Castanea mollissima
Encyclopedia
Castanea mollissima is a member of the family Fagaceae
Fagaceae
The family Fagaceae, or beech family, comprises about 900 species of both evergreen and deciduous trees and shrubs, which are characterized by alternate simple leaves with pinnate venation, unisexual flowers in the form of catkins, and fruit in the form of cup-like nuts. Fagaceous leaves are often...

, and a species of chestnut
Chestnut
Chestnut , some species called chinkapin or chinquapin, is a genus of eight or nine species of deciduous trees and shrubs in the beech family Fagaceae, native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce.-Species:The chestnut belongs to the...

 native to China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

.

Description

It is a deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 growing to 20 m tall with a broad crown. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are alternate, simple, 10–22 cm long and 4.5–8 cm broad, with a toothed margin. The flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s are produced in catkin
Catkin
A catkin or ament is a slim, cylindrical flower cluster, with inconspicuous or no petals, usually wind-pollinated but sometimes insect pollinated . They contain many, usually unisexual flowers, arranged closely along a central stem which is often drooping...

s 4–20 cm long, with the female flowers at the base of the catkin and males on the rest. The fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 is a densely spiny cupule
Cupule
The calybium and the cupule make up the accessory fruit of flowering plants in the family Fagaceae. These two parts derive from different flower components....

 4–8 cm diameter, containing two or three glossy brown nut
Nut (fruit)
A nut is a hard-shelled fruit of some plants having an indehiscent seed. While a wide variety of dried seeds and fruits are called nuts in English, only a certain number of them are considered by biologists to be true nuts...

s; these are 2–3 cm diameter on wild trees. The scientific name mollissima derives from the softly downy shoots and young leaves.

Taxonomy

Synonyms:Castanea bungeana Blume; C. duclouxii Dode; C. fargesii Dode; C. formosana (Hayata) Hayata; C. hupehensis Dode; C. mollissima var. pendula X. Y. Zhou & Z. D. Zhou; C. sativa Miller var. formosana Hayata; C. sativa var. mollissima (Blume) Pampanini; C. vulgaris Lamarck var. yunnanensis Franchet.

In Vietnam
Vietnam
Vietnam – sometimes spelled Viet Nam , officially the Socialist Republic of Vietnam – is the easternmost country on the Indochina Peninsula in Southeast Asia. It is bordered by China to the north, Laos to the northwest, Cambodia to the southwest, and the South China Sea –...

, Chinese chestnut (Vietnamese language
Vietnamese language
Vietnamese is the national and official language of Vietnam. It is the mother tongue of 86% of Vietnam's population, and of about three million overseas Vietnamese. It is also spoken as a second language by many ethnic minorities of Vietnam...

: hạt dẻ, Tày language
Tày language
Tày or Tho is the major Tai language of Vietnam, in the northeast near the Chinese border....

: mác lịch) which are grown in Trùng Khánh
Trung Khanh
Trung Khanh is a district of Cao Bang Province in the Northeast region of Vietnam.As of 2003 the district had a population of 50,189. The district covers an area of 469 km²...

 district, Cao Bằng province (Castanea mollissima Bl.) have highest quality with 3,3-5,4% glucose
Glucose
Glucose is a simple sugar and an important carbohydrate in biology. Cells use it as the primary source of energy and a metabolic intermediate...

, 43,36- 46,47% glucid
Carbohydrate
A carbohydrate is an organic compound with the empirical formula ; that is, consists only of carbon, hydrogen, and oxygen, with a hydrogen:oxygen atom ratio of 2:1 . However, there are exceptions to this. One common example would be deoxyribose, a component of DNA, which has the empirical...

, 1,16 – 2% lipid
Lipid
Lipids constitute a broad group of naturally occurring molecules that include fats, waxes, sterols, fat-soluble vitamins , monoglycerides, diglycerides, triglycerides, phospholipids, and others...

, 3,12 – 3,62% protein
Protein
Proteins are biochemical compounds consisting of one or more polypeptides typically folded into a globular or fibrous form, facilitating a biological function. A polypeptide is a single linear polymer chain of amino acids bonded together by peptide bonds between the carboxyl and amino groups of...

 analyzed by Vietnam National Vegetable and Fruit Researching Institution in 1999.

Distribution and habitat

In the provinces of Anhui
Anhui
Anhui is a province in the People's Republic of China. Located in eastern China across the basins of the Yangtze River and the Huai River, it borders Jiangsu to the east, Zhejiang to the southeast, Jiangxi to the south, Hubei to the southwest, Henan to the northwest, and Shandong for a tiny...

, Fujian
Fujian
' , formerly romanised as Fukien or Huguing or Foukien, is a province on the southeast coast of mainland China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, and Guangdong to the south. Taiwan lies to the east, across the Taiwan Strait...

, Gansu
Gansu
' is a province located in the northwest of the People's Republic of China.It lies between the Tibetan and Huangtu plateaus, and borders Mongolia, Inner Mongolia, and Ningxia to the north, Xinjiang and Qinghai to the west, Sichuan to the south, and Shaanxi to the east...

, Guangdong
Guangdong
Guangdong is a province on the South China Sea coast of the People's Republic of China. The province was previously often written with the alternative English name Kwangtung Province...

, Guangxi
Guangxi
Guangxi, formerly romanized Kwangsi, is a province of southern China along its border with Vietnam. In 1958, it became the Guangxi Zhuang Autonomous Region of the People's Republic of China, a region with special privileges created specifically for the Zhuang people.Guangxi's location, in...

, Guizhou
Guizhou
' is a province of the People's Republic of China located in the southwestern part of the country. Its provincial capital city is Guiyang.- History :...

, Hebei
Hebei
' is a province of the People's Republic of China in the North China region. Its one-character abbreviation is "" , named after Ji Province, a Han Dynasty province that included what is now southern Hebei...

, Henan
Henan
Henan , is a province of the People's Republic of China, located in the central part of the country. Its one-character abbreviation is "豫" , named after Yuzhou , a Han Dynasty state that included parts of Henan...

, Hubei
Hubei
' Hupeh) is a province in Central China. The name of the province means "north of the lake", referring to its position north of Lake Dongting...

, Hunan
Hunan
' is a province of South-Central China, located to the south of the middle reaches of the Yangtze River and south of Lake Dongting...

, Jiangsu
Jiangsu
' is a province of the People's Republic of China, located along the east coast of the country. The name comes from jiang, short for the city of Jiangning , and su, for the city of Suzhou. The abbreviation for this province is "苏" , the second character of its name...

, Jiangxi
Jiangxi
' is a southern province in the People's Republic of China. Spanning from the banks of the Yangtze River in the north into hillier areas in the south, it shares a border with Anhui to the north, Zhejiang to the northeast, Fujian to the east, Guangdong to the south, Hunan to the west, and Hubei to...

, Liaoning
Liaoning
' is a province of the People's Republic of China, located in the northeast of the country. Its one-character abbreviation is "辽" , a name taken from the Liao River that flows through the province. "Níng" means "peace"...

, Nei Mongol, Qinghai
Qinghai
Qinghai ; Oirat Mongolian: ; ; Salar:) is a province of the People's Republic of China, named after Qinghai Lake...

, Shaanxi
Shaanxi
' is a province in the central part of Mainland China, and it includes portions of the Loess Plateau on the middle reaches of the Yellow River in addition to the Qinling Mountains across the southern part of this province...

, Shandong
Shandong
' is a Province located on the eastern coast of the People's Republic of China. Shandong has played a major role in Chinese history from the beginning of Chinese civilization along the lower reaches of the Yellow River and served as a pivotal cultural and religious site for Taoism, Chinese...

, Shanxi
Shanxi
' is a province in Northern China. Its one-character abbreviation is "晋" , after the state of Jin that existed here during the Spring and Autumn Period....

, Sichuan
Sichuan
' , known formerly in the West by its postal map spellings of Szechwan or Szechuan is a province in Southwest China with its capital in Chengdu...

, Xizang, Yunnan
Yunnan
Yunnan is a province of the People's Republic of China, located in the far southwest of the country spanning approximately and with a population of 45.7 million . The capital of the province is Kunming. The province borders Burma, Laos, and Vietnam.Yunnan is situated in a mountainous area, with...

, and Zhejiang
Zhejiang
Zhejiang is an eastern coastal province of the People's Republic of China. The word Zhejiang was the old name of the Qiantang River, which passes through Hangzhou, the provincial capital...

, and also to Taiwan
Taiwan
Taiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...

 and Korea
Korea
Korea ) is an East Asian geographic region that is currently divided into two separate sovereign states — North Korea and South Korea. Located on the Korean Peninsula, Korea is bordered by the People's Republic of China to the northwest, Russia to the northeast, and is separated from Japan to the...

. It grows close to sea level in the north of its range, and at altitudes of up to 2,800 m in the south of the range. The species prefers full sun and acidic, loamy soil
Soil
Soil is a natural body consisting of layers of mineral constituents of variable thicknesses, which differ from the parent materials in their morphological, physical, chemical, and mineralogical characteristics...

, and has a medium growth rate.

Ecology

When cultivated close to other species of chestnut (including Japanese chestnut
Japanese Chestnut
Japanese Chestnut is a species of chestnut originally native to Japan and South Korea. It is a small to medium-sized deciduous tree growing to 10-15 m tall. The leaves are similar to those of the Sweet Chestnut, though usually a little smaller, 8-19 cm long and 3-5 cm broad...

, C. crenata; American chestnut
American Chestnut
The American Chestnut is a large, deciduous tree of the beech family native to eastern North America. Before the species was devastated by the chestnut blight, a fungal disease, it was one of the most important forest trees throughout its range...

, C. dentata; and sweet chestnut
Sweet Chestnut
Castanea sativa is a species of the flowering plant family Fagaceae, the tree and its edible seeds are referred to by several common names such Sweet Chestnut or Marron. Originally native to southeastern Europe and Asia Minor, it is now widely dispersed throughout Europe and parts of Asia, such as...

, C. sativa), Chinese chestnut readily cross-pollinates with them to form hybrids.

Chinese chestnuts have evolved over a long period of time in coexistence with the bark fungal
Fungus
A fungus is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds , as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi, which is separate from plants, animals, and bacteria...

 disease chestnut blight
Chestnut blight
The pathogenic fungus Cryphonectria parasitica is a member of the ascomycota category, and is the main cause of chestnut blight, a devastating disease of the American chestnut tree that caused a mass extinction in the early 1900s of this once plentiful tree from its historic range in the eastern...

 (Cryphonectria parasitica, formerly Endothia parasitica), and have evolved a very successful resistance to the blight, probably more so than any other species of chestnut, so that, although it is not immune, it typically sustains no more than minor damage when infected. This is in stark contrast to the American chestnut, which had no resistance to the blight, and was nearly wiped out by it after its introduction from Asia to North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

. An active program has been pursued in North America to cross-breed the Chinese and American chestnuts to try to maximize various desirable traits of the American chestnut, such as larger stature, larger leaf size, larger nut size, and greater nut sweetness, while also isolating and carrying the blight resistance from the Chinese chestnut.

Uses

The nuts are edible, and the tree is widely cultivated in eastern Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

; over 300 cultivar
Cultivar
A cultivar'Cultivar has two meanings as explained under Formal definition. When used in reference to a taxon, the word does not apply to an individual plant but to all those plants sharing the unique characteristics that define the cultivar. is a plant or group of plants selected for desirable...

s have been selected for nut production, subdivided into five major regional groups: Northern, Yangtze River
Yangtze River
The Yangtze, Yangzi or Cháng Jiāng is the longest river in Asia, and the third-longest in the world. It flows for from the glaciers on the Tibetan Plateau in Qinghai eastward across southwest, central and eastern China before emptying into the East China Sea at Shanghai. It is also one of the...

 Valley, Sichuan
Sichuan
' , known formerly in the West by its postal map spellings of Szechwan or Szechuan is a province in Southwest China with its capital in Chengdu...

 and Guizhou
Guizhou
' is a province of the People's Republic of China located in the southwestern part of the country. Its provincial capital city is Guiyang.- History :...

, Southern and Southwestern. Besides that, the Dandong chestnut (belonging to the Japanese chestnut
Japanese Chestnut
Japanese Chestnut is a species of chestnut originally native to Japan and South Korea. It is a small to medium-sized deciduous tree growing to 10-15 m tall. The leaves are similar to those of the Sweet Chestnut, though usually a little smaller, 8-19 cm long and 3-5 cm broad...

 – Castanea crenata) is a major cultivar in Liaoning
Liaoning
' is a province of the People's Republic of China, located in the northeast of the country. Its one-character abbreviation is "辽" , a name taken from the Liao River that flows through the province. "Níng" means "peace"...

 Province. Some cultivars, such as 'Kuling', 'Meiling', and 'Nanking', have large nuts up to 4 cm diameter. The nuts are sweet, and considered by some to have the best taste of any chestnut, though others state they are not as good as the American Chestnut
American Chestnut
The American Chestnut is a large, deciduous tree of the beech family native to eastern North America. Before the species was devastated by the chestnut blight, a fungal disease, it was one of the most important forest trees throughout its range...

. The nuts also provide a significant food source for wildlife
Wildlife
Wildlife includes all non-domesticated plants, animals and other organisms. Domesticating wild plant and animal species for human benefit has occurred many times all over the planet, and has a major impact on the environment, both positive and negative....

.
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