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Homo heidelbergensis

 
Homo Heidelbergensis

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Homo heidelbergensis



 
 
Homo heidelbergensis ("Heidelberg Man") is an extinct species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
 of the genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 Homo
Homo (genus)

Homo is the genus that includes anatomically modern humanss and their close relatives. The genus is estimated to be about 2.5 million years old, evolving from Australopithecine ancestors with the appearance of Homo habilis....
 which may be the direct ancestor of Homo neanderthalensis in Europe
Europe

Europe is, conventionally, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally divided from Asia to its east by the water divide of the Ural Mountains, the Ural , the Caspian Sea, and by the Caucasus Mountains to the southeast....
. The best evidence found for these hominins date between 600,000 and 400,000 years ago. H. heidelbergensis stone tool technology was considerably close to that of the Acheulean
Acheulean

Acheulean is the name given to an archaeological industry of stone tool manufacture associated with prehistoric hominins during the Lower Palaeolithic era across Africa and much of West Asia and Europe....
 tools used by Homo erectus
Homo Erectus

Homo Erectus is a 2007 comedy film about cavemen that was written and directed by Adam Rifkin, and starring Giuseppe Andrews, Gary Busey, David Carradine, Ron Jeremy, Ali Larter, Hayes MacArthur, Adam Rifkin, and Talia Shire....
.

H.






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Homo heidelbergensis ("Heidelberg Man") is an extinct species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
 of the genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 Homo
Homo (genus)

Homo is the genus that includes anatomically modern humanss and their close relatives. The genus is estimated to be about 2.5 million years old, evolving from Australopithecine ancestors with the appearance of Homo habilis....
 which may be the direct ancestor of Homo neanderthalensis in Europe
Europe

Europe is, conventionally, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally divided from Asia to its east by the water divide of the Ural Mountains, the Ural , the Caspian Sea, and by the Caucasus Mountains to the southeast....
. The best evidence found for these hominins date between 600,000 and 400,000 years ago. H. heidelbergensis stone tool technology was considerably close to that of the Acheulean
Acheulean

Acheulean is the name given to an archaeological industry of stone tool manufacture associated with prehistoric hominins during the Lower Palaeolithic era across Africa and much of West Asia and Europe....
 tools used by Homo erectus
Homo Erectus

Homo Erectus is a 2007 comedy film about cavemen that was written and directed by Adam Rifkin, and starring Giuseppe Andrews, Gary Busey, David Carradine, Ron Jeremy, Ali Larter, Hayes MacArthur, Adam Rifkin, and Talia Shire....
.

Morphology and interpretations

Both H. antecessor and H. heidelbergensis are likely descended from the morphologically very similar Homo ergaster
Homo ergaster

Homo ergaster is an extinct hominin species which lived throughout eastern and southern Africa between 1.9 to 1.4 million years ago with the advent of the lower Pleistocene and the cooling of the global climate....
 from Africa. But because H. heidelbergensis had a larger brain-case with a typical cranial average of modern humans and had more advanced tools and had more behavior, it has been given a separate species classification. The species was 1.8 meters tall on average, and more muscular than modern humans.

RAO hypothesis

According to the "Recent Out of Africa" theory, similar "Archaic Homo sapiens" found in Africa
Africa

Africa is the world's second-largest and second most-populous continent, after Asia. At about 30.2 million km? including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area....
 (ie. Homo sapiens idaltu
Homo sapiens idaltu

'Homo sapiens idaltu' is an extinct subspecies of human that lived almost Lower Paleolithic in Pleistocene Africa. is the Afar language word for "elder, first born"....
 only 160,000 years old), existing in Africa as a part of the operation of the Saharan pump
Sahara pump theory

The Sahara Pump Theory explains how Floristic province and Biomes left Africa to penetrate the Middle East and beyond to Europe and Asia. African pluvial periods are associated with a "wet Sahara" phase during which larger lakes and more rivers exist....
, and not the European forms of Homo heidelbergensis, are thought to be direct ancestors of modern Homo sapiens.

Social behavior

In theory recent findings in Atapuerca
Atapuerca

The 'Atapuerca Mountains' is an ancient karst topography region of Spain, near the town of Atapuerca and Ibeas de Juarros, containing several caves, where fossils and stone tools of the earliest known Homininas in West Europe have been found....
 also suggest that H. heidelbergensis may have been the first species of the Homo genus to bury their dead, but that is contested at this time. Some experts believe that H. heidelbergensis, like its descendant H. neanderthalensis acquired a primitive form of language. No forms of art or sophisticated artifacts other than stone tools have been uncovered, although red ochre
Red ochre

Red ochre and yellow ochre are pigments made from naturally tinted clay. It has been used worldwide since prehistoric times. Chemically, it is hydrated iron oxide....
, a mineral that can be used to create a red pigment which is useful as a paint, has been found at Terra Amata
Terra Amata

Terra Amata is an archaeology site near the France town of Nice.Terra Amata was an open site where Acheulean flint tools were found, dating it to the Lower Paleolithic....
 excavations in the south of France.

Language

The morphology of the outer and middle ear suggests they had an auditory sensitivity similar to modern humans and very different from chimpanzees. They were probably able to differentiate between many different sounds.

Evidence of hunting

Cut marks found on wild deer
Deer

Deer are the ruminant mammals forming the family Cervidae . A number of broadly similar animals from related families within the order even-toed ungulate are often also called deer....
, elephant
Elephant

Elephants are large land mammals of the order Proboscidea and the family Elephantidae. There are three living species: the African Bush Elephant, the African Forest Elephant and the Asian Elephant ....
s, rhinos and horse
Horse

The horse is a hoofed mammal, a subspecies of one of seven extant species of the family Equidae. The horse has evolution of the horse over the past 45 to 55 million years from a small multi-toed creature into the large, odd-toed ungulate animal of today....
s demonstrate that they were butchered, some of the animals weighed as much as 700 kg (1,500 lb) or possibly larger. During this era, now-extinct wild animals such as mammoth
Mammoth

A mammoth is any species of the extinct genus Mammuthus. These proboscideans are members of the Elephantidae and close relatives of modern elephants....
s, European lion
European lion

The European lion could be an extinct subspecies of lion that inhabited southern Europe until historic times. This population is generally considered part of the Asiatic lion , but others consider it a separate subspecies, the European lion ....
s and Irish elk
Irish Elk

The Irish Elk or Giant Deer, Megaloceros giganteus was a species of Megaloceros and one of the largest deer that ever lived....
 roamed the European continent.

Moreover, a number of 400,000 year old wooden projectile spears were found at Schöningen
Schöningen

Sch?ningen is a city of 13,000 inhabitants in the district of Helmstedt , Lower Saxony, Germany. In its current form, it was created in 1974 by joining the municipalities of Esbeck, Hoiersdorf, and Sch?ningen....
 in northern Germany. These are thought to have been made by Homo erectus
Homo Erectus

Homo Erectus is a 2007 comedy film about cavemen that was written and directed by Adam Rifkin, and starring Giuseppe Andrews, Gary Busey, David Carradine, Ron Jeremy, Ali Larter, Hayes MacArthur, Adam Rifkin, and Talia Shire....
 or Homo heidelbergensis. Generally, projectile weapons are more commonly associated with H. sapiens. The lack of projectile weaponry is an indication of different sustenance methods, rather than inferior technology or abilities. The situation is identical to that of native New Zealand Maori - modern Homo sapiens, who also rarely threw objects, but used spears and clubs instead.

Discovery


The first fossil discovery of this species was made on October 21, 1907 and came from Mauer
Mauer (Baden)

Mauer is a village in south western Germany. It is located between Heidelberg and Sinsheim in the Rhein-Neckar-Kreis district in the state of Baden-W?rttemberg....
 where the workman Daniel Hartmann spotted a jaw in a sandpit. The jaw (Mauer 1
Mauer 1

Mauer 1 is a fossilized mandible and the type specimen of the species Homo heidelbergensis. It was discovered by Daniel Hartmann in Mauer , Germany in 1907....
) was in good condition except for the missing premolar teeth, which were eventually found near the jaw. The workman gave it to professor Otto Schoetensack
Otto Schoetensack

Otto Schoetensack was a Germany industrialist and later professor of anthropology. During an archeological dig he directed the worker Daniel Hartmann found the lower jaw of a hominid, which Schoetensack later named the Homo heidelbergensis....
 from the University of Heidelberg, who identified and named the fossil.

The next Homo heidelbergensis remains were found in Steinheim an der Murr
Steinheim an der Murr

Steinheim an der Murr is a town in the Ludwigsburg , Baden-W?rttemberg, Germany. It is situated on the river Murr, 9 km northeast of Ludwigsburg. The Steinheim Skull was found near Steinheim an der Murr....
, Germany
Germany

Germany , officially the Federal Republic of Germany , is a country in Central Europe. It is bordered to the north by the North Sea, Denmark, and the Baltic Sea; to the east by Poland and the Czech Republic; to the south by Austria and Switzerland; and to the west by France, Luxembourg, Belgium, and the Netherlands....
 (the Steinheim Skull
Steinheim Skull

The Steinheim skull is a fossilized skull of an archaic Archaic Homo sapiens or Homo heidelbergensis found in 1933 near Steinheim an der Murr ....
, 350kya), Arago, France
France

France , officially the French Republic , is a country whose Metropolitan France is located in Western Europe and that also comprises various Overseas departments and territories of France....
 (Arago 21), Petralona, Greece
Greece

Greece , officially the Hellenic Republic , is a country in southeastern Europe, situated on the southern end of the Balkans. It has borders with Albania, Bulgaria and the former Yugoslav Republic of Macedonia to the north, and Turkey to the east....
 and Ciampate del Diavolo
Ciampate del Diavolo

Ciampate del Diavolo is the name of a locality near Roccamonfina , in the neighbourhood of an extinct volcano with the same name. It takes its name from several human fossil....
, Italy
Italy

Italy , officially the Italian Republic , is a country located on the Italian Peninsula in Southern Europe and on the two largest islands in the Mediterranean Sea, Sicily and Sardinia....
.

Boxgrove Man

In 1994 British
United Kingdom

The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom , the UK or Britain,is a sovereign state located off the northwestern coast of continental Europe....
 scientists had unearthed a lower hominin tibia bone just a few kilometres away from the English Channel
English Channel

The English Channel is an Arm of the Atlantic Ocean that separates England from northern France, and joins the North Sea to the Atlantic. It is about long and varies in width from at its widest, to only in the Strait of Dover....
 including hundreds of ancient hand axes at the Boxgrove Quarry
Boxgrove Quarry

Boxgrove Quarry is a gravel quarry and Lower Palaeolithic archaeology archaeological site at Boxgrove in the England county of West Sussex.Parts of the site complex were excavated between 1983 and 1996 by a team led by Mark Roberts of University College London....
 site. A partial leg bone is dated to 478,000 and 524,000 years old. Homo heidelbergensis was the early proto-human species that occupied both France
France

France , officially the French Republic , is a country whose Metropolitan France is located in Western Europe and that also comprises various Overseas departments and territories of France....
 and Great Britain
Great Britain

Great Britain is an island lying to the northwest of Continental Europe. It is the List of islands by area, and the largest in Europe. With a population of 58.9 million people it is List of islands by population....
 at that time; both locales were connected by a landmass during that epoch
Geologic time scale

File:Geologic clock.jpgThe geologic time scale is a chronology schema relating stratigraphy to time that is used by geologys and other earth sciences scientists to describe the timing and relationships between events that have occurred during the history of the Earth....
. Prior to Gran Dolina, Boxgrove offered the earliest hominid occupants in Europe.

The tibia had been gnawed by a large carnivore, suggesting that he had been killed by a lion or wolf or that his unburied corpse had been scavenged after death .

Sima de los Huesos

Beginning in 1997, a Spanish
Spain

Spain or the Kingdom of Spain , is a country located in Southern Europe on the Iberian Peninsula.The Spanish constitution does not establish any official denomination of the country, even though Espa?a , Estado espa?ol and Naci?n espa?ola are used interchangeably....
 team has located more than 5,500 human bones dated to an age of at least 350,000 years old in the Sima de los Huesos site in the Sierra de Atapuerca
Atapuerca

The 'Atapuerca Mountains' is an ancient karst topography region of Spain, near the town of Atapuerca and Ibeas de Juarros, containing several caves, where fossils and stone tools of the earliest known Homininas in West Europe have been found....
 in northern Spain. The pit contains fossils of perhaps 28 individuals together with remains of Ursus deningeri and other carnivores and a biface
Biface

In archaeology, a biface is a two-sided stone tool, manufactured through a process of lithic reduction, that displays lithic flake scars on both sides....
 called Excalibur
Excalibur

Excalibur is the legendary sword of King Arthur sometimes attributed with magical powers or associated with the rightful sovereignty of Great Britain....
. It is hypothesized that this Acheulean
Acheulean

Acheulean is the name given to an archaeological industry of stone tool manufacture associated with prehistoric hominins during the Lower Palaeolithic era across Africa and much of West Asia and Europe....
 axe made of red quartzite to be some kind of ritual offering for a funeral. Ninety percent of the known Homo heidelbergensis remains has been obtained from this site. The fossil pit bones include:
  • A complete cranium (Skull 5), nicknamed Miguelón
    Miguelón

    Miguel?n is the popular nickname for the most complete skull of an Homo heidelbergensis ever found. More than 5,500 human fossils of this species, which are considered to be the direct ancestor of Homo neanderthalensis, have been found in the Sima de los Huesos site in the Sierra de Atapuerca in northern Spain....
    , and fragments of other craniums, as Skull 4, nicknamed Agamenón and skull 6, nicknamed Rui (from El Cid
    El Cid

    Rodrigo D?az de Vivar , known as El Cid Campeador, was a Kingdom of Castile nobleman, a gifted military leader and diplomat who, after being exiled, conquered and governed the city of Valencia ....
    , a local hero).
  • A complete pelvis
    Pelvis

    The pelvis or pelvic girdle is the irregular bone structure located at the base of the spine . In the adult human, it is formed by the sacrum and the coccyx, the caudal part of the axial skeleton, and a pair of hip bones, part of the appendicular skeleton or human leg....
     (Pelvis 1), nicknamed Elvis, in remembrance of Elvis Presley
    Elvis Presley

    Elvis Aaron Presley was an United Statesn singer, actor, and musician. A cultural icon, he is commonly known simply as "Elvis", and is also sometimes referred to as "List of honorific titles in popular music" or "The King"....
    .
  • Mandibles, teeth, a lot of postcranial bones (femur
    Femur

    The femur, or thigh bone, is the most proximal bone of the leg in vertebrates capable of walking or jumping, such as most land mammals, birds, many reptiles such as lizards, and amphibians such as frogs....
    s, hand and foot bones, vertebrae
    Vertebral column

    In human anatomy, the vertebral column is a column of 24 vertebrae, the sacrum, intervertebral discs, and the coccyx situated in the dorsum aspect of the torso, separated by spinal discs....
    , ribs, etc.)
Indeed, nearby sites contain the only known and controversial Homo antecessor
Homo antecessor

Homo antecessor is an extinct hominin and a potential distinct species dating from 1.2 million to 800,000 years ago, that was discovered by Eudald Carbonell, Juan Luis Arsuaga and J....
 fossils.

Extinction or evolution

Some believe that H. heidelbergensis is an extinct species, some that is a cladistic ancestor to other Homo forms sometimes improperly linked to distinct species in terms of populational genetics.

Some scenarios of survival include
  • H heidelbergensis > H. neanderthalensis > H sapiens sapiens
  • H. heidelbergensis > H. rhodesiensis > H. idaltu > H sapiens sapiens


Those supporting multiregional origin of modern humans envise fertile reproduction between many evolutionary stages; Homo walking, or gene transfer between adjacent populations due to gene passage and spreading in successive generations.

See also

  • List of fossil sites
    List of fossil sites

    This is a worldwide list of important and/or well-known localities where fossils have been found. Such locations may either be a geological formation or a single site....
     (with link directory)
  • List of hominina (hominid) fossils (with images)
  • Saldanha man
    Saldanha man

    Saldanha man also known as Saldanha cranium or Elandsfontein cranium are fossilized remains of a hominid species believed to be Homo heidelbergensis....


External links