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Vaccinium

Vaccinium

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Vaccinium is a genus
Genus
In biology, a genus is a taxonomic unit used in the classification of living and fossil organisms. The term comes from Latin genus "descent, family, type, gender" , cognate with – genos, "race, stock, kin" ..In addition, genus is a taxonomic rank in the hierarchy In biology, a genus (plural:...

 of shrub
Shrub
A shrub or bush is a horticultural rather than strictly botanical category of woody plant, distinguished from a tree by its multiple stems and lower height, usually less than 5-6 m tall. A large number of plants can be either shrubs or trees, depending on the growing conditions they experience...

s or dwarf shrubs in the plant Family Ericaceae
Ericaceae
Ericaceae, the heath family or the heather family is a plant family, comprising mostly calcifuge plants that thrive in acidic soils. Many well-known plants of the Ericaceae live in temperate climates, such as cranberry, blueberry, various heaths and heathers , huckleberry, azalea and rhododendron...

. The fruit
Fruit
The term fruit has different meanings dependent on context, and the term is not synonymous in food preparation and biology. Fruits are the means by which flowering plants disseminate seeds, and the presence of seeds indicates that a structure is most likely a fruit, though not all seeds come from...

 of many species are eaten by humans and some are of commercial importance, including the cranberry
Cranberry
Cranberries are a group of evergreen dwarf shrubs or trailing vines in the genus Vaccinium subgenus Oxycoccos, or in some treatments, in the distinct genus Oxycoccos...

, blueberry
Blueberry
Blueberries are flowering plants of the genus Vaccinium with dark-blue berries. Species in the section Cyanococcus are the most common fruits sold as "blueberries" and are mainly native to North America. They are usually erect but sometimes prostrate shrubs varying in size from to tall...

, bilberry
Bilberry
Bilberry is any of several species of low-growing shrubs in the genus Vaccinium , bearing edible fruits. The species most often referred to is Vaccinium myrtillus L., but there are several other closely related species....

 or whortleberry, lingonberry or cowberry, and huckleberry. Like many other ericaceous plants, they are generally restricted to acid
Acid
An acid is traditionally considered any chemical compound that, when dissolved in water, gives a solution with a hydrogen ion activity greater than in pure water, i.e. a pH less than 7.0...

ic soils.

Distribution


The genus contains about 450 species, which are found mostly in the cooler areas of the Northern Hemisphere
Northern Hemisphere
The Northern Hemisphere is the half of a planet that is north of the equator—the word hemisphere literally means 'half sphere'. It is also that half of the celestial sphere north of the celestial equator...

, although there are tropical species from areas as widely separated as Madagascar
Madagascar
Madagascar, or Republic of Madagascar , is an island nation in the Indian Ocean off the southeastern coast of Africa. The main island, also called Madagascar, is the fourth-largest island in the world, and is home to 5% of the world's plant and animal species, of which more than 80% are endemic to...

 and Hawaii
Hawaii
Hawaii is the newest of the 50 U.S. states, and is the only state made up entirely of islands. It is located on an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of Australia. The state was admitted to the Union on August...

.

Etymology


The name vaccinium was used in classical Latin
Classical Latin
Classical Latin in simplest terms is the sociolinguistic register of the Latin language regarded by the enfranchised and empowered populations of the late Roman republic and the Roman empire as good Latin. Most writers during this time made use of it...

 for a type of berry (probably the bilberry
Bilberry
Bilberry is any of several species of low-growing shrubs in the genus Vaccinium , bearing edible fruits. The species most often referred to is Vaccinium myrtillus L., but there are several other closely related species....

 
V. myrtillus), but its ultimate derivation is obscure; it has nothing to do with vaccinum "of or pertaining to cows", but may be a corruption of Latin bacca, berry
Berry
The botanical definition of a berry is a simple fruit produced from a single ovary, such as a grape or a tomato. The berry is the most common type of fleshy fruit in which the entire ovary wall ripens into an edible pericarp. The flowers of these plants have a superior ovary formed by the fusion of...

.

Characteristics


The plants require acid
Acid
An acid is traditionally considered any chemical compound that, when dissolved in water, gives a solution with a hydrogen ion activity greater than in pure water, i.e. a pH less than 7.0...

ic soils, and as wild plants they live in habitats such as heath
Heath (habitat)
A heath or heathland is a dwarf-shrub habitat found on mainly infertile acidic soils, characterised by open, low growing woody vegetation, often dominated by plants of the Ericaceae...

, bog
Bog
A bog or mire is a wetland type that accumulates acidic peat, a deposit of dead plant material—usually mosses, but also lichens in Arctic climates....

 and acidic woodland
Woodland
Ecologically, a woodland is an area covered in trees, usually at low density, forming an open habitat, allowing sunlight to penetrate between the trees, and limiting shade. Woodland may support an understory of shrubs and herbaceous plants including grasses. Woodland may form a transition to...

. The plant structure varies between species – some trail along the ground, some are dwarf shrubs, and some are larger shrubs perhaps tall. The fruit
Fruit
The term fruit has different meanings dependent on context, and the term is not synonymous in food preparation and biology. Fruits are the means by which flowering plants disseminate seeds, and the presence of seeds indicates that a structure is most likely a fruit, though not all seeds come from...

 develops from an inferior ovary, and is a false berry
False berry
An epigynous berry or false berry is an accessory fruit found in certain plant species with an inferior ovary, distinguishing it from a true berry. In these species other parts of the flower can ripen along with the ovary, forming the false berry...

; it is usually brightly coloured, often being red or bluish with purple juice.

Food uses


Vaccinium species are used as food plants by the larva
Larva
A larva is a young form of animal with indirect development, going through or undergoing metamorphosis ....

e of a number of Lepidoptera
Lepidoptera
Lepidoptera is an order of insects that includes moths and butterflies. It is one of the most speciose orders in the class Insecta, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...

 (butterfly
Butterfly
A butterfly is an insect of the order Lepidoptera. Like all Lepidoptera, butterflies are notable for their unusual life cycle with a larval caterpillar stage, an inactive pupal stage, and a spectacular metamorphosis into a familiar and colourful winged adult form. Most species are day-flying so...

 and moth
Moth
A moth is an insect closely related to the butterfly, both being of the order Lepidoptera. The differences between butterflies and moths are more than just taxonomy. Sometimes the names "Rhopalocera" and "Heterocera" are used to formalize the popular distinction...

) species – see list of Lepidoptera that feed on Vaccinium.

Taxonomy


The taxonomy
Taxonomy
Taxonomy is the practice and science of classification. The word finds its roots in the Greek , taxis and , nomos...

 is complex, and still under investigation. A number of the Asian species are more closely related to
Agapetes than to other Vaccinium species. A second group includes most of Orthaea and Notopora, at least some of Gaylussacia
Gaylussacia
Gaylussacia is a genus of about fifty species of flowering plants in the family Ericaceae, native to the Americas, where they occur in eastern North America and in South America in the Andes and the mountains of southeastern Brazil...

(huckleberry), and a number of species from Vaccinium, such as Vaccinium crassifolium
Vaccinium crassifolium
Creeping blueberry is a species of Vaccinium native to the southeastern United States. It is evergreen with shiny dark green to bronze leaves...

. Other parts of Vaccinium form other groups, sometimes together with species of other genera.

Subgenera



A classification which predates molecular phylogeny
Molecular phylogeny
Molecular phylogenetics, also known as molecular systematics, is the use of the structure of molecules to gain information on an organism's evolutionary relationships. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree....

 divides
Vaccinium into subgenera, and several sections:
Subgenus Oxycoccus: The cranberries
Cranberry
Cranberries are a group of evergreen dwarf shrubs or trailing vines in the genus Vaccinium subgenus Oxycoccos, or in some treatments, in the distinct genus Oxycoccos...

, with slender, trailing, wiry non-woody shoots and strongly reflexed flower petals. Some botanists treat
Oxycoccus as a distinct genus.
  • Sect. Oxycoccus
    • Vaccinium macrocarpon
      Cranberry
      Cranberries are a group of evergreen dwarf shrubs or trailing vines in the genus Vaccinium subgenus Oxycoccos, or in some treatments, in the distinct genus Oxycoccos...

      (American Cranberry)
    • Vaccinium microcarpum
      Cranberry
      Cranberries are a group of evergreen dwarf shrubs or trailing vines in the genus Vaccinium subgenus Oxycoccos, or in some treatments, in the distinct genus Oxycoccos...

      (Small Cranberry)
    • Vaccinium oxycoccus
      Cranberry
      Cranberries are a group of evergreen dwarf shrubs or trailing vines in the genus Vaccinium subgenus Oxycoccos, or in some treatments, in the distinct genus Oxycoccos...

      (Common Cranberry)
  • Sect. Oxycoccoides
    • Vaccinium erythrocarpum
      Southern Mountain Cranberry
      Southern Mountain Cranberry , is a type of cranberry native to the southeastern United States. It is a deciduous shrub....

      (Southern Mountain Cranberry)

Subgenus Vaccinium: All the other species, with thicker, upright woody shoots and bell-shaped flowers.
  • Sect. Batodendron
    • Vaccinium arboreum
      Vaccinium arboreum
      Vaccinium arboreum is a species of Vaccinium native to the southeastern United States, from southern Virginia west to southeastern Missouri, and south to Florida and eastern Texas....

      (Sparkleberry) (images)
    • Vaccinium crassifolium (Creeping blueberry)
  • Sect. Brachyceratium
    • Vaccinium dependens
  • Sect. Bracteata
    • Vaccinium acrobracteatum
    • Vaccinium barandanum
    • Vaccinium bracteatum
    • Vaccinium coriaceum
    • Vaccinium cornigerum
    • Vaccinium cruentum
    • Vaccinium hooglandii
    • Vaccinium horizontale
    • Vaccinium laurifolium
    • Vaccinium lucidum
    • Vaccinium myrtoides
    • Vaccinium phillyreoides
    • Vaccinium reticulatovenosum
    • Vaccinium sparsum
    • Vaccinium varingifolium
  • Sect. Ciliata
    • Vaccinium ciliatum
    • Vaccinium oldhamii
  • Sect. Cinctosandra
    • Vaccinium exul
  • Sect. Conchophyllum
    • Vaccinium corymbodendron
    • Vaccinium delavayi
    • Vaccinium emarginatum
    • Vaccinium griffithianum
    • Vaccinium meridionale
    • Vaccinium moupinense (Himalayan Blueberry)
    • Vaccinium neilgherrense
    • Vaccinium nummularia
    • Vaccinium retusum
  • Sect. Cyanococcus (blueberries
    Blueberry
    Blueberries are flowering plants of the genus Vaccinium with dark-blue berries. Species in the section Cyanococcus are the most common fruits sold as "blueberries" and are mainly native to North America. They are usually erect but sometimes prostrate shrubs varying in size from to tall...

    )
    • Vaccinium angustifolium (Lowbush Blueberry)
    • Vaccinium boreale (Northern Blueberry)
    • Vaccinium caesariense (New Jersey Blueberry)
    • Vaccinium corymbosum (Highbush Blueberry)
    • Vaccinium darrowii
      Vaccinium darrowii
      Vaccinium darrowii is a species of Vaccinium in the blueberry group .-Habitat:...

      (Evergreen Blueberry)
    • Vaccinium elliottii
      Vaccinium elliottii
      Vaccinum elliottii is a species of Vaccinium in the blueberry group . It is native to the southeastern United States, from southeastern Virginia south to Florida, and west to Arkansas and Texas.-Growth:It is a deciduous shrub 2-4 m tall, with small, simple ovoid-acute leaves 15-30 mm long...

      (Elliott's Blueberry)
    • Vaccinium formosum
    • Vaccinium fuscatum (Black Highbush Blueberry; syn. V. atrococcum)
    • Vaccinium hirsutum
    • Vaccinium koreanum
    • Vaccinium myrsinites (Evergreen Blueberry)
    • Vaccinium myrtilloides (Canadian Blueberry)
    • Vaccinium pallidum (Dryland Blueberry) (images)
    • Vaccinium simulatum
    • Vaccinium tenellum
    • Vaccinium virgatum (Rabbiteye Blueberry; syn. V. ashei)
  • Sect. Eococcus
    • Vaccinium fragile
  • Sect. Epigynium
    • Vaccinium vacciniaceum
  • Sect. Galeopetalum
    • Vaccinium chunii
    • Vaccinium dunalianum
    • Vaccinium glaucoalbum
    • Vaccinium urceolatum
  • Sect. Hemimyrtillus
    • Vaccinium arctostaphylos
    • Vaccinium cylindraceum
    • Vaccinium hirtum
    • Vaccinium padifolium
    • Vaccinium smallii
  • Sect. Myrtillus
    • Vaccinium calycinum Sm.
      James Edward Smith
      Sir James Edward Smith was an English botanist and founder of the Linnean Society.Smith was born in Norwich in 1759, the son of a wealthy wool merchant. He displayed a precocious interest in the natural world...

      - Ōhelo kau laau (Hawaii
      Hawaii
      Hawaii is the newest of the 50 U.S. states, and is the only state made up entirely of islands. It is located on an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of Australia. The state was admitted to the Union on August...

      )
    • Vaccinium cespitosum
      Vaccinium cespitosum
      Vaccinium cespitosum, , the dwarf bilberry, is a species of flowering shrub in the genus which includes blueberries and cranberries. It is a low-lying plant rarely reaching half a meter in height which forms a carpetlike stand in rocky mountainous meadows. It is found across Canada and into the...

      (Dwarf bilberry)
    • Vaccinium deliciosum (Cascade Bilberry)
    • Vaccinium dentatum Sm.
      James Edward Smith
      Sir James Edward Smith was an English botanist and founder of the Linnean Society.Smith was born in Norwich in 1759, the son of a wealthy wool merchant. He displayed a precocious interest in the natural world...

      - Ōhelo (Hawaii)
    • Vaccinium membranaceum
    • Vaccinium myrtillus
      Vaccinium myrtillus
      Vaccinium myrtillus is a species of bilberry....

      (Bilberry)
    • Vaccinium ovalifolium
      Vaccinium ovalifolium
      Vaccinium ovalifolium is a plant found in coastal forests throughout southern Alaska and the Aleutian Islands.-Growth:...

      (Alaska Blueberry; syn. V. alaskaense)
    • Vaccinium parvifolium
      Vaccinium parvifolium
      Red Huckleberry is a species of Vaccinium native to the Pacific Northwest of North America, where it is common in forests from southeastern Alaska and British Columbia south through Washington and Oregon to central California...

      (Red Huckleberry)
    • Vaccinium praestans - Krasnika in Russian
    • Vaccinium reticulatum
      Vaccinium reticulatum
      The Ōhelo ai is a species of flowering plant in the cranberry family, Ericaceae, that is endemic to Hawaii. It where it grows at altitudes of on lava flows and freshly disturbed volcanic ash on Maui and Hawaii, and less commonly on Kauai, Oahu, and Molokai...

      Sm.
      James Edward Smith
      Sir James Edward Smith was an English botanist and founder of the Linnean Society.Smith was born in Norwich in 1759, the son of a wealthy wool merchant. He displayed a precocious interest in the natural world...

      - Ōhelo ai (Hawaii)
    • Vaccinium scoparium (Grouse Whortleberry)
  • Sect. Neurodesia
    • Vaccinium crenatum
  • Sect. Oarianthe
    • Vaccinium ambyandrum
    • Vaccinium cyclopense
  • Sect. Oreades
    • Vaccinium poasanum
  • Sect. Pachyanthum
    • Vaccinium fissiflorum
  • Sect. Polycodium
    • Vaccinium stamineum
      Vaccinium stamineum
      Vaccinium stamineum, commonly known as deerberry, squaw huckleberry or gooseberry, is a flowering shrub in the heath family. The plant is native to eastern North America from Ontario in the north, south to Florida and west to Texas...

      L.
      Carolus Linnaeus
      Carl Linnaeus was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature...

      - Deerberry; syn.
      V. caesium (Eastern North America) (images)
  • Sect. Pyxothamnus
    • Vaccinium consanguineum
    • Vaccinium floribundum
    • Vaccinium ovatum
      Vaccinium ovatum
      Vaccinium ovatum is a species of flowering shrub known by the common names evergreen huckleberry and California huckleberry.-Distribution and ecology:...

      Pursh
      Frederick Traugott Pursh
      Frederick Traugott Pursh was a German-American botanist. Born in Grossenhain, Saxony, to the name Friedrich Traugott Pursh, he was educated at Dresden Botanical Gardens, and emigrated to the United States in 1799...

      - California Huckleberry (Northwestern North America)
  • Sect. Vaccinium
    • Vaccinium uliginosum
      Vaccinium uliginosum
      Vaccinium uliginosum is a flowering plant in the genus Vaccinium.It is native to cool temperate regions of the Northern Hemisphere, at low altitudes in the Arctic, and at high altitudes south to the Pyrenees, the Alps, and the Caucasus in Europe, the mountains of Mongolia, northern China and...

      L.
      Carolus Linnaeus
      Carl Linnaeus was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature...

      - Northern (or Bog) Bilberry; syn.
      V. occidentale (Northern North America and Eurasia)
  • Sect. Vitis-idaea
    • Vaccinium vitis-idaea
      Vaccinium vitis-idaea
      The Vaccinium vitis-idaea – often called lingonberry and also called cowberry , foxberry, mountain cranberry, red whortleberry, lowbush cranberry, mountain bilberry, partridgeberry , and redberry – is a small evergreen shrub in the flowering plant family Ericaceae that...

       L.
      Carolus Linnaeus
      Carl Linnaeus was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature...

      - Cowberry, Lingonberry (Northern North America and Eurasia)

Production




Production tonnes. Figures 2003-2004
FAOSTAT data (FAO
Food and Agriculture Organization
The is a specialised agency of the United Nations that leads international efforts to defeat hunger. Serving both developed and developing countries, FAO acts as a neutral forum where all nations meet as equals to negotiate agreements and debate policy...

)
United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

280,503 80 % 270,000 78 %
Canada
Canada
Canada is a country occupying most of northern North America, extending from the Atlantic Ocean in the east to the Pacific Ocean in the west and northward into the Arctic Ocean...

52,651 15 % 53,400 16 %
Belarus
Belarus
Belarus is a landlocked country in Eastern Europe, bordered by Russia to the north and east, Ukraine to the south, Poland to the west, and Lithuania and Latvia to the north. Its capital is Minsk; other major cities include Brest, Grodno , Gomel , Mahilyow and Vitebsk...

8,000 2 % 10,000 3 %
Latvia
Latvia
Latvia , officially the Republic of Latvia is a country in the Baltic region of Northern Europe. It is bordered to the north by Estonia , to the south by Lithuania , to the east by the Russian Federation , and to the southeast by Belarus . Across the Baltic Sea to the west lies Sweden...

8,000 2 % 8,000 2 %
Azerbaijan
Azerbaijan
Azerbaijan , formally the Republic of Azerbaijan , is a country in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded by the Caspian Sea to the east, Russia to the north, Georgia to the northwest, Armenia to the west, and Iran to the south...

2,000 1 % 1,500 0 %
Ukraine
Ukraine
Ukraine is a country in Eastern Europe. It is bordered by Russia to the east; Belarus to the north; Poland, Slovakia, and Hungary to the west; Romania and Moldova to the southwest; and the Black Sea and Sea of Azov to the south. The city of Kiev is both the capital and the largest city of...

1,000 0 % 1,000 0 %
Tunisia
Tunisia
Tunisia , officially the Tunisian Republic , is a country located in North Africa. It is bordered by Algeria to the west and Libya to the southeast. Tunisia is located southwest of the island of Sicily and south of Sardinia. Its size is almost 165,000 km² with an estimated population of just...

50 0 % 50 0 %
Turkey
Turkey
Turkey , known officially as the Republic of Turkey , is a Eurasian country that stretches across the Anatolian peninsula in Western Asia and Thrace in the Balkan region of southeastern Europe...

50 0 % 50 0 %
Total 352 254 100 % 344 000 100 %

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