Vaccinium membranaceum
Encyclopedia

Geographic Distribution

Vaccinium membranaceum is a species within the group of Vaccinium commonly referred to as huckleberry
Huckleberry
Huckleberry is a common name used in North America for several species of plants in two closely related genera in the family Ericaceae:* Vaccinium* GaylussaciaHuckleberry may also refer to:-Plants:...

. This particular species is known by the common names thinleaf huckleberry, tall huckleberry, big huckleberry, mountain huckleberry, square-twig blueberry, and (ambiguously) as "black huckleberry". It is native to western North America, with a range extending in the northern from southern Alaska
Alaska
Alaska is the largest state in the United States by area. It is situated in the northwest extremity of the North American continent, with Canada to the east, the Arctic Ocean to the north, and the Pacific Ocean to the west and south, with Russia further west across the Bering Strait...

 and northwestern Canada to the northern mountains of California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

 to the south. It can be found from the mountains next to the Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

 in the west, to the Rocky Mountains
Rocky Mountains
The Rocky Mountains are a major mountain range in western North America. The Rocky Mountains stretch more than from the northernmost part of British Columbia, in western Canada, to New Mexico, in the southwestern United States...

 in the east. Isolated populations of this species have been found in Arizona
Arizona
Arizona ; is a state located in the southwestern region of the United States. It is also part of the western United States and the mountain west. The capital and largest city is Phoenix...

, North Dakota
North Dakota
North Dakota is a state located in the Midwestern region of the United States of America, along the Canadian border. The state is bordered by Canada to the north, Minnesota to the east, South Dakota to the south and Montana to the west. North Dakota is the 19th-largest state by area in the U.S....

, Minnesota
Minnesota
Minnesota is a U.S. state located in the Midwestern United States. The twelfth largest state of the U.S., it is the twenty-first most populous, with 5.3 million residents. Minnesota was carved out of the eastern half of the Minnesota Territory and admitted to the Union as the thirty-second state...

, the Upper Peninsula of Michigan
Upper Peninsula of Michigan
The Upper Peninsula of Michigan is the northern of the two major land masses that make up the U.S. state of Michigan. It is commonly referred to as the Upper Peninsula, the U.P., or Upper Michigan. It is also known as the land "above the Bridge" linking the two peninsulas. The peninsula is bounded...

, and Ontario
Ontario
Ontario is a province of Canada, located in east-central Canada. It is Canada's most populous province and second largest in total area. It is home to the nation's most populous city, Toronto, and the nation's capital, Ottawa....

, Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

 .

Ecology & Physiology

Vaccinium membranaceum grows at higher elevations in subalpine
Subalpine
The subalpine zone is the biotic zone immediately below tree line around the world. Species that occur in this zone depend on the location of the zone on the Earth, for example, Snow Gum in Australia, or Subalpine Larch, Mountain Hemlock and Subalpine Fir in western North America.Trees in the...

 and alpine
Alpine
The term alpine refers to the Alps, a European mountain range.Alpine may also refer to:- Geography and geology :* Alpine tundra, the biological community that occurs above the tree line at high altitude...

 environments. It occurs in both pine
Pine
Pines are trees in the genus Pinus ,in the family Pinaceae. They make up the monotypic subfamily Pinoideae. There are about 115 species of pine, although different authorities accept between 105 and 125 species.-Etymology:...

 and spruce
Spruce
A spruce is a tree of the genus Picea , a genus of about 35 species of coniferous evergreen trees in the Family Pinaceae, found in the northern temperate and boreal regions of the earth. Spruces are large trees, from tall when mature, and can be distinguished by their whorled branches and conical...

 dominated forests and in open meadow ecosystems. In forests V. membranaceumoften dominates the forest understory
Understory
Understory is the term for the area of a forest which grows at the lowest height level below the forest canopy. Plants in the understory consist of a mixture of seedlings and saplings of canopy trees together with understory shrubs and herbs...

 during early to mid stages of succession
Succession
Succession is the act or process of following in order or sequence. It may further refer to:*Order of succession, in politics, the ascension to power by one ruler, official, or monarch after the death, resignation, or removal from office of another, usually in a clearly defined order*Succession...

. Vaccinium membranceum is fire adapted. The leaves and stems of the huckleberry are resistant to low-intensity fires, and if burned away they will resprout vigorously from rhizomes buried under the soil.

Vaccinium membranaceum is an erect shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

 growing up to 1.5 meters in maximum height. The new twigs are yellow-green and somewhat angled. The deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 leaves are alternately arranged. The very thin to membranous, oval leaf blades are up to 5 centimeters long. The edges are serrated, with each tiny tooth tipped with a glandular hair. Solitary flowers occur in the leaf axils. Each is around 6 millimeters long, urn
Urn
An urn is a vase, ordinarily covered, that usually has a narrowed neck above a footed pedestal. "Knife urns" placed on pedestals flanking a dining-room sideboard were an English innovation for high-style dining rooms of the late 1760s...

-shaped to cylindrical, and pale pink to waxy bronze in color. They are pollinated
Pollination
Pollination is the process by which pollen is transferred in plants, thereby enabling fertilisation and sexual reproduction. Pollen grains transport the male gametes to where the female gamete are contained within the carpel; in gymnosperms the pollen is directly applied to the ovule itself...

 by bee
Bee
Bees are flying insects closely related to wasps and ants, and are known for their role in pollination and for producing honey and beeswax. Bees are a monophyletic lineage within the superfamily Apoidea, presently classified by the unranked taxon name Anthophila...

s. The fruit mature fruit ranges in colour from red through bluish-purple to a dark, almost black berry
Berry
The botanical definition of a berry is a fleshy fruit produced from a single ovary. Grapes are an example. The berry is the most common type of fleshy fruit in which the entire ovary wall ripens into an edible pericarp. They may have one or more carpels with a thin covering and fleshy interiors....

 about a centimeter wide. Each fruit contains an average of 47 tiny seeds. The plant rarely reproduces via seed, rather, it usually spreads by cloning
Vegetative reproduction
Vegetative reproduction is a form of asexual reproduction in plants. It is a process by which new individuals arise without production of seeds or spores...

 itself from its rhizome
Rhizome
In botany and dendrology, a rhizome is a characteristically horizontal stem of a plant that is usually found underground, often sending out roots and shoots from its nodes...

 or shoots. The seeds do germinate
Germination
Germination is the process in which a plant or fungus emerges from a seed or spore, respectively, and begins growth. The most common example of germination is the sprouting of a seedling from a seed of an angiosperm or gymnosperm. However the growth of a sporeling from a spore, for example the...

 if dispersed
Biological dispersal
Biological dispersal refers to species movement away from an existing population or away from the parent organism. Through simply moving from one habitat patch to another, the dispersal of an individual has consequences not only for individual fitness, but also for population dynamics, population...

 by animals, however, as evidenced by populations of the plant growing on the recovering section of Mount St. Helens
Mount St. Helens
Mount St. Helens is an active stratovolcano located in Skamania County, Washington, in the Pacific Northwest region of the United States. It is south of Seattle, Washington and northeast of Portland, Oregon. Mount St. Helens takes its English name from the British diplomat Lord St Helens, a...

. Other than the study by Yang et al. (2008) reports of V. membranaceum sprouting from seed are quite rare with other scientists who have studied this species reporting only 6 seedlings observed during 18 years in the field.

Uses

In a good year Vaccinium membranaceumshrubs produce a lot of fruit. The amount of fruit produced by these shrubs is legendary, with stories being told of mountain sides turned purple by all of the fruit, or shrubs being weighed to the ground by large, and abundant berries Both humans and wildlife enjoy feasting on this fruit in the late summer and early fall. People have been eating the fruit of this species for thousands of years. It was and continues to be widely used for food by Native Americans
Indigenous peoples of the Americas
The indigenous peoples of the Americas are the pre-Columbian inhabitants of North and South America, their descendants and other ethnic groups who are identified with those peoples. Indigenous peoples are known in Canada as Aboriginal peoples, and in the United States as Native Americans...

. Alaska Natives
Alaska Natives
Alaska Natives are the indigenous peoples of Alaska. They include: Aleut, Inuit, Tlingit, Haida, Tsimshian, Eyak, and a number of Northern Athabaskan cultures.-History:In 1912 the Alaska Native Brotherhood was founded...

 consumed it in bread and pies as a source of vitamin C
Vitamin C
Vitamin C or L-ascorbic acid or L-ascorbate is an essential nutrient for humans and certain other animal species. In living organisms ascorbate acts as an antioxidant by protecting the body against oxidative stress...

, the Coeur d'Alene people ate the fruit fresh, dried, mashed, cooked, and added it to soup or froze it for later use, and many other groups relished it and stored it frozen, dried, pressed into cakes, or canned for winter use. Vaccinium membranaceum is this species that is the most commonly collected of all of the wild western huckleberries, and it has great commercial importance.

The plant also provides a key food source for black and grizzly bear
Grizzly Bear
The grizzly bear , also known as the silvertip bear, the grizzly, or the North American brown bear, is a subspecies of brown bear that generally lives in the uplands of western North America...

s, which eat the leaves, stems, roots, and fruit. Elk
Elk
The Elk is the large deer, also called Cervus canadensis or wapiti, of North America and eastern Asia.Elk may also refer to:Other antlered mammals:...

, moose
Moose
The moose or Eurasian elk is the largest extant species in the deer family. Moose are distinguished by the palmate antlers of the males; other members of the family have antlers with a dendritic configuration...

, and white-tailed deer
White-tailed Deer
The white-tailed deer , also known as the Virginia deer or simply as the whitetail, is a medium-sized deer native to the United States , Canada, Mexico, Central America, and South America as far south as Peru...

 also browse the plant. The thickets provide cover for many species of small animals.

Management

Some Native American groups lit carefully planned controlled burn
Controlled burn
Controlled or prescribed burning, also known as hazard reduction burning or Swailing is a technique sometimes used in forest management, farming, prairie restoration or greenhouse gas abatement. Fire is a natural part of both forest and grassland ecology and controlled fire can be a tool for...

s in wild huckleberry patches to promote fruit production by eliminating competing plants and by stimulating the huckleberry to sprout and spread. Native American groups throughout the Pacific Northwest
Pacific Northwest
The Pacific Northwest is a region in northwestern North America, bounded by the Pacific Ocean to the west and, loosely, by the Rocky Mountains on the east. Definitions of the region vary and there is no commonly agreed upon boundary, even among Pacific Northwesterners. A common concept of the...

 still utilize this plant as an important cultural food and are active in its management.

Interesting Facts

The huckleberry is the official state fruit of Idaho
Idaho
Idaho is a state in the Rocky Mountain area of the United States. The state's largest city and capital is Boise. Residents are called "Idahoans". Idaho was admitted to the Union on July 3, 1890, as the 43rd state....

, with this particular species assumed to be the huckleberry in question.

External links

The source of this article is wikipedia, the free encyclopedia.  The text of this article is licensed under the GFDL.
 
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