Violet (plant)
Encyclopedia
Viola
is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

s in the violet family Violaceae
Violaceae
Violaceae are a family of flowering plants consisting of about 800 species in 21 genera. It takes its name from the genus Viola, the violets and pansies.Older classifications such as the Cronquist system placed Violaceae in an order named after it, the Violales...

, with around 400–500 species distributed around the world. Most species are found in the temperate Northern Hemisphere
Northern Hemisphere
The Northern Hemisphere is the half of a planet that is north of its equator—the word hemisphere literally means “half sphere”. It is also that half of the celestial sphere north of the celestial equator...

; however, viola species (commonly called violets, pansies or heartsease) are also found in widely divergent areas such as Hawaii
Hawaii
Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...

, Australasia
Australasia
Australasia is a region of Oceania comprising Australia, New Zealand, the island of New Guinea, and neighbouring islands in the Pacific Ocean. The term was coined by Charles de Brosses in Histoire des navigations aux terres australes...

, and the Andes
Andes
The Andes is the world's longest continental mountain range. It is a continual range of highlands along the western coast of South America. This range is about long, about to wide , and of an average height of about .Along its length, the Andes is split into several ranges, which are separated...

 in South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

.

Most Viola species are perennial plant
Perennial plant
A perennial plant or simply perennial is a plant that lives for more than two years. The term is often used to differentiate a plant from shorter lived annuals and biennials. The term is sometimes misused by commercial gardeners or horticulturalists to describe only herbaceous perennials...

s, some are annual plant
Annual plant
An annual plant is a plant that usually germinates, flowers, and dies in a year or season. True annuals will only live longer than a year if they are prevented from setting seed...

s, and a few are small shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

s. A number of species are grown for their ornamental flowers in borders and rock gardens; the garden pansy
Pansy
The Pansy is a large group of hybrid plants cultivated as garden flowers. Pansies are derived from Viola species Viola tricolor hybridized with other viola species, these hybrids are referred to as Viola × wittrockiana or less commonly Viola tricolor hortensis...

 in particular is an extensively used spring and autumn/winter bedding and pot plant. Viola and violetta are terms used by gardeners and generally in horticulture
Horticulture
Horticulture is the industry and science of plant cultivation including the process of preparing soil for the planting of seeds, tubers, or cuttings. Horticulturists work and conduct research in the disciplines of plant propagation and cultivation, crop production, plant breeding and genetic...

 for neat, small-flowered hybrid plants intermediate in size between pansies and violets.

Description

Viola species typically have heart-shaped, scalloped leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

, though a number have palmate leaves or other shapes. The vast majority of Viola species are herbaceous
Herbaceous plant
A herbaceous plant is a plant that has leaves and stems that die down at the end of the growing season to the soil level. They have no persistent woody stem above ground...

, and a substantial number are acaulescent in habit - meaning they lack any noticeable stems and the foliage and flowers appear to rise from the ground; the remaining species have short stems with foliage and flowers produced in the axils of the leaves. The simple leaves of plants with either habit are arranged alternately; the acaulescent species produce basal rosettes. Plants always have leaves with stipules that are often leaf-like.
The flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s of the vast majority of the species are zygomorphic with bilateral symmetry. The flowers are formed from five petals; four are upswept or fan-shaped petals with two per side, and there is one broad, lobed lower petal pointing downward. The shape of the petals and placement defines many species, for example, some Viola species have a "spur" on the end of each petal while most have a spur on the lower petal.

Solitary flowers end long stalks with a pair of bracteoles. The flowers have 5 sepals that persist after blooming, and in some species the sepals enlarge after blooming. The flowers have five free stamens with short filaments that are oppressed against the ovary, only the lower two stamens have nectary spurs that are inserted on the lowest petal into the spur or a pouch. The flower styles are thickened near the top and the stigmas are head-like, narrowed or often beaked. The flowers have a superior ovary with one cell, which has three placentae, containing many ovules.

Viola flowers are most often spring blooming with chasmogamous flowers with well-developed petals pollinated by insects. Many species also produce self-pollinated cleistogamous flowers in summer and autumn that do not open and lack petals. In some species the showy chasmogamous flowers are infertile (e.g.,Viola papilionacea).

After flowering, fruit capsules are produced that split open by way of three valves. On drying, the capsules may eject seeds with considerable force to distances of several meters. The nutlike seeds have straight embryos, flat cotyledons, and soft fleshy endosperm that is oily.
The seeds of some species have elaiosome
Elaiosome
Elaiosomes are fleshy structures that are attached to the seeds of many plant species. The elaiosome is rich in lipids and proteins, and may be variously shaped. Many plants have elaiosomes to attract ants, which take the seed to their nest and feed the elaiosome to their larvae...

s and are dispersed by ants.

Flower colours vary in the genus, ranging from violet
Violet (color)
As the name of a color, violet is synonymous with a bluish purple, when the word "purple" is used in the common English language sense of any color between blue and red, not including either blue or red...

, as their common name suggests, through various shades of blue, yellow, white, and cream, whilst some types are bicolored, often blue and yellow. Many cultivars and hybrids have been bred
Plant breeding
Plant breeding is the art and science of changing the genetics of plants in order to produce desired characteristics. Plant breeding can be accomplished through many different techniques ranging from simply selecting plants with desirable characteristics for propagation, to more complex molecular...

 in a greater spectrum of colours. Flowering is often profuse, and may last for much of the spring and summer.

One quirk of some viola is the elusive scent of their flowers; along with terpene
Terpene
Terpenes are a large and diverse class of organic compounds, produced by a variety of plants, particularly conifers, though also by some insects such as termites or swallowtail butterflies, which emit terpenes from their osmeterium. They are often strong smelling and thus may have had a protective...

s, a major component of the scent is a ketone
Ketone
In organic chemistry, a ketone is an organic compound with the structure RCR', where R and R' can be a variety of atoms and groups of atoms. It features a carbonyl group bonded to two other carbon atoms. Many ketones are known and many are of great importance in industry and in biology...

 compound called ionone
Ionone
The ionones are a series of closely related chemical substances that are part of a group of compounds known as rose ketones, which also includes damascones and damascenones. Ionones are aroma compounds found in a variety of essential oils, including rose oil...

, which temporarily desensitises the receptors
Olfactory receptor neuron
An olfactory receptor neuron , also called an olfactory sensory neuron , is a transduction cell within the olfactory system. J. Rospars, Dendritic integration in olfactory sensory neurons: a steady-state analysis of how the neuron structure and neuron environment influence the coding of odor...

 of the nose, thus preventing any further scent being detected from the flower until the nerves recover.

Selected species

See List of Viola species for a more complete list.
  • Viola arvensis
    Viola arvensis
    Viola arvensis is a species of violet known by the common name field pansy. It is native to Europe, western Asia, and North Africa, and it is known on other continents as an introduced species and a weed of disturbed and cultivated areas....


    field pansy
  • Viola banksii
    Viola banksii
    Viola banksii, the native violet, is sold and grown throughout garden nurseries and grown and loved in gardens around Australia, especially in the east. For many years it was known as Viola hederacea, however the species complex was revised in 2004 by Kevin Thiele...


    Australian native violet, ivy-leaved violet
  • Viola biflora
    Viola biflora
    Viola biflora is a species of the genus Viola. It is also called twoflower violet. It is found in Europe, Siberia, Central Asia, Pakistan, Western & Northern China, North Korea, Japan, and Western North America....


    yellow wood violet, twoflower violet
  • Viola canina
    Viola canina
    Viola canina is a species of the genus Viola, native to Europe, where it is found in the uplands of box hill in Dorset, heaths, fens, and moist woodlands, especially on acidic soils....


    heath dog violet
  • Viola cheiranthifolia
    Viola cheiranthifolia
    Viola cheiranthifolia is a species of the genus Viola. It is also known as the Teide Violet and Teide Daisy.This plant is exclusively found in the dry and stony caldera of the volcano Teide on the Canary Island of Tenerife. It survives there in altitudes of 2000-3000m. Most of the time it is hiding...


    Teide violet
  • Viola corsica
    Corsican pansy
  • Viola hederacea
    Viola hederacea
    Viola hederacea is a species of violet which is native to Australia. It is common and widespread in Victoria and Tasmania, along the Great Dividing Range in New South Wales north at least to the Barrington Tops area, in the far south-east of South Australia, and in a small area of the Adelaide...


    Australian native violet, ivy-leaved violet
  • Viola hirta
    Viola hirta
    Viola hirta is a species of the plant genus Viola. It is also called Hairy violet. As with the Sweet Violet no fossil seeds of this species have been found. It is confined to the cold Temperate Zone, in Europe, N. and W. Asia, extending as far as N.-W. India...


    hairy violet
  • Viola labradorica
    Viola labradorica
    Viola labradorica is a perennial native to Greenland, eastern Canada, and the United States. The plant sold as Viola labradorica by nurseries is Viola riviniana.-Culinary:...


    alpine violet
  • Viola odorata
    sweet violet
  • Viola pedunculata
    yellow pansy
  • Viola persicifolia
    fen violet
  • Viola pubescens
    Viola pubescens
    Viola pubescens, often called Downy yellow violet, is a plants species of the genus Viola. It is found in dry mixed and deciduous woodland areas of the United States and Canada; from Minnesota and Ontario to Nova Scotia, South to Virginia...


    downy yellow violet
  • Viola riviniana
    Viola riviniana
    Viola riviniana, the Common Dog-violet, is a species of the genus Viola. It is also called wood violet or dog violet. It is a perennial herb of woodland rides, grassland and shady hedge banks...


    common dog violet
  • Viola rostrata
    Viola rostrata
    Viola rostrata, is a stemmed herbaceous perennial plant that is native to eastern North America. It is known by a number of common names including long-spurred violet.-General description:...


    long-spurred violet
  • Viola sororia
    Viola sororia
    Viola sororia, known commonly as the Common Blue Violet, is a stemless herbaceous perennial plant that is native to eastern North America...


    common blue violet, hooded violet
  • Viola tricolor
    wild pansy, heartsease


  • Note: Neither Saintpaulia
    Saintpaulia
    Saintpaulia, commonly known as African violet, is a genus of six species of herbaceous perennial flowering plants in the family Gesneriaceae, native to Tanzania and adjacent southeastern Kenya in eastern tropical Africa, with a concentration of species in the Nguru mountains of Tanzania...

     ("African violets") nor Erythronium dens-canis
    Erythronium dens-canis
    Erythronium dens-canis is a small herbaceous flowering plant in the Liliaceae, native to Europe, where it is the only naturally occurring species....

     ("dogtooth violets") are related to the true Viola.

    The genus includes dog violet
    Dog violet
    Dog violet is the common name for various species of the plant genus Viola with unscented flowers. The term arose to differentiate them from the scented sweet violet. Species so named include:*Viola canina - heath dog violet...

    s, a group of scentless species which are the most common viola in many areas, sweet violet (Viola odorata) (named from its sweet scent), and many other species whose common name includes the word "violet". Several species are known as pansies, including the yellow pansy (Viola pedunculata) of the Pacific
    Pacific Ocean
    The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

     coast.

    Common blue violet Viola sororia
    Viola sororia
    Viola sororia, known commonly as the Common Blue Violet, is a stemless herbaceous perennial plant that is native to eastern North America...

     is the state flower of Wisconsin
    Wisconsin
    Wisconsin is a U.S. state located in the north-central United States and is part of the Midwest. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michigan to the northeast, and Lake Superior to the north. Wisconsin's capital is...

    , Rhode Island
    Rhode Island
    The state of Rhode Island and Providence Plantations, more commonly referred to as Rhode Island , is a state in the New England region of the United States. It is the smallest U.S. state by area...

    , Illinois
    Illinois
    Illinois is the fifth-most populous state of the United States of America, and is often noted for being a microcosm of the entire country. With Chicago in the northeast, small industrial cities and great agricultural productivity in central and northern Illinois, and natural resources like coal,...

    , and New Jersey
    New Jersey
    New Jersey is a state in the Northeastern and Middle Atlantic regions of the United States. , its population was 8,791,894. It is bordered on the north and east by the state of New York, on the southeast and south by the Atlantic Ocean, on the west by Pennsylvania and on the southwest by Delaware...

    .

    Australia is home to a number of Viola species, including Viola hederacea
    Viola hederacea
    Viola hederacea is a species of violet which is native to Australia. It is common and widespread in Victoria and Tasmania, along the Great Dividing Range in New South Wales north at least to the Barrington Tops area, in the far south-east of South Australia, and in a small area of the Adelaide...

    , Viola betonicifolia
    Viola betonicifolia
    Viola betonicifolia, commonly known as the Arrowhead Violet, Showy Violet or Mountain Violet, is a small perennial herbaceous shrub of the Viola, which contains pansies and violets. It occurs from India and Pakistan in southern Asia throughout eastern Australia and Tasmania...

     and Viola banksii
    Viola banksii
    Viola banksii, the native violet, is sold and grown throughout garden nurseries and grown and loved in gardens around Australia, especially in the east. For many years it was known as Viola hederacea, however the species complex was revised in 2004 by Kevin Thiele...

    , first collected by Joseph Banks
    Joseph Banks
    Sir Joseph Banks, 1st Baronet, GCB, PRS was an English naturalist, botanist and patron of the natural sciences. He took part in Captain James Cook's first great voyage . Banks is credited with the introduction to the Western world of eucalyptus, acacia, mimosa and the genus named after him,...

     and Daniel Solander
    Daniel Solander
    Daniel Carlsson Solander or Daniel Charles Solander was a Swedish naturalist and an apostle of Carl Linnaeus. Solander was the first university educated scientist to set foot on Australian soil.-Biography:...

     on the Cook
    James Cook
    Captain James Cook, FRS, RN was a British explorer, navigator and cartographer who ultimately rose to the rank of captain in the Royal Navy...

     voyage to Botany Bay
    Botany Bay
    Botany Bay is a bay in Sydney, New South Wales, a few kilometres south of the Sydney central business district. The Cooks River and the Georges River are the two major tributaries that flow into the bay...

    .

    The modern garden pansy
    Pansy
    The Pansy is a large group of hybrid plants cultivated as garden flowers. Pansies are derived from Viola species Viola tricolor hybridized with other viola species, these hybrids are referred to as Viola × wittrockiana or less commonly Viola tricolor hortensis...

     (V. × wittrockiana) is a plant of complex hybrid origin involving at least three species, V. tricolor (wild pansy or heartsease), V. altaica and V. lutea
    Viola lutea
    Viola lutea , also known as Mountain Pansy, is a species in the genus Viola. It is native to Europe....

     (mountain pansy).

    Species and cultivars

    Cultivars of Viola cornuta, Viola cucullata
    Viola cucullata
    Viola cucullata is a species of the genus Viola native to eastern North America, from Newfoundland west to Ontario and Minnesota, and south to Georgia....

    , Viola odorata, are commonly grown from seed. Other species often grown include Viola labradorica
    Viola labradorica
    Viola labradorica is a perennial native to Greenland, eastern Canada, and the United States. The plant sold as Viola labradorica by nurseries is Viola riviniana.-Culinary:...

    , Viola pedata
    Viola pedata
    Viola pedata is an ornamental plant in the Violaceae family. Monophyletic in stature, this violet, also known as the "mountain pansy" is endemic to eastern North America, where it favors well drained, acidic soils in full to partial sun environments...

     and Viola rotundifolia.

    Bedding plants

    The plants known to gardeners as "violas" are common bedding and pot plants world wide. In 2005 in the United States, violas (including pansies) were one of the top three bedding plant crops and 111 million dollars worth of flats of violas were produced for the bedding flower market. Pansies and violas used for bedding are generally raised from seed, and F1 hybrid
    F1 hybrid
    F1 hybrid is a term used in genetics and selective breeding. F1 stands for Filial 1, the first filial generation seeds/plants or animal offspring resulting from a cross mating of distinctly different parental types....

     seed strains have been developed which produce compact plants of reasonably consistent flower colouring and appearance. Bedding plants are usually discarded after one growing season.

    Perennial cultivars

    There are hundreds of perennial viola and violetta cultivar
    Cultivar
    A cultivar'Cultivar has two meanings as explained under Formal definition. When used in reference to a taxon, the word does not apply to an individual plant but to all those plants sharing the unique characteristics that define the cultivar. is a plant or group of plants selected for desirable...

    s; many of these do not "come true" from seed and therefore have to be propagated
    Plant propagation
    Plant propagation is the process of creating new plants from a variety of sources: seeds, cuttings, bulbs and other plant parts. Plant propagation can also refer to the artificial or natural dispersal of plants.-Sexual propagation :...

     from cuttings. Violettas can be distinguished from violas by the lack of ray markings on their petals. A few popular examples include:
    • Viola 'Ardross Gem' (viola)
    • Viola 'Buttercup' (violetta)
    • Viola 'Columbine' (viola)
    • Viola 'Dawn' (violetta)
    • Viola 'Etain' (viola)
    • Viola 'Irish Molly' (viola)
    • Viola 'Jackanapes' (viola)
    • Viola 'Maggie Mott' (viola)
    • Viola 'Martin' (viola)
    • Viola 'Molly Sanderson' (viola)
    • Viola 'Rebecca' (violetta)
    • Viola 'Vita' (viola)
    • Viola 'Zoe' (violetta)
    • viola 'Blackjack'

    Weed control

    Not all violets are desired, and wild violets are considered weeds in North American lawns by some people. Others tolerate or welcome the presence of violets in lawns. Wild violets are regarded as a problem in shady fescue
    Fescue
    Festuce is a genus of about 300 species of perennial tufted grasses, belonging to the grass family Poaceae . The genus has a cosmopolitan distribution, although the majority of the species are found in cool temperate areas...

     lawns in North America. Violets thrive in the part/full shade and are not susceptible to most herbicides used to kill common lawn weeds. Triclopyr
    Triclopyr
    Triclopyr is a systemic, foliar herbicide in the pyridine group. It is used to control broadleaf weeds while leaving grasses and conifers unaffected....

    , a weedkiller, has been found to be an effective method of controlling wild violets in fescue lawns.

    Culinary

    When newly opened, Viola flowers may be used to decorate salads or in stuffings for poultry or fish. Soufflés, cream and similar desserts can be flavoured with essence of Viola flowers. The young leaves are edible raw or cooked as a somewhat bland leaf vegetable
    Leaf vegetable
    Leaf vegetables, also called potherbs, green vegetables, greens, leafy greens or salad greens, are plant leaves eaten as a vegetable, sometimes accompanied by tender petioles and shoots...

    . The flowers and leaves of the cultivar 'Rebecca', one of the Violetta violets, has a distinct vanilla flavor with hints of wintergreen.

    A candied
    Candied fruit
    Candied fruit, also known as crystallized fruit or Glacé fruit, has been around since the 14th century. Whole fruit, smaller pieces of fruit, or pieces of peel, are placed in heated sugar syrup, which absorbs the moisture from within the fruit and eventually preserves it...

     violet
    or crystallized violet is a flower, usually of Viola odorata, preserved by a coating of egg white and crystallised sugar. Alternatively, hot syrup is poured over the fresh flower (or the flower is immersed in the syrup) and stirred until the sugar recrystallizes and has dried. This method is still used for rose
    Rose
    A rose is a woody perennial of the genus Rosa, within the family Rosaceae. There are over 100 species. They form a group of erect shrubs, and climbing or trailing plants, with stems that are often armed with sharp prickles. Flowers are large and showy, in colours ranging from white through yellows...

     petals and was applied to orange
    Orange (fruit)
    An orange—specifically, the sweet orange—is the citrus Citrus × sinensis and its fruit. It is the most commonly grown tree fruit in the world....

     flowers in the past (when almond
    Almond
    The almond , is a species of tree native to the Middle East and South Asia. Almond is also the name of the edible and widely cultivated seed of this tree...

    s or orange peel are treated this way they are called praline
    Praline
    Praline is a family of confections made from nuts and sugar syrup.-Europe:As originally inspired in France at the Château of Vaux-le-Vicomte by the cook of the 17th-century sugar industrialist Marshal du Plessis-Praslin , early pralines were whole almonds individually coated in caramelized sugar,...

    s). Candied violets are still made commercially in Toulouse
    Toulouse
    Toulouse is a city in the Haute-Garonne department in southwestern FranceIt lies on the banks of the River Garonne, 590 km away from Paris and half-way between the Atlantic Ocean and the Mediterranean Sea...

    , France, where they are known as violettes de Toulouse. They are used as decorating or included in aromatic desserts.

    The French are also known for their violet syrup
    Syrup
    In cooking, a syrup is a thick, viscous liquid consisting primarily of a solution of sugar in water, containing a large amount of dissolved sugars but showing little tendency to deposit crystals...

    , most commonly made from an extract of violets. In the United States, this French violet syrup is used to make violet scone
    Scone
    -Food:* Scone , a type of quick-bread, typically eaten with jam and cream.* Drop-scone, British word for a small pancake-People:*Barbara Young, Baroness Young of Old Scone , Labour member of the House of Lords...

    s and marshmallow
    Marshmallow
    The marshmallow is a confection that, in its modern form, typically consists of sugar, corn syrup, water, gelatin that has been softened in hot water, dextrose, vanilla flavourings, and sometimes colouring, whipped to a spongy consistency. Some marshmallow recipes call for egg whites...

    s.

    Viola essence flavours the liqueurs Creme Yvette
    Creme Yvette
    Creme Yvette, also called Creme d'Yvette, is a proprietary liqueur made from parma violet petals and possibly vanilla & other spices. It was once manufactured by Charles Jacquin et Cie in the United States, who purchased the brand formerly made by Sheffield Company of Connecticut. It became almost...

    , Creme de Violette
    Creme de Violette
    Crème de Violette, also known as liqueur de violette, is a generic term for a liqueur with natural and/or artificial violet flower flavoring and coloring with either a brandy base, a neutral spirit base, or a combination of the two. The taste profile and aroma are distinctly floral and sweet, and...

    , and Parfait d'Amour
    Parfait d'Amour
    Parfait d'Amour is a liqueur. Purple in colour, it is often used in cocktails primarily for its colour, and is generally created from a Curaçao liqueur base....

    . It is also used in Parma Violets
    Parma Violets
    Parma Violets are a British violet-flavoured tablet confectionery manufactured by the Derbyshire company Swizzels Matlow. The sweets are hard, biconcave disc-shaped sweets, similar to the Fizzers product from the same company but without their fizziness...

     confectionery.

    Medicinal

    The flowers, leaves and roots of various Viola species are used for medicinal purposes, being rich in vitamin
    Vitamin
    A vitamin is an organic compound required as a nutrient in tiny amounts by an organism. In other words, an organic chemical compound is called a vitamin when it cannot be synthesized in sufficient quantities by an organism, and must be obtained from the diet. Thus, the term is conditional both on...

    s A and C. They also contain a type of antioxidant
    Antioxidant
    An antioxidant is a molecule capable of inhibiting the oxidation of other molecules. Oxidation is a chemical reaction that transfers electrons or hydrogen from a substance to an oxidizing agent. Oxidation reactions can produce free radicals. In turn, these radicals can start chain reactions. When...

     called an anthocyanin
    Anthocyanin
    Anthocyanins are water-soluble vacuolar pigments that may appear red, purple, or blue according to pH...

    . Viola flowers are also used to make an herbal tea that is used in Chinese herbal medicine
    Chinese herbology
    Chinese Herbology is the theory of Traditional Chinese herbal therapy, which accounts for the majority of treatments in Traditional Chinese medicine ....

    . Most violas (all tested) and many plants of the Violaceae plant family contain cyclotides
    Cyclotides
    Cyclotides are small disulfide rich peptides isolated from plants. Typically containing 28-37 amino acids, they are characterized by their head-to-tail cyclised peptide backbone and the interlocking arrangement of their three disulfide bonds. These combined features have been termed the cyclic...

    , which have a diverse range of biological activities, including uterotonic, anti-HIV, antimicrobial, and insecticidal activities.

    Viola canescens, a species from from India, exhibited in vitro
    In vitro
    In vitro refers to studies in experimental biology that are conducted using components of an organism that have been isolated from their usual biological context in order to permit a more detailed or more convenient analysis than can be done with whole organisms. Colloquially, these experiments...

     activity against Trypanosoma cruzi
    Trypanosoma cruzi
    Trypanosoma cruzi is a species of parasitic euglenoid trypanosomes. This species causes the trypanosomiasis diseases in humans and animals in America...

    .

    Fourteen anthocyanins from V. yedoensis and V. prionantha have been identified. Some anthocyanins show strong antioxidant activities.

    Perfume

    Viola odorata is used as a source for scents in the perfume industry. Violet is known to have a 'flirty' scent as its fragrance comes and goes. Ionone
    Ionone
    The ionones are a series of closely related chemical substances that are part of a group of compounds known as rose ketones, which also includes damascones and damascenones. Ionones are aroma compounds found in a variety of essential oils, including rose oil...

     is present in the flowers, which turns off the ability for humans to smell the fragrant compound for moments at a time.

    Uses by other species

    Viola species are used as food plants by the larva
    Larva
    A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

    e of some Lepidoptera
    Lepidoptera
    Lepidoptera is a large order of insects that includes moths and butterflies . It is one of the most widespread and widely recognizable insect orders in the world, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...

     species, including the Giant Leopard Moth
    Giant Leopard Moth
    The Giant Leopard Moth or Eyed Tiger Moth is a moth of the family Arctiidae. It is distributed throughout the Southern and Eastern United States from New England to Mexico. The obsolete name Ecpantheria scribonia is still occasionally encountered.This species has a wingspan of 3 inches...

    , Large Yellow Underwing
    Large Yellow Underwing
    The Large Yellow Underwing is a moth, the type species for the family Noctuidae. It is an abundant species throughout Europe, one of the most common and most familiar moths of the region. In some years the species is highly migratory with large numbers appearing suddenly in marginal parts of the...

    , Lesser Broad-bordered Yellow Underwing
    Lesser Broad-bordered Yellow Underwing
    The Lesser Broad-bordered Yellow Underwing is a moth of the family Noctuidae. It is distributed throughout southern and central Europe, and southern Sweden....

    , High Brown Fritillary
    High Brown Fritillary
    The High Brown Fritillary is a butterfly of the Nymphalidae family, native from Europe across mainland Asia to Japan.The adults fly in July/August and lay eggs near to the larval food plants which are species of violets,...

    , Small Pearl-bordered Fritillary
    Small Pearl-bordered Fritillary
    The Small Pearl-bordered Fritillary , called the Silver-bordered Fritillary in North America, is a butterfly of the Nymphalidae family...

    , Pearl-bordered fritillary
    Pearl-bordered Fritillary
    The Pearl-bordered Fritillary is a butterfly of the Nymphalidae family.It is orange with black spots on the upperside of its wing and has a wingspan of 38–46 mm. On the underside of the wings there is a row of silver pearly markings along the edge, which give the species its name...

     and Setaceous Hebrew Character
    Setaceous Hebrew Character
    The Setaceous Hebrew Character is a moth of the family Noctuidae. It is found in the Palearctic ecozone. It is a common species throughout Europe. It is also found in North America, from coast to coast across Canada and the northern United States to western Alaska. It occurs in the Rocky Mountains...

    .

    External references

    1. ITIS (Accessed December 2, 2002)
    2. The Oxford Companion to Food, by Alan Davidson, Oxford University Press
      Oxford University Press
      Oxford University Press is the largest university press in the world. It is a department of the University of Oxford and is governed by a group of 15 academics appointed by the Vice-Chancellor known as the Delegates of the Press. They are headed by the Secretary to the Delegates, who serves as...

      . ISBN 0-19-211579-0
    3. Larousse Gastronomique, by Prosper Montagne (Ed.), Clarkson Potter, 2001. ISBN 0-609-60971-8

    External links

    • Violaceae in Topwalks
    • Viola charlestonensis type sheet from Louis-Marie herbarium (Laval University; Isotype, holotype
      Holotype
      A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

       is at University of California
      University of California
      The University of California is a public university system in the U.S. state of California. Under the California Master Plan for Higher Education, the University of California is a part of the state's three-tier public higher education system, which also includes the California State University...

      ).
    • Images of Japanese Viola Flavon's art gallery
    • The American Violet Society
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