Homo erectus is an extinct
speciesIn biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...
of hominid that lived from the end of the Pliocene epoch to the later Pleistocene, about . The species originated in
AfricaAfrica is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...
and spread as far as
IndiaIndia , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...
,
ChinaChinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...
and
JavaJava is an island of Indonesia. With a population of 135 million , it is the world's most populous island, and one of the most densely populated regions in the world. It is home to 60% of Indonesia's population. The Indonesian capital city, Jakarta, is in west Java...
. There is still disagreement on the subject of the classification, ancestry, and progeny of H. erectus, with two major alternative
hypothesesA hypothesis is a proposed explanation for a phenomenon. The term derives from the Greek, ὑποτιθέναι – hypotithenai meaning "to put under" or "to suppose". For a hypothesis to be put forward as a scientific hypothesis, the scientific method requires that one can test it...
: erectus may be another name for
Homo ergasterHomo ergaster is an extinct chronospecies of Homo that lived in eastern and southern Africa during the early Pleistocene, about 2.5–1.7 million years ago.There is still disagreement on the subject of the classification, ancestry, and progeny of H...
, and therefore the direct ancestor of later hominids such as
Homo heidelbergensisHomo heidelbergensis is an extinct species of the genus Homo which may be the direct ancestor of both Homo neanderthalensis in Europe and Homo sapiens. The best evidence found for these hominins date between 600,000 and 400,000 years ago. H...
, Homo neanderthalensis, and Homo sapiens; or it may be an Asian species distinct from African ergaster.
Origin
The first theory is that H. erectus migrated from
AfricaAfrica is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...
during the
Early PleistoceneCalabrian is a subdivision of the Pleistocene Epoch of the Geologic time scale. ~1.8 Ma.—781,000 years ago ± 5,000 years, a period of ~.The end of the stage is defined by the last magnetic pole reversal and plunge in to an ice age and global drying possibly colder and drier than the late Miocene ...
, possibly as a result of the operation of the
Saharan pumpThe Sahara pump theory is a hypothesis that explains how flora and fauna migrated between Eurasia and Africa via a Levantine land bridge. The theory observes that extended periods of abundant rainfall lasting many thousands of years in Africa are associated with a "wet Sahara" phase, during which...
, around 2.0 million years ago, and dispersed throughout much of the
Old WorldThe Old World consists of those parts of the world known to classical antiquity and the European Middle Ages. It is used in the context of, and contrast with, the "New World" ....
. Fossilized remains have been found in Africa (e.g.,
Lake TurkanaLake Turkana , formerly known as Lake Rudolf, is a lake in the Great Rift Valley in Kenya, with its far northern end crossing into Ethiopia. It is the world's largest permanent desert lake and the world's largest alkaline lake...
and
Olduvai GorgeThe Olduvai Gorge is a steep-sided ravine in the Great Rift Valley that stretches through eastern Africa. It is in the eastern Serengeti Plains in northern Tanzania and is about long. It is located 45 km from the Laetoli archaeological site...
),
EuropeEurope is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...
(
GeorgiaGeorgia is a sovereign state in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded to the west by the Black Sea, to the north by Russia, to the southwest by Turkey, to the south by Armenia, and to the southeast by Azerbaijan. The capital of...
and
SpainSpain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...
),
IndonesiaIndonesia , officially the Republic of Indonesia , is a country in Southeast Asia and Oceania. Indonesia is an archipelago comprising approximately 13,000 islands. It has 33 provinces with over 238 million people, and is the world's fourth most populous country. Indonesia is a republic, with an...
(e.g., Sangiran and Trinil),
VietnamVietnam – sometimes spelled Viet Nam , officially the Socialist Republic of Vietnam – is the easternmost country on the Indochina Peninsula in Southeast Asia. It is bordered by China to the north, Laos to the northwest, Cambodia to the southwest, and the South China Sea –...
,
ChinaChinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...
(e.g.,
Shaanxi' is a province in the central part of Mainland China, and it includes portions of the Loess Plateau on the middle reaches of the Yellow River in addition to the Qinling Mountains across the southern part of this province...
) and
IndiaIndia , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...
.
The second theory is H. erectus evolved in
AsiaAsia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...
and then migrated to Africa. The species occupied a West Asian site called
DmanisiDmanisi is a townlet and archaeological site in Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera.- History :...
, in Georgia, from 1.85 million to 1.77 million years ago, at the same time or slightly before the earliest evidence in Africa. Excavations found 73 stone tools for cutting and chopping and 34 bone fragments from unidentified
creatureA creature is a created thing, being or any product of creative action. The term is used most frequently to refer to biological organisms. The term is derived from the traditional belief that all things are created by God....
s.
Discovery and representative fossils
The Dutch anatomist
Eugene DuboisMarie Eugène François Thomas Dubois was a Dutch paleoanthropologist. He earned worldwide fame for his discovery of Pithecanthropus erectus , or 'Java Man'...
, who was fascinated especially by
DarwinCharles Robert Darwin FRS was an English naturalist. He established that all species of life have descended over time from common ancestry, and proposed the scientific theory that this branching pattern of evolution resulted from a process that he called natural selection.He published his theory...
's theory of evolution as applied to man, set out to Asia (the place accepted then, despite Darwin, as the cradle of human evolution), to find a human ancestor in 1886. In 1891, his team discovered a human fossil on the island of Java, Indonesia; he described the species as Pithecanthropus erectus (from the Greek πίθηκος, "ape", and ἄνθρωπος, "man"), based on a calotte (skullcap) and a femur like that of H. sapiens found from the bank of the Solo River at
TrinilTrinil is a palaeoanthropological site on the banks of the Bengawan Solo River in Ngawi Regency, East Java Province, Indonesia. It was at this site in 1891 that the Dutch anatomist Eugène Dubois discovered the first early hominid remains to be found outside of Europe: the famous "Java Man" specimen....
, in
East JavaEast Java is a province of Indonesia. It is located on the eastern part of the island of Java and includes neighboring Madura and islands to its east and to its north East Java is a province of Indonesia. It is located on the eastern part of the island of Java and includes neighboring Madura and...
. (This species is now regarded as Homo erectus.)
The find became known as
Java ManJava Man is the name given to fossils discovered in 1891 at Trinil - Ngawi Regency on the banks of the Solo River in East Java, Indonesia, one of the first known specimens of Homo erectus...
. Thanks to Canadian anatomist
Davidson BlackDavidson Black, FRS was a Canadian paleoanthropologist, best known for his naming of Sinanthropus pekinensis . He was Chairman of the Geological Survey of China and a Fellow of the Royal Society...
's (1921) initial description of a lower molar, which was dubbed Sinanthropus pekinensis, however, most of the early and spectacular discoveries of this taxon took place at
ZhoukoudianZhoukoudian or Choukoutien is a cave system in Beijing, China. It has yielded many archaeological discoveries, including one of the first specimens of Homo erectus, dubbed Peking Man, and a fine assemblage of bones of the gigantic hyena Pachycrocuta brevirostris...
in
ChinaChinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...
. German anatomist
Franz Weidenreich-External references:*...
provided much of the detailed description of this material in several monographs published in the journal Palaeontologica Sinica (Series D).
Nearly all of the original specimens were lost during
World War IIWorld War II, or the Second World War , was a global conflict lasting from 1939 to 1945, involving most of the world's nations—including all of the great powers—eventually forming two opposing military alliances: the Allies and the Axis...
; however, authentic Weidenreichian casts do exist at the
American Museum of Natural HistoryThe American Museum of Natural History , located on the Upper West Side of Manhattan in New York City, United States, is one of the largest and most celebrated museums in the world...
in
New YorkNew York is a state in the Northeastern region of the United States. It is the nation's third most populous state. New York is bordered by New Jersey and Pennsylvania to the south, and by Connecticut, Massachusetts and Vermont to the east...
and at the
Institute of Vertebrate Paleontology and PaleoanthropologyThe Institute of Vertebrate Paleontology and Paleoanthropology of China is a prominent research institution and collections repository for fossils, including many dinosaur and pterosaurand cat poo specimens...
in
BeijingBeijing , also known as Peking , is the capital of the People's Republic of China and one of the most populous cities in the world, with a population of 19,612,368 as of 2010. The city is the country's political, cultural, and educational center, and home to the headquarters for most of China's...
, and are considered to be reliable evidence.
Throughout much of the 20th century, anthropologists debated the role of H. erectus in
human evolutionHuman evolution refers to the evolutionary history of the genus Homo, including the emergence of Homo sapiens as a distinct species and as a unique category of hominids and mammals...
. Early in the century, however, due to discoveries on Java and at Zhoukoudian, it was believed that modern humans first evolved in Asia. A few naturalists (Charles Darwin most prominent among them) predicted that humans' earliest ancestors were African: he pointed out that chimpanzees and gorillas, who are human relatives, live only in Africa.
From 1950s to 1970s, however, numerous fossil finds from East Africa yielded evidence that the oldest
homininsHomininae is a subfamily of Hominidae, which includes humans, gorillas and chimpanzees, and some extinct relatives; it comprises all those hominids, such as Australopithecus, that arose after the split from orangutans . Our family tree, which has 3 main branches leading to chimpanzees, humans and...
originated there. It is now believed that H. erectus is a descendant of earlier genera such as
ArdipithecusArdipithecus is a very early hominin genus. Two species are described in the literature: A. ramidus, which lived about 4.4 million years ago during the early Pliocene, and A. kadabba, dated to approximately 5.6 million years ago ....
and
AustralopithecusAustralopithecus is a genus of hominids that is now extinct. From the evidence gathered by palaeontologists and archaeologists, it appears that the Australopithecus genus evolved in eastern Africa around 4 million years ago before spreading throughout the continent and eventually becoming extinct...
, or early Homo-species such as H. habilis or H. ergaster. H. habilis and H. erectus coexisted for several thousand years, and may represent separate lineages of a common ancestor.
The skull of Tchadanthropus uxoris, discovered in 1961 by Yves Coppens in Chad, is the earlier fossil human discovered in Africa. This fossil "had been so eroded by wind-blown sand that it mimicked the appearance of an australopith, a primitive type of hominid". Though some first considered it to be a specimen of H. habilis, it is no longer considered to be a valid taxon, and scholars rather consider it to represent H. erectus.
Classification and special distinction
Many paleoanthropologists still debate the definition of H. erectus and H. ergaster as separate species. Several scholars suggested to drop the taxon Homo erectus, and suggested instead to equate Homo erectus with archaic Homo sapiens Some call H. ergaster the direct African ancestor of H. erectus, proposing that it emigrated out of Africa and migrated into
AsiaAsia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...
, branching into a distinct species. Most dispense with the species-name ergaster, making no distinction between such fossils as the
Turkana BoyTurkana Boy, also occasionally, Nariokotome Boy is the common name of fossil KNM-WT 15000, a nearly complete skeleton of a hominid who died in the early Pleistocene. This specimen is the most complete early human skeleton ever found. It is 1.5 million years old...
and
Peking ManPeking Man , Homo erectus pekinensis, is an example of Homo erectus. A group of fossil specimens was discovered in 1923-27 during excavations at Zhoukoudian near Beijing , China...
. Though "Homo ergaster" has gained some acceptance as a valid taxon, these two are still usually defined as distinct African and Asian populations of the larger species H. erectus.
While some have argued (and insisted) that
Ernst MayrErnst Walter Mayr was one of the 20th century's leading evolutionary biologists. He was also a renowned taxonomist, tropical explorer, ornithologist, historian of science, and naturalist...
's biological species definition cannot be used here to test the above hypotheses, one can, however, examine the amount of morphological cranial variation within known H. erectus / H. ergaster specimens, and compare it to what one sees in disparate extant groups of primates with similar geographical distribution or close evolutionary relationship. Thus, if the amount of variation between H. erectus and H. ergaster is greater than what one sees within a species of, say,
macaqueThe macaques constitute a genus of Old World monkeys of the subfamily Cercopithecinae. - Description :Aside from humans , the macaques are the most widespread primate genus, ranging from Japan to Afghanistan and, in the case of the barbary macaque, to North Africa...
s, then H. erectus and H. ergaster may be considered two different species.
The extant model of comparison is very important, and selecting appropriate species can be difficult. (For example, the morphological variation among the global population of H. sapiens is incredibly small, as attested by the
Mitochondrial EveIn the field of human genetics, Mitochondrial Eve refers to the matrilineal "MRCA" . In other words, she was the woman from whom all living humans today descend, on their mother's side, and through the mothers of those mothers and so on, back until all lines converge on one person...
hypothesis, and our own special diversity may not be a trustworthy comparison.)
Another closely related, and possibly ancestral, species is
Homo georgicusHomo georgicus is a species of Homo that was suggested in 2002 to describe fossil skulls and jaws found in Dmanisi, Georgia in 1999 and 2001, which seem intermediate between Homo habilis and H. erectus. A partial skeleton was discovered in 2001. The fossils are about 1.8 million years old...
: fossils examples of this species were found in
DmanisiDmanisi is a townlet and archaeological site in Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera.- History :...
in the Republic of Georgia, and are considered to display traits intermediate between H. erectus and H. habilis.
H. erectus had a cranial capacity greater than that of
Homo habilisHomo habilis is a species of the genus Homo, which lived from approximately at the beginning of the Pleistocene period. The discovery and description of this species is credited to both Mary and Louis Leakey, who found fossils in Tanzania, East Africa, between 1962 and 1964. Homo habilis Homo...
(although the Dmanisi specimens have distinctively small crania): the earliest remains show a cranial capacity of 850 cm³, while the latest Javan specimens measure up to 1100 cm³, overlapping that of H. sapiens.; the
frontal boneThe frontal bone is a bone in the human skull that resembles a cockleshell in form, and consists of two portions:* a vertical portion, the squama frontalis, corresponding with the region of the forehead....
is less sloped and the dental arcade smaller than the
australopithecineThe term australopithecine refers generally to any species in the related genera Australopithecus or Paranthropus. These species occurred in the Plio-Pleistocene era, and were bipedal and dentally similar to humans, but with a brain size not much larger than modern apes, lacking the...
s'; the face is more orthognatic (less protrusive) than either the australopithecines' or H. habiliss, with large brow-ridges and less prominent
zygomaThe term zygoma generally refers to the zygomatic bone, a bone of the human skull commonly referred to as the cheekbone or malar bone, but it may also refer to:...
ta (cheekbones). These early hominins stood about , and were more robust than modern humans.
The
sexual dimorphismSexual dimorphism is a phenotypic difference between males and females of the same species. Examples of such differences include differences in morphology, ornamentation, and behavior.-Examples:-Ornamentation / coloration:...
between males and females was slightly greater than seen in H. sapiens, with males being about 25% larger than females. However, their dimorphism is drastically lesser than that of the earlier
AustralopithecusAustralopithecus is a genus of hominids that is now extinct. From the evidence gathered by palaeontologists and archaeologists, it appears that the Australopithecus genus evolved in eastern Africa around 4 million years ago before spreading throughout the continent and eventually becoming extinct...
genus. The discovery of the skeleton KNM-WT 15000, "
Turkana boyTurkana Boy, also occasionally, Nariokotome Boy is the common name of fossil KNM-WT 15000, a nearly complete skeleton of a hominid who died in the early Pleistocene. This specimen is the most complete early human skeleton ever found. It is 1.5 million years old...
" (Homo ergaster), made near
Lake TurkanaLake Turkana , formerly known as Lake Rudolf, is a lake in the Great Rift Valley in Kenya, with its far northern end crossing into Ethiopia. It is the world's largest permanent desert lake and the world's largest alkaline lake...
,
KenyaKenya , officially known as the Republic of Kenya, is a country in East Africa that lies on the equator, with the Indian Ocean to its south-east...
by
Richard LeakeyRichard Erskine Frere Leakey is a politician, paleoanthropologist and conservationist. He is second of the three sons of the archaeologists Louis Leakey and Mary Leakey, and is the younger brother of Colin Leakey...
and
Kamoya KimeuKamoya Kimeu, is one of the world's most successful fossil collectors who, together with paleontologists Meave Leakey and Richard Leakey, is responsible for some of the most significant paleoanthropological discoveries...
in 1984, is one of the most complete hominid-skeletons discovered, and has contributed greatly to the interpretation of human physiological evolution.
For the remainder of this article, the name Homo erectus will be used to describe a distinct species for the convenience of continuity.
Use of tools
Homo ergaster used more diverse and sophisticated
stone toolA stone tool is, in the most general sense, any tool made either partially or entirely out of stone. Although stone tool-dependent societies and cultures still exist today, most stone tools are associated with prehistoric, particularly Stone Age cultures that have become extinct...
s than its predecessors: H. erectus, however, used comparatively primitive tools. This is possibly because H. ergaster first used tools of Oldowan technology and later progressed to the
AcheuleanAcheulean is the name given to an archaeological industry of stone tool manufacture associated with early humans during the Lower Palaeolithic era across Africa and much of West Asia, South Asia and Europe. Acheulean tools are typically found with Homo erectus remains...
: while the use of Acheulean tools began ca.
1.8 million years ago, the line of H. erectus diverged some 200,000 years before the general innovation of Acheulean technology. Thus the Asian migratory descendants of H. ergaster made no use of any Acheulean technology. In addition, it has been suggested that H. erectus may have been the first hominid to use rafts to travel over oceans.
Use of fire
Sites in Europe and Asia seem to indicate controlled use of
fireFire is the rapid oxidation of a material in the chemical process of combustion, releasing heat, light, and various reaction products. Slower oxidative processes like rusting or digestion are not included by this definition....
by H. erectus, some dating back 1.5 million years ago. A presentation at the
PaleoanthropologyPaleoanthropology, which combines the disciplines of paleontology and physical anthropology, is the study of ancient humans as found in fossil hominid evidence such as petrifacted bones and footprints.-19th century:...
Society annual meeting in Montreal, Canada in March 2004 stated that there is evidence for controlled fires in excavations in northern
IsraelThe State of Israel is a parliamentary republic located in the Middle East, along the eastern shore of the Mediterranean Sea...
from about 690,000 to 790,000 years ago. A site called
Terra AmataTerra Amata is an archeological site in open air located on the slopes of Mount Boron in Nice, at a level 26 meters above the current sea level of the Mediterranean. It was discovered and excavated in 1966 by Henry de Lumley...
, located on the
French RivieraThe Côte d'Azur, pronounced , often known in English as the French Riviera , is the Mediterranean coastline of the southeast corner of France, also including the sovereign state of Monaco...
, which lies on an ancient
beachA beach is a geological landform along the shoreline of an ocean, sea, lake or river. It usually consists of loose particles which are often composed of rock, such as sand, gravel, shingle, pebbles or cobblestones...
, seems to have been occupied by H. erectus; it contains the earliest evidence of controlled fire, dated at around 300,000 years BC. Excavations dating from approximately 790,000 years ago in Israel suggest that H. erectus not only controlled fire but could light fires. Despite these examples, some scholars continue to assert that the controlled use of fire was not typical of H. erectus, but only of later species of Homo, such as H. heidelbergensis, H. neanderthalensis, and H. sapiens).
Sociality
Homo erectus was probably the first hominid to live in a
hunter-gathererA hunter-gatherer or forage society is one in which most or all food is obtained from wild plants and animals, in contrast to agricultural societies which rely mainly on domesticated species. Hunting and gathering was the ancestral subsistence mode of Homo, and all modern humans were...
society, and anthropologists such as
Richard LeakeyRichard Erskine Frere Leakey is a politician, paleoanthropologist and conservationist. He is second of the three sons of the archaeologists Louis Leakey and Mary Leakey, and is the younger brother of Colin Leakey...
believe that it was socially more like modern humans than the more
AustralopithecusAustralopithecus is a genus of hominids that is now extinct. From the evidence gathered by palaeontologists and archaeologists, it appears that the Australopithecus genus evolved in eastern Africa around 4 million years ago before spreading throughout the continent and eventually becoming extinct...
-like species before it. Likewise, increased cranial capacity generally coincides with the more sophisticated tools occasionally found with fossils.
The discovery of
Turkana boyTurkana Boy, also occasionally, Nariokotome Boy is the common name of fossil KNM-WT 15000, a nearly complete skeleton of a hominid who died in the early Pleistocene. This specimen is the most complete early human skeleton ever found. It is 1.5 million years old...
(H. ergaster) in 1984 gave evidence that, despite its Homo-sapiens-like anatomy, it may not have been capable of producing sounds comparable to modern human speech. Ergaster likely communicated in a proto-language lacking the fully developed structure of modern human language but more developed than the non-verbal communication used by
chimpanzeeChimpanzee, sometimes colloquially chimp, is the common name for the two extant species of ape in the genus Pan. The Congo River forms the boundary between the native habitat of the two species:...
s. Such inference has been challenged by the discovery of H. ergaster/erectus vertebrae some 150,000 years older than the Turkana Boy in Dmanisi, Georgia, that reflect vocal capabilities within the range of H. sapiens. Both brain-size and the presence of the
Broca's areaBroca's area is a region of the hominid brain with functions linked to speech production.The production of language has been linked to the Broca’s area since Pierre Paul Broca reported impairments in two patients. They had lost the ability to speak after injury to the posterior inferior frontal...
also support the use of articulate language.
H. erectus was probably the first hominid to live in small, familiar band-societies similar to modern hunter-gatherer band-societies. H. erectus/ergaster is thought to be the first hominid to hunt in coordinated groups, use complex tools, and care for infirm or weak companions.
There has been some debate as to whether H. erectus, and possibly the later Neanderthals may have interbred with anatomically modern humans in
EuropeEurope is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...
and
AsiaAsia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...
. See Neanderthal admixture theory.
Descendants and subspecies
Homo erectus remains one of the most successful and long-lived species of Homo. As a distinct Asian species, however, no consensus has been reached as to whether it is ancestral to H. sapiens or any later hominids.
- Homo erectus
- Homo erectus erectus
- Homo erectus yuanmouensis
Yuanmou Man , Homo erectus yuanmouensis, refers to a member of the homo genus whose remnants, two incisors, were discovered near Danawu Village in Yuanmou County in southwestern province of Yunnan, China. Later, stone artifacts, pieces of animal bone showing signs of human work and ash from...
- Homo erectus lantianensis
Lantian Man , formerly Sinanthropus lantianensis is a subspecies of Homo erectus. Its discovery in 1963 was first described by J. K...
- Homo erectus wushanensis
Wushan Man is a controversial taxon. Originally considered a subspecies of Homo erectus, it is now thought by one of the scientists, Russell Ciochon, that first described it to be based upon fossilized fragments of an extinct non-hominin ape...
- Homo erectus nankinensis
Nanjing Man a subspecies of Homo erectus found in China. A male and a female skull of Nanjing Man were discovered in 1993 in Tangshan Cave near Nanjing, and they have been dated to be about 580,000 to 620,000 years old. The skull is now preserved in Nanjing Museum and is under in-depth research by...
- Homo erectus pekinensis
Peking Man , Homo erectus pekinensis, is an example of Homo erectus. A group of fossil specimens was discovered in 1923-27 during excavations at Zhoukoudian near Beijing , China...
- Homo erectus palaeojavanicus
Meganthropus is a name commonly given to several large jaw and skull fragments from Sangiran, Central Java. The original scientific name was Meganthropus palaeojavanicus, and while it is commonly considered invalid today, the genus name has survived as something of an informal nickname for the...
- Homo erectus soloensis
Homo erectus soloensis, known as Solo Man and formerly classified as Homo sapiens soloensis, is generally now regarded as a subspecies of the extinct hominin, Homo erectus. The only known specimens of this anomalous hominid were retrieved from sites along the Bengawan Solo River, on the Indonesian...
- Homo erectus tautavelensis
Related species
- Homo ergaster
Homo ergaster is an extinct chronospecies of Homo that lived in eastern and southern Africa during the early Pleistocene, about 2.5–1.7 million years ago.There is still disagreement on the subject of the classification, ancestry, and progeny of H...
- Homo georgicus
Homo georgicus is a species of Homo that was suggested in 2002 to describe fossil skulls and jaws found in Dmanisi, Georgia in 1999 and 2001, which seem intermediate between Homo habilis and H. erectus. A partial skeleton was discovered in 2001. The fossils are about 1.8 million years old...
- Homo floresiensis
Homo floresiensis is a possible species, now extinct, in the genus Homo. The remains were discovered in 2003 on the island of Flores in Indonesia. Partial skeletons of nine individuals have been recovered, including one complete cranium...
- Homo antecessor
Homo antecessor is an extinct human species dating from 1.2 million to 800,000 years ago, that was discovered by Eudald Carbonell, Juan Luis Arsuaga and J. M. Bermúdez de Castro. H. antecessor is one of the earliest known human varieties in Europe. Various archaeologists and anthropologists have...
- Homo heidelbergensis
Homo heidelbergensis is an extinct species of the genus Homo which may be the direct ancestor of both Homo neanderthalensis in Europe and Homo sapiens. The best evidence found for these hominins date between 600,000 and 400,000 years ago. H...
- Homo neanderthalensis
The Neanderthal is an extinct member of the Homo genus known from Pleistocene specimens found in Europe and parts of western and central Asia...
- Homo sapiens
- Homo rhodesiensis
Homo rhodesiensis is a hominin species described from the fossil Kabwe skull. Other morphologically-comparable remains have been found from the same, or earlier, time period in southern Africa , East Africa and North Africa...
- Homo cepranensis
Homo cepranensis is a proposed name for a human species, known from only one skull cap discovered in 1994. The fossil was discovered by archeologist Italo Biddittu and was nicknamed "Ceprano Man" after a nearby town in the province of Frosinone, 89 kilometers Southeast of Rome, Italy.The age of...
The discovery of
Homo floresiensisHomo floresiensis is a possible species, now extinct, in the genus Homo. The remains were discovered in 2003 on the island of Flores in Indonesia. Partial skeletons of nine individuals have been recovered, including one complete cranium...
in 2003 and of the recentness of its extinction has raised the possibility that numerous descendant species of Homo erectus may have existed in the islands of
Southeast AsiaSoutheast Asia, South-East Asia, South East Asia or Southeastern Asia is a subregion of Asia, consisting of the countries that are geographically south of China, east of India, west of New Guinea and north of Australia. The region lies on the intersection of geological plates, with heavy seismic...
and await fossil discovery (see
Orang PendekOrang Pendek is the most common name given to a cryptid, or cryptozoological animal, that reportedly inhabits remote, mountainous forests on the island of Sumatra....
). Homo erectus soloensis, who lived on Java at least as late as about 50,000 years ago, would be one of them. Some scientists are skeptical of the claim that Homo floresiensis is a descendant of Homo erectus. One explanation holds that the fossils are of a modern human with
microcephalyMicrocephaly is a neurodevelopmental disorder in which the circumference of the head is more than two standard deviations smaller than average for the person's age and sex. Microcephaly may be congenital or it may develop in the first few years of life...
, while another one holds that they are from a group of
pygmyPygmy is a term used for various ethnic groups worldwide whose average height is unusually short; anthropologists define pygmy as any group whose adult men grow to less than 150 cm in average height. A member of a slightly taller group is termed "pygmoid." The best known pygmies are the Aka,...
s.
Individual fossils
Some of the major Homo erectus fossils:
- Indonesia (island of Java): Trinil 2 (holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
), SangiranSangiran is an archaeological excavation site on the island of Java in Indonesia. The area comprises about 48 km² and is located in Central Java, about 15 kilometers north of Surakarta in the Solo River valley. In 1996 it was accepted as World Heritage by the UNESCO.-History:In 1934 the...
collection, Sambungmachan collection, Ngandong collection
- China: Lantian (Gongwangling and Chenjiawo), Yunxian, Zhoukoudian
Zhoukoudian or Choukoutien is a cave system in Beijing, China. It has yielded many archaeological discoveries, including one of the first specimens of Homo erectus, dubbed Peking Man, and a fine assemblage of bones of the gigantic hyena Pachycrocuta brevirostris...
, Nanjing, Hexian
- India: Narmada (taxonomic status debated!)
- Kenya: WT 15000 (Nariokotome), ER 3883, ER 3733
- Tanzania: OH 9
- Vietnam: Northern, Tham Khuyen, Hoa Binh
- Republic of Georgia: Dmanisi collection
- Ethiopia: Daka calvaria
The Daka Calvaria , specimen number BOU-VP-2/66, is a Homo erectus specimen from the Daka Member of the Bouri Formation in the Middle Awash Study Area of the Awash valley of the Ethiopia Rift. It was discovered in 1997 by Henry Gilbert...
- Eritrea: Buia cranium
- Denizli Province
Denizli Province is a province of Turkey in Western Anatolia, on high ground above the Aegean coast. Neighbouring provinces are Uşak to the north, Burdur, Isparta, Afyon to the east, Aydın, Manisa to the west and Muğla to the south. It is located between the coordinates 28° 30’ and 29° 30’ E and...
,Turkey: Kocabas fossil
See also
- Homo ergaster
Homo ergaster is an extinct chronospecies of Homo that lived in eastern and southern Africa during the early Pleistocene, about 2.5–1.7 million years ago.There is still disagreement on the subject of the classification, ancestry, and progeny of H...
- Java Man
Java Man is the name given to fossils discovered in 1891 at Trinil - Ngawi Regency on the banks of the Solo River in East Java, Indonesia, one of the first known specimens of Homo erectus...
- List of fossil sites (with link directory)
- List of primate and hominin fossils (with images)
External links