All Topics  
Jackdaw

 
Jackdaw

   Email Print
   Bookmark   Link






 

Jackdaw



 
 
The Jackdaw (Corvus monedula), sometimes known as the Eurasian Jackdaw, European Jackdaw, Western Jackdaw, or formerly simply the daw, is one of the smallest species (34–39 cm in length) in the genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 of crow
Crow

The true crows are large passerine birds that form the genus Corvus in the family Corvidae. Ranging in size from the relatively small dove-sized jackdaws to the Common Raven of the Holarctic region and Thick-billed Raven of the highlands of Ethiopia, the 40 or so members of this genus occur on all temperate continents and several offsh...
s and raven
Raven

Raven is the common name given to several larger-bodied members of the genus Corvus —but in Europe and North America the Common Raven is normally implied....
s. It is a black-plumaged bird with grey nape and distinctive white irises. Like all corvids, it is omnivorous. It is found across Europe, western Asia and North Africa.






Discussion
Ask a question about 'Jackdaw'
Start a new discussion about 'Jackdaw'
Answer questions from other users
Full Discussion Forum



Encyclopedia


The Jackdaw (Corvus monedula), sometimes known as the Eurasian Jackdaw, European Jackdaw, Western Jackdaw, or formerly simply the daw, is one of the smallest species (34–39 cm in length) in the genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 of crow
Crow

The true crows are large passerine birds that form the genus Corvus in the family Corvidae. Ranging in size from the relatively small dove-sized jackdaws to the Common Raven of the Holarctic region and Thick-billed Raven of the highlands of Ethiopia, the 40 or so members of this genus occur on all temperate continents and several offsh...
s and raven
Raven

Raven is the common name given to several larger-bodied members of the genus Corvus —but in Europe and North America the Common Raven is normally implied....
s. It is a black-plumaged bird with grey nape and distinctive white irises. Like all corvids, it is omnivorous. It is found across Europe, western Asia and North Africa. Four subspecies are currently recognised.

Taxonomy


The Jackdaw was one of the many species originally described by Linnaeus
Carolus Linnaeus

Carl Linnaeus was a Sweden botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern alpha taxonomy, and is also considered one of the fathers of modern ecology....
 in his 18th century work, Systema Naturae
Systema Naturae

The book Systema Naturae was one of the major works of the Sweden botanist, zoologist and physician Carolus Linnaeus. Its full title is Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis or translated: "System of nature through the three kingdoms of...
, and it still bears its original name of Corvus monedula. The species name monedula is the Latin
Latin

Latin is an Italic language, historically spoken in Latium and Ancient Rome. Through the Military history of the Roman Empire, Latin spread throughout the Mediterranean and a large part of Europe....
 for jackdaw.

The common name jackdaw first appears in the 16th century, and is a compound of the forename Jack used in animal names to signify a small form (e.g. jack-snipe) and the native English word daw. Formerly jackdaws were simply called daws. Claims that the metallic chyak call is the origin of the jack part of the common name are not supported by the Oxford English Dictionary
Oxford English Dictionary

The Oxford English Dictionary , published by the Oxford University Press , is a comprehensive dictionary of the English language. Two fully-bound print editions of the OED have been published under its current name, in 1928 and 1989; as of December 2008 the dictionary's current editors have completed a quarter of the third edition....
.

Daw is first attested in the 15th century, which the Oxford English Dictionary conjectures to be derived from an unattested Old English dawe, citing cognate
Cognate

Cognates in linguistics are words that have a common etymology origin.An example of cognates within the same language would be English shirt vs....
s in Old High German
Old High German

The term Old High German refers to the earliest stage of the German language and it conventionally covers the period from around 500 to 1050. Coherent written texts do not appear until the second half of the 8th century, and some treat the period before 750 as 'prehistoric' and date the start of Old High German proper to 750 for this reason...
 tâha, Middle High German
Middle High German

Middle High German , abbreviated MHG , is the term used for the period in the history of the German language between 1050 and 1350. It is preceded by Old High German and followed by Early New High German....
 tâhe and modern German dialect dähi, däche, dacha.

The original Old English name was ceo (pronounced with initial ch). Although now reserved for corvids of the genus Pyrrhocorax, the word chough
Chough

The Red-billed Chough or Chough , Pyrrhocorax pyrrhocorax, is a bird in the crow family; it is one of only two species in the genus Pyrrhocorax....
 originally referred to the jackdaw; Chaucer
Geoffrey Chaucer

Geoffrey Chaucer was an English author, poet, philosopher, Bureaucracy, Noble court and diplomat. Although he wrote many works, he is best remembered for his unfinished frame narrative The Canterbury Tales....
 and Shakespeare
William Shakespeare

William Shakespeare was an English people poet and playwright, widely regarded as the greatest writer in the English language and the world's preeminent dramatist....
 sometimes used this word.

English dialect names are numerous. Scottish and north England dialect has had ka or kae since the 14th century. The midlands form of this was co or coo. Caddow is potentially a compound of ka and dow, a variant of daw. Other dialect or obsolete names include caddesse, cawdaw, caddy, chauk, college-bird (from dialect college = cathedral), jackerdaw, jacko, ka-wattie, chimney-sweep bird, from their nesting propensities, and sea-crow, from their frequenting coasts. It was also frequently known quasi-nominally as Jack.

An archaic collective noun for a group of jackdaws is a "clattering". Another term used is "train," however, in practice, most people use the more generic term "flock".

Subspecies


There are four recognised subspecies. All European subspecies intergrade where their populations meet. C. m. monedula integrates into C. m. soemmerringii with the transition zone running from Finland south across the Baltic, east Poland to Romania and Croatia.

  • C. m. monedula (Linnaeus, 1758), the nominate subspecies, breeds in south-east Norway, southern Sweden and northern and eastern Denmark, with occasional wintering birds in England and France. It has a pale nape and side of the neck, dark throat, and a light grey partial collar of variable extent.
  • C. m. spermologus (Vieillot, 1817) occurs in western and central Europe, and winters in the Canary Islands and Corsica. It is darker in colour and lacks the whitish border at the base of the grey collar.
  • C. m. soemmerringii (Fischer, 1811) is found in north-eastern Europe, and north and central Asia, from the former Soviet Union to Lake Baikal
    Lake Baikal

    Lake Baikal is in southern Siberia in Russia, located between Irkutsk Oblast to the northwest and the Buryatia to the southeast, near the city of Irkutsk....
     and north-west Mongolia and south to Turkey, Israel and the eastern Himalayas. It winters in Iran and northwestern India (Kashmir
    Kashmir

    Kashmir is the northwestern region of the Indian subcontinent. Until the mid-19th century, the term "Kashmir" referred only to the valley lying between the Great Himalayas and the Pir Panjal range; since then, it has been used for a larger area that today includes the Indian administerd state of Jammu and Kashmir consisting of the Kashmir...
    ). It is distinguished by its paler nape and side of the neck creating a contrasting black crown, and lighter grey partial collar.
  • C. m. cirtensis (Rothschild and Hartert, 1912) is found in Morocco
    Morocco

    Morocco , officially the Kingdom of Morocco , is a country located in North Africa with a population of nearly 34 million and an area just under 447,000 km2....
     and Algeria
    Algeria

    Algeria , officially the People's Democratic Republic of Algeria, is a country located in North Africa. It is the largest country of the Mediterranean sea, second largest in the Arab World, and the second largest on the African continent and the eleventh-largest country in the world in terms of land area....
     in North Africa. The plumage is duller and more uniform dark grey, with the paler nape less distinct.


Description


Measuring 34–39 cm (14-15 in), the jackdaw is the smallest species in the genus Corvus. Most of the plumage is black or greyish black except for the cheeks, nape and neck, which are light grey to greyish silver. The iris of adults is greyish white or silvery white, the only member of the genus outside of the Australasia
Australasia

Australasia is a region of Oceania: New Zealand, Australia, Papua New Guinea, and neighbouring islands in the Pacific Ocean. The term was coined by Charles de Brosses in Histoire des navigations aux terres australes ....
n region to have this feature. The iris of juvenile jackdaws is light blue.

In flight, jackdaws are separable from other corvids by their smaller size, faster and deepers wingbeats and proportionately narrower and less fingered wings. They also have a shorter, thicker neck, a much shorter bill and frequently fly in tighter flocks. Underwing is uniformly grey, unlike chough
Chough

The Red-billed Chough or Chough , Pyrrhocorax pyrrhocorax, is a bird in the crow family; it is one of only two species in the genus Pyrrhocorax....
s.

On the ground, jackdaws strut about briskly and have an upright posture.

Sexes and ages are alike.

Voice


Jackdaws are voluble birds. The call, frequently given in flight, is a metallic and somewhat squeaky, "chyak-chyak" or "kak-kak". Perched birds often chatter together, and before settling for the night large roosting flocks make a cackling noise. Jackdaws also have a hoarse, drawn-out alarm-call.

Distribution and habitat


Jackdaws are resident over a large area stretching from North West Africa through virtually all of Europe, including the British Isles and southern Scandinavia, westwards through central Asia to the eastern Himalayas and Lake Baikal
Lake Baikal

Lake Baikal is in southern Siberia in Russia, located between Irkutsk Oblast to the northwest and the Buryatia to the southeast, near the city of Irkutsk....
. They are resident throughout Turkey, the Caucasus, Iran, Iraq, Afghanistan, Pakistan and north-west India.

The species has a large range, with an estimated global extent of between 1,000,000 and 10,000,000 km˛. It has a large global population, with an estimated 10 to 29 million individuals in Europe..

Jackdaws are mostly resident, but the northern and eastern populations are more migratory. Their range expands northwards into Russia to Siberia during summer, and retracts in winter. They are winter vagrants to Lebanon, first recorded there in 1962. In Syria they are winter vagrants and rare residents with some confirmed breeding. The soemmerringii race occurs in south-central Siberia and extreme northwest China and is accidental to Hokkaido
Hokkaido

, formerly known as Ezo, Yezo, Yeso, or Yesso, is Japan's second largest island and the largest, northernmost of its 47 prefectures of Japan....
, Japan.

A small number of Jackdaws reached the northwest of North America in the 1980s, presumedly ship-assisted, and have been found from Atlantic Canada to Pennsylvania. They have also occurred as vagrants in Canada, the Faroe Islands
Faroe Islands

The Faroe Islands or Faeroe Islands or simply Faroe or Faeroes are an island group situated between the Norwegian Sea and the North Atlantic Ocean, approximately half way between Scotland and Iceland....
, Gibraltar
Gibraltar

Gibraltar is a British overseas territory located near the southernmost tip of the Iberian Peninsula overlooking the Strait of Gibraltar. The territory shares a border with Spain to the north....
, Iceland, Mauritania
Mauritania

Mauritania , officially the Islamic Republic of Mauritania, is a country in northwest Africa. It is bordered by the Atlantic Ocean on the west, by Senegal on the southwest, by Mali on the east and southeast, by Algeria on the northeast, and by the Morocco-controlled Western Sahara on the northwest....
 and Saint Pierre and Miquelon. Jackdaws are regionally extinct in Malta
Malta

Malta , officially the Republic of Malta , is a densely populated developed country European microstates microstate in the European Union....
 and Tunisia
Tunisia

Tunisia , officially the Tunisian Republic , is a country located in North Africa. It is bordered by Algeria to the west and Libya to the southeast....
. The Jackdaw is reported to have once occurred in Egypt.

They inhabit wooded steppe
Steppe

In physical geography, a steppe , pronounced , is a grassland plain without trees . The prairie can be considered a steppe. It may be semi-desert, or covered with Poaceae or shrubs or both, depending on the season and latitude....
s, woodland, cultivated land, pasture, coastal cliffs and villages and towns.

Behavior


Jackdaws are highly gregarious and are generally seen in small to large flocks, though males and females pair-bond for life and pairs stay together within flocks. Flock sizes increase in autumn and large flocks group together at dusk for communal roosting. They become sexually mature in the first breeding season, and there is little evidence for divorce or extra pair coupling in jackdaws, even after multiple instances of reproductive failure.

Jackdaws frequently congregate with Hooded Crow
Hooded Crow

The Hooded Crow is a Eurasian bird species in the crow genus. Widely distributed, it is also known locally as Scotch Crow, Danish Crow, and Grey Crow in Ireland, which is what its Welsh name, :cy:Br?n Lwyd, translates as....
 (Corvus cornix), and during migration often accompany Rook
Rook (bird)

The Rook is a member of the Corvidae family in the passerine order of birds. Named by Linnaeus in 1758, the species name frugilegus is Latin for "food-gathering"....
s (C. frugilegus).

Like magpies, jackdaws are known to steal shiny objects such as jewelry to hoard in nests. John Gay
John Gay

John Gay was an English people poet and dramatist. He is best remembered for The Beggar's Opera , set to music by Johann Christoph Pepusch....
 in his Beggar's Opera notes that "A covetous fellow, like a jackdaw, steals what he was never made to enjoy, for the sake of hiding it" and in Tobias Smollett
Tobias Smollett

Tobias George Smollett was a Scotland poet and author. He was best known for his picaresque novels, such as The Adventures of Roderick Random and The Adventures of Peregrine Pickle , which influenced later novelists such as Charles Dickens....
's The Expedition of Humphry Clinker a scathing character assassination by Mr. Bramble runs "He is ungracious as a hog, greedy as a vulture, and thievish as a jackdaw."

Feeding


The jackdaw mostly takes food from the ground but does take some food in trees.

In terms of animal food, jackdaws tend to feed upon small invertebrates found above ground between 2 and 18 mm in length, including imagines, larvae and pupae of Curculionidae
Curculionidae

Curculionidae is the family of the "true" weevils . With over 60,000 species described worldwide, it is the largest of the beetle families.They are recognized by their distinctive long snout and geniculate antennae with small clubs; beyond that curculionids have considerable diversity of form and size, with adult lengths ranging from 1&nbs...
, Coleoptera, Diptera and Lepidoptera
Lepidoptera

Lepidoptera is an order of insect that includes moths and butterfly. It is one of the most speciose orders in the class Insecta, encompassing moths and the three superfamilies of butterfly, skipper , and Hedylidae....
. Snails, spiders and some other insects also make up part of their animal diet. Unlike rook
Rook

Rook may refer to:...
s and carrion crow
Carrion Crow

The Carrion Crow is a member of the passerine order of birds and the crow family which is native to western Europe and eastern Asia....
s, jackdaws do not generally feed on carrion, though they will eat stranded fish on the shore. The vegetal diet of jackdaws consists of farm grains (barley, wheat and oats), seeds of weeds, elderberries, acorns and various cultivated fruits. Jackdaws also take scraps of human food in towns, and will more readily take food from bird tables than other Corvus species.

Jackdaws employ various feeding methods, such as jumping, pecking, clod-turning and scattering, probing the soil, and rarely digging. Flies around cow pats are caught by jumping from the ground or at times by dropping vertically from a few metres above onto the cow pat. Earthworms are not usually extracted from the ground by jackdaws but are eaten from freshly plough soil.

Jackdaws practice active food sharing, where the initiative for the transfer lies with the donor, with a number of individuals, regardless of sex and kinship. They also share more of a preferred food than a less preferred food.

Infant jackdaws are altricial and thus are completely dependent on being fed by their parents until they fledge.

Corvus Monedula

Breeding


Jackdaws usually nest in colonies with monogamous pairs collaborating to locate a nest site which they then defend from other pairs and predators most of the year.

Jackdaws nest in cavities of trees, cliffs or ruined, and sometimes inhabited, buildings, often in chimneys, and even in dense conifers. They are famous for using church steeples for nesting, a fact reported in verse by William Cowper
William Cowper

William Cowper was an English poet and hymnodist. One of the most popular poets of his time, Cowper changed the direction of 18th century nature poetry by writing of everyday life and scenes of the English countryside....
A great frequenter of the church,
Where, bishoplike, he finds a perch,
And dormitory too.


Nests are usually constructed by a mated pair blocking up the crevice by dropping sticks into it; the nest is then built atop the platform formed. This behaviour has led to blocked chimneys and even nests, with the jackdaw present, crashing down into fireplaces. Nest platforms can attain great size - John Mason Neale notes that a "Clerk was allowed by the Churchwarden to have for his own use all that the caddows had brought into the Tower: and he took home, at one time, two cart-loads of good firewood, besides a great quantity of rubbish which he threw away."

Gilbert White, in his popular book The Natural History of Selborne
The Natural History and Antiquities of Selborne

The Natural History and Antiquities of Selborne, or just The Natural History of Selborne was a book by pioneering England natural history and Ornithology Gilbert White first published in 1789....
, notes that jackdaws used to nest in crevices beneath the lintels of Stonehenge
Stonehenge

Stonehenge is a prehistoric monument located in the England county of Wiltshire, about west of Amesbury and north of Salisbury. One of the most famous sites in the world, Stonehenge is composed of Earthworks surrounding a circular setting of large standing stones and sits at the centre of the densest complex of Neolithic and Bronze Age mon...
, and describes a curious example of jackdaws using rabbit burrows for nest sites.

Nests are lined with hair, rags, bark, soil, and many other materials. Jackdaws nest in colonies and often close to rooks
Rook (bird)

The Rook is a member of the Corvidae family in the passerine order of birds. Named by Linnaeus in 1758, the species name frugilegus is Latin for "food-gathering"....
. The egg
Egg (biology)

In most birds and reptiles, an egg is the zygote, resulting from fertilization of the ovum. To enable incubation the egg is usually kept within a favourable temperature range as it nourishes and protects the growing embryo....
s are smooth, glossy pale blue speckled with dark brown, measuring approximately 36 x 26 mm. Clutches of normally 4-5 eggs, are incubated by the female for 17-18 days and fledge after 28-35 days, when they are fed by both parents.

Jackdaws hatch asynchronously and incubation begins before clutch completion, often leading to the death of the last-hatched young. The young which die in the nest do so quickly which minimises parental investment, and hence the brood size comes to fit the available food supply.

Social behavior


The jackdaw is a highly sociable species outside of the breeding season, occurring in flocks that can contain hundreds of birds.

Konrad Lorenz
Konrad Lorenz

Konrad Zacharias Lorenz was an Austrian zoology, animal psychology, ornithologist and Nobel Prize winner. He is often regarded as one of the founders of modern ethology, developing an approach that began with an earlier generation, including his teacher Oskar Heinroth....
 studied the complex social interactions that occur in groups of jackdaws and published his detailed observations of their social behavior in his book King Solomon's Ring
King Solomon's Ring (nonfiction)

King Solomon's Ring is a zoology book for the general audience, written by the Austrian scientist Konrad Lorenz in 1949. The first English-language edition appeared in 1952....
. To study jackdaws, Lorenz put coloured rings on the legs of the jackdaws that lived around his house in Altenberg
Sankt Andrä-Wördern

Sankt Andr?-W?rdern is a municipality in the district of Tulln in Lower Austria, Austria....
, Austria
Austria

Austria , officially the Republic of Austria , is a landlocked country in Central Europe. It borders both Germany and the Czech Republic to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the west....
 for identification, and he caged them in the winter because of their annual migration
Bird migration

Bird migration refers to the regular seasonal journeys undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather....
 away from Austria
Austria

Austria , officially the Republic of Austria , is a landlocked country in Central Europe. It borders both Germany and the Czech Republic to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the west....
. His book describes his observations on jackdaws' hierarchical group structure
Dominance hierarchy

A dominance hierarchy is the organization of individuals in a group that occurs when competition of resources lead to aggression. Schjelderup-Ebbe, who studied the often-cited example of the pecking order in chickens, found that such social structures lead to more stable flocks in which aggression was reduced among individuals....
, in which the higher-ranking birds are dominant over lower ranked birds. The book also records his observations on jackdaws' strong male–female bonding; he noted that each bird of a pair both have about the same rank in the hierarchy, and that a low-ranked female jackdaw rocketed up the jackdaw social ladder when she became the mate of a high-ranking male.

Jackdaws have been observed sharing food and objects. The active giving of food is rare in primates, and in birds is found mainly in the context of parental care and courtship. Jackdaws show much higher levels of active giving than documented for chimpanzee
Chimpanzee

Chimpanzee, sometimes colloquially known as a chimp, is the common name for the two Extant taxon species of ape in the genus Pan where the Congo River forms the boundary between the native habitat of the two species:...
s. The function of this behaviour is not fully understood, although it has been found to be compatible with hypotheses of mutualism, reciprocity and harassment avoidance.

Occasionally the flock makes 'mercy killings', in which a sick or injured bird is mobbed until it is killed.

Cultural depictions and folklore


  • In some cultures, a jackdaw on the roof is said to predict a new arrival; alternatively, a jackdaw settling on the roof of a house or flying down a chimney is an omen of death and coming across one is considered a bad omen. In the 11th century William of Malmesbury
    William of Malmesbury

    William of Malmesbury , English historians in the Middle Ages, was born about the year 1080/1095, in Wiltshire. His father was Normans and his mother English....
     records the story of a woman who upon hearing a jackdaw chattering "more loudly than usual", grew pale and became fearful of suffering a "dreadful calamity", and that "while yet speaking, the messenger of her misfortunes arrived."
  • A jackdaw standing on the vanes of a cathedral tower is meant to prognosticate rain. Czech superstition formerly held that if jackdaws are seen quarrelling, war will follow, and that jackdaws will not build nests at Sázava
    Sázava (Benešov District)

    S?zava is a town in the Central Bohemian Region of the Czech Republic. It has around 3,800 inhabitants.Villages Belokozly, Cern? Budy, Cerenice and Dojetrice are administrative parts of S?zava....
     having been banished by Saint Procopius
    Procopius of Sázava

    Saint Procopius of S?zava was a Czechs saint. He studied at Prague where he was also ordination. He was a Canon and a hermit and then became the founding abbot of S?zava near Prague....
    .
  • Ancient Greek authors tell how a jackdaw, being a social creature, may be caught with a dish of oil which it falls into while looking at its own reflection.
  • Pliny
    Pliny the Elder

    Gaius Plinius Secundus , better known as Pliny the Elder, was an ancient author, naturalist or natural philosopher and naval and military commander of some importance who wrote Natural History ....
     notes how the Thessalians, Illyrians
    Illyrians

    Illyrians has come to refer to a broad, ill-defined "Indo-European languages" group of peoples who inhabited the western Balkans and even possibly Messapia in Southern Italy ....
     and Lemnians cherished jackdaws for destroying grasshoppers' eggs. The Veneti
    Veneti

    Veneti may refer to:*Veneti , a Celtic tribe who once lived in what is now Brittany, France*Adriatic Veneti, a bygone people of northeastern Italy who spoke an unclassified Indo-European language...
     are fabled to have bribed the jackdaws to spare their crops.
  • The Ingoldsby Legends
    The Ingoldsby Legends

    The Ingoldsby Legends are a collection of myths, legends, ghost stories and poetry supposedly written by Thomas Ingoldsby of Tappington Manor, actually a pen-name of Richard Harris Barham....
     (1837) contains a poem named The Jackdaw of Rheims, which is about a jackdaw who steals a cardinal's ring and is made a saint.
  • An ancient Greek and Roman adage runs "The swans will sing when the jackdaws are silent" meaning that educated or wise people will speak after foolish prattlers finally run out of talk.
  • The sentence "Jackdaws love my big sphinx of quartz" is a commonly used example of a pangram
    Pangram

    A pangram , or holoalphabetic sentence, is a sentence using every letter of the alphabet at least once. Pangrams are used to display typefaces and test equipment....
    , (i.e. a sentence that contains all 26 letters of the English alphabet
    English alphabet

    The modern English alphabet is a Latin-based alphabet consisting of 26 letters, like in the Basic modern Latin alphabet:The exact shape of printed letters varies depending on the typeface....
    ), while the sentence itself is only 31 letters long.


External links

  • on the Internet Bird Collection


Cited texts


Further reading


Identification


Harrop, Andrew (2000) Identification of Jackdaw forms in northwestern Europe Birding World
Birding World

Birding World is a monthly birdwatching magazine published in the United Kingdom. It is the magazine of the Bird Information Service, based at Cley next the Sea, Norfolk....
 vol. 13, no. 7 pages 290-295