973 — Mexico
    • Tyrannochthonius parvus Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
    • Tyrannochthonius pecki Muchmore, 1996 — Alabama
    • Tyrannochthonius perpusillus Beier, 1951 — Vietnam
    • Tyrannochthonius philippinus (Beier, 1966) — Philippines
    • Tyrannochthonius pholeter Muchmore, 1996 — Alabama
    • Tyrannochthonius pluto Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
    • Tyrannochthonius procerus Mahnert, 1978 — Republic of Congo
    • Tyrannochthonius proximus Hoff, 1959 — Jamaica, Dominican Republic
    • Tyrannochthonius pugnax Mahnert, 1978 — Republic of Congo
    • Tyrannochthonius pupukeanus Muchmore, 1983 — Hawai'i
    • Tyrannochthonius pusillimus Beier, 1951 — Vietnam
    • Tyrannochthonius pusillus Beier, 1955 — Peru
    • Tyrannochthonius queenslandicus (Beier, 1969) — Australia
    • Tyrannochthonius rahmi Beier, 1976 — Nepal, Bhutan
    • Tyrannochthonius rex Harvey, 1989 — Australia
    • Tyrannochthonius riberai Mahnert, 1984 — Peru
    • Tyrannochthonius robustus Beier, 1951 — Vietnam
    • Tyrannochthonius rotundimanus Mahnert, 1985 — Brazil
    • Tyrannochthonius satan Muchmore, 1996 — Alabama
    • Tyrannochthonius semidentatus (Redikorzev, 1924) — Kenya
    • Tyrannochthonius semihorridus (Beier, 1969) — Australia
    • Tyrannochthonius setiger Mahnert, 1997 — Tenerife, Cueva del Sobrado, Canary Islands
    • Tyrannochthonius sheltae Muchmore, 1996 — Alabama
    • Tyrannochthonius similidentatus Sato, 1984 — Japan
    • Tyrannochthonius simillimus Beier, 1951 — Cambodia
    • Tyrannochthonius simulans Mahnert, 1986 — Kenya
    • Tyrannochthonius skeletonis Muchmore, 1996 — Alabama
    • Tyrannochthonius sokolovi (Redikorzev, 1924) — central Africa
    • Tyrannochthonius sparsedentatus Beier, 1959 — Democratic Republic of Congo
    • Tyrannochthonius spinatus Hong, in Yong Hong, Tae Heung Kim & Won Koo Le 1996 — South Korea
    • Tyrannochthonius steevesi Muchmore, 1996 — Tennessee
    • Tyrannochthonius stonei Muchmore, 1989 — Hawai'i
    • Tyrannochthonius strinatii (Beier, 1974) — Guatemala
    • Tyrannochthonius stygius Muchmore, 1996 — Alabama
    • Tyrannochthonius superstes Mahnert, 1986 — Canary Islands
    • Tyrannochthonius suppressalis Hong, in Yong Hong, Tae Heung Kim & Won Koo Le 1996 — South Korea
    • Tyrannochthonius swiftae Muchmore, 1993 — Hawai'i
    • Tyrannochthonius tartarus Muchmore, 1996 — Alabama
    • Tyrannochthonius tekauriensis Moyle, 1989 — New Zealand
    • Tyrannochthonius tenuis Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
    • Tyrannochthonius terribilis (With, 1906) — southeast Asia
  • Tyrannochthonius terribilis terribilis (With, 1906) — southeast Asia
  • Tyrannochthonius terribilis malaccensis Beier, 1952 — Malaysia
    • Tyrannochthonius tlilapanensis Muchmore, 1986 — Mexico
    • Tyrannochthonius torodei Muchmore, 1996 — Alabama
    • Tyrannochthonius troglobius Muchmore, 1969 — Mexico
    • Tyrannochthonius troglodytes Muchmore, 1986 — Texas
    • Tyrannochthonius troglophilus (Beier, 1968) — New Caledonia
    • Tyrannochthonius vampirorum Muchmore, 1986 — Mexico
    • Tyrannochthonius volcancillo Muchmore, 1986 — Mexico
    • Tyrannochthonius volcanus Muchmore, 1977 — Mexico
    • Tyrannochthonius wittei Beier, 1955
    • Tyrannochthonius wlassicsi (Daday, 1897) — New Guinea
    • Tyrannochthonius zicsii Mahnert, 1978 — Republic of Congo
    • Tyrannochthonius zonatus (Beier, 1964) — New Caledonia

  • Vulcanochthonius Muchmore, 2000
  • Vulcanochthonius aa Muchmore, 2000 — Hawai'i, Manuka Natural Area Reserve, Visor Cave, Pacific Islands
  • Vulcanochthonius howarthi (Muchmore, 1979) — Hawai'i
  • Vulcanochthonius pohakuloae Muchmore, 2000 — Hawai'i, Mauna Kea, Pohakuloa Training Area, Bobcat Trail, Pacific Islands
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