Newly named insects

Name	Status	Authors	Age	Unit	Location	Notes	Images
Electromyrmococcus Electromyrmococcus Electromyrmococcus is an extinct genus of mealybug in the Pseudococcidae subfamily Rhizoecinae. The genus currently contains three species, all from the early Miocene, Burdigalian stage, Dominican amber deposits on the island of Hispaniola....	Valid	Williams	Burdigalian Burdigalian The Burdigalian is, in the geologic timescale, an age or stage in the early Miocene. It spans the time between 20.43 ± 0.05 Ma and 15.97 ± 0.05 Ma...  (Miocene Miocene The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene... )	Dominican amber Dominican amber Dominican amber is amber from the Dominican Republic. Resin from the extinct species Hymenaea protera is the source of Dominican amber and probably of most amber found in the tropics....	Dominican Republic
Tainosia Tainosia Tainosia is an extinct monotypic genus of planthopper in the Nogodinidae subfamily Nogodininae and at present, it contains the single species Tainosia quisqueyae...	Valid	Szwedo Stroiński	Burdigalian Burdigalian The Burdigalian is, in the geologic timescale, an age or stage in the early Miocene. It spans the time between 20.43 ± 0.05 Ma and 15.97 ± 0.05 Ma...  (Miocene Miocene The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene... )	Dominican amber Dominican amber Dominican amber is amber from the Dominican Republic. Resin from the extinct species Hymenaea protera is the source of Dominican amber and probably of most amber found in the tropics....	Dominican Republic

Newly named bony fishes

Name	Status	Authors	Age	Unit	Location	Notes	Images
Achoania Achoania Achoania is an extinct genus of primitive lobe-finned fish which lived during the Early Devonian period. Fossils have been discovered in China....	Valid	Zhu Yu Ahlberg	Early Devonian	Xitun Formation Xitun Formation The Xitun Formation is a palaeontological formation which named after the Xitun, a location in South China. This formation it includes many remains of fossilized fish of Early Devonian period ....	Mainland China	The type species is Achoania jarvikii.
Westollrhynchus Westollrhynchus Westollrhynchus is an extinct genus of prehistoric sarcopterygian or lobe-finned fish.-See also:* Sarcopterygii* List of sarcopterygians* List of prehistoric bony fish...	Valid	Schultze	Early Devonian	Bear Rock Formation	Canada	A new genus for "Speonesydrion" lehmanni.

Newly named amphibians

Name	Status	Authors	Age	Unit	Location	Notes	Images
Mesophryne Mesophryne Mesophryne beipiaoensis is an extinct species of frog, from the Cretaceous Yixian Formation of Liaoning , and the only species in the genus Mesophryne. It is known from a single specimen collected near Heitizigou, south of Beipiao, from which the specific epithet derives. The specimen has a...	Valid	Gao Shubin	Late Jurassic	Yixian Formation Yixian Formation The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...	Mainland China	The type species is Mesophryne beipiaoensis.
Nezpercius Nezpercius Nezpercius is an extinct genus of prehistoric amphibian.-See also:* Prehistoric amphibian* List of prehistoric amphibians...	Valid	Blob Carrano Rogers Forster Espinoza	Upper Cretaceous	Judith River Formation Judith River Formation The Judith River Formation is a fossil-bearing geologic formation in Montana, and is part of the Judith River Group. It dates to the upper Cretaceous, between 80 and 75 million years ago, corresponding to the "Judithian" land vertebrate age...	United States	The type species is Nezpercius dodsoni.
Sinerpeton Sinerpeton Sinerpeton is an extinct genus of prehistoric amphibian.-See also:* Prehistoric amphibian* List of prehistoric amphibians...	Valid	Gao Shubin	Late Jurassic	Zhangjiakou Formation	Mainland China	The type species is Sinerpeton fengshanensis.

Newly named turtles

Name	Status	Authors	Age	Unit	Location	Notes	Images
Caribemys Caribemys Caribemys is an extinct genus of turtle which existed in the Jurassic period. Its fossils have been found in the Jagua Formation in western Cuba. The type species of Caribemys is C. oxfordiensis.-External links:...	Valid	de la Fuente Iturralde-Vinent	Upper Jurassic	Jagua Formation Jagua Formation The Jagua Formation is a Late Jurassic geologic formation in Pinar del Rio Province, western Cuba. Plesiosaur, pliosaur, pterosaur, metriorhynchid, and turtle remains are among the fossils that have been recovered from its strata....	Cuba	The type species is Caribemys oxfordiensis.
Cearachelys Cearachelys Cearachelys is an extinct genus of pleurodiran turtle which existed some 80-million years ago. The genus is monotypic, with only type species Cearachelys placidoi known....	Valid	Gaffney Campos Hirayama	Early Cretaceous	Santana Formation Santana Formation The Santana Formation is a geologic Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, named after the village of Santana do Cariri, lies at the base of the Araripe Plateau...	Brazil	The type species is Cearachelys placidoi.
Kurmademys Kurmademys Kurmademys is an extinct genus of side-necked turtle which existed in India during the late Cretaceous period. It was first named in 2001, by Eugene S. Gaffney, Sankar Chatterjee, and Dhiraj K. Rudra, and contains the species Kurmademys kallamedensis. The species name is derived from the Kallamedu...	Valid	Gaffney Chatterjee Rudra	Late Cretaceous	Kallamedu Formation	India	The type species is Kurmademys kallamedensis.

Ichthyosaurs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Metashastasaurus	Synonymy	Nichollis Manabe	Late Triassic	Pardonet Formation	Canada	A new genus for "Shastasaurus" neoscapularis. A junior synonymy of Callawayia Callawayia Callawayia is an extinct genus of ichthyosaur. It contains the species Callawayia neoscapularis and C. altispinus.... Maisch & Matzke, 2000.

Newly named basal lepidosauromorphs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Hypuronector Hypuronector Hypuronector is a genus of extinct reptile from the Triassic Period that lived in what is now New Jersey. The etymology of the name translates as "deep-tailed swimmer from the lake." A member of the Simiosauria, Hypuronector is related to the arboreal Megalancosaurus. It was a small animal,...	Valid	Colbert Olsen	Late Triassic	Lockatong Formation Lockatong Formation The Triassic Lockatong Formation is a mapped bedrock unit in Pennsylvania, New Jersey, and New York. It is named after the Lockatong Creek in Hunterdon County, New Jersey.-Description:...	United States	The type species is Hypuronector limnaios.

Newly named plesiosaurs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Hauffiosaurus Hauffiosaurus Hauffiosaurus is an extinct genus of Early Jurassic pliosaurid plesiosaur known from Holzmaden of Germany and from Yorkshire of the United Kingdom. It was first named by Frank Robin O’Keefe in 2001 and the type species is Hauffiosaurus zanoni. In 2011, two additional species were assigned to this...	Valid	O'Keefe	Lower Jurassic	Posidonia Shale Formation	Germany	The type species is Hauffiosaurus zanoni.

Newly named sphenodonts

Name	Status	Authors	Age	Unit	Location	Notes	Images
Godavarisaurus Godavarisaurus Godavarisaurus was a sphenodontid reptile from the Early Jurassic Kota Formation of Andhra Pradesh, India....	Valid	Evans Prasad Manhas	Early Jurassic	Kota Formation Kota Formation The Kota Formation is a geological formation in India. It dates back to the Hettangian-Pliensbachian.-Vertebrate paleofauna:Indeterminate thyreophoran remains geographically present in Andhra Pradesh State, India.-Invertebrate paleofauna:...	India	The type species is Godavarisaurus latefi.
Rebbanasaurus	Valid	Evans Prasad Manhas	Early Jurassic	Kota Formation Kota Formation The Kota Formation is a geological formation in India. It dates back to the Hettangian-Pliensbachian.-Vertebrate paleofauna:Indeterminate thyreophoran remains geographically present in Andhra Pradesh State, India.-Invertebrate paleofauna:...	India	The type species is Rebbanasaurus jani.

Newly named basal archosauromorphs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Yonghesuchus Yonghesuchus Yonghesuchus is an extinct genus of Late Triassic archosauriform reptile. Remains have been found from the early Late Triassic Tongchuan Formation in Shanxi, China. It is named after Yonghe County, the county where fossils were found. Currently only one species, Y. sangbiensis, is known...	Valid	Wu Liu Li	Upper Triassic	Tongchuan Formation	Mainland China	The type species is Yonghesuchus sangbiensis.

Newly named crurotarsans

Name	Status	Authors	Age	Unit	Location	Notes	Images
Pabwehshi Pabwehshi Pabwehshi is an extinct genus of mesoeucrocodylian. It is based on GSP-UM 2000, a partial snout and corresponding lower jaw elements, with another snout assigned to it...	Valid	Wilson Malkani Gingerich	Upper Cretaceous	Pab Formation Pab Formation The Pab Formation is a Mesozoic geologic formation in Balochistan, in western Pakistan. Dinosaur remains are among the fossils that have been recovered from the formation.-Paleofauna:* Vitakridrinda* Pakisaurus* Balochisaurus* Brohisaurus...	Pakistan	The type species is Pabwehshi pakistanensis.
Rugosuchus Rugosuchus Rugosuchus is an extinct genus of neosuchian crocodyliform from the late Early Cretaceous of China. It is known from most of a skull, a partial postcranial skeleton, and a second partial skeleton including part of the hips. It was described by Wu and colleagues in 2001, with R...	Valid	Wu Wu Russell	Lower Cretaceous	?Nenjiang Formation	Mainland China	The type species is Rugosuchus nonganensis.
Stratiotosuchus Stratiotosuchus Stratiotosuchus is an extinct genus of baurusuchid mesoeucrocodylian which existed in the Adamantina Formation, Brazil during the late Cretaceous period . It was first named by Campos, D.A., Suarez, J.M., Riff, D. and Kellner, A.W.A. in 2001 and the type species is Stratiotosuchus maxhechti...	Valid	Campos Suarez Riff Kellner	Upper Cretaceous	Adamantina Formation Adamantina Formation The Adamantina Formation is a geological formation in Brazil whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation...	Brazil	The type species is Stratiotosuchus maxhechti.

Newly named dinosaurs

Data courtesy of George Olshevsky

George Olshevsky

George Olshevsky is a freelance editor, writer, publisher, amateur paleontologist, and mathematician living in San Diego, California.Olshevsky maintains the comprehensive online Dinosaur Genera List...

's dinosaur genera list and Dr. Jeremy Montague's dinosaur genus database.

Name	Status	Authors	Discovery year	Age	Unit	Location	Notes	Images
"Alashansaurus"	Nomen ex dissertatione	Daniel Chure			Ulansuhai Formation Ulansuhai Formation The Ulansuhai Formation is a geological formation in Inner Mongolia, north China. Dinosaur remains are among the fossils that have been recovered from the formation....	Mainland China	Would later be formally named Shaochilong Shaochilong Shaochilong is a genus of carcharodontosaurid dinosaur from the mid Cretaceous Ulansuhai Formation of China . The type species, S... in 2009 2009 in paleontology -Anomalocaridids:-Arachnids:-Insects:-Cephalopods:Three new species of extinct Octopoda discovered in 2009. The species - Keuppia hyperbolaris, Keuppia levante, and Styletoctopus annae - lived about 95 million years ago, and bear a strong resemblance to modern octopuses, suggesting that the... .
Aletopelta Aletopelta Aletopelta coombsi was an ankylosaurian ornithischian dinosaur whose fossils were found in Southern California.Etymologically, the generic name is composed of the Greek terms aletes and pelte, meaning, respectively "wandering" and "shield"...	Valid taxon	Ford James Kirkland			Point Loma Formation Point Loma Formation The Point Loma Formation is a sedimentary geological formation in Southern California. The strata date back to the Late Cretaceous. The formation is named after Point Loma, California.-Description:...	United States	A 20 feet (6.1 m) ankylosaurid. Apparently, before being fossilized, the animal's bloated carcass had floated out to sea and formed a miniature reef environment after it sunk to the bottom.
Bienosaurus Bienosaurus Bienosaurus was a dinosaur from the Early Jurassic. The genus was an armoured dinosaur from the Lower Lufeng Formation in Yunnan Province in China.-Discovery and species:...	Valid taxon	Dong Zhiming Dong Zhiming Dong Zhiming , from the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing, is one of China's leading paleontologists. He began working at the IVPP in 1962, learning from Yang Zhongjian who was director at the time...			Lower Lufeng Formation	Mainland China	A primitive scelidosaurid known from scant remains.
Cedarpelta Cedarpelta Cedarpelta is an extinct genus of basal ankylosaurid ankylosaur, based on material recovered from the Lower Cretaceous of North America. The skull lacks extensive cranial ornamentation, a trait which has been interpreted as plesiomorphic for ankylosaurs....	Valid taxon	Kenneth Carpenter Kenneth Carpenter Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life... James Kirkland et al.			Cedar Mountain Formation Cedar Mountain Formation The Cedar Mountain Formation is the name given to distinctive sedimentary rocks in eastern Utah that occur between the underlying Morrison Formation and overlying Naturita Formation . It is composed of non-marine sediments, that is, sediments deposited in rivers, lakes and on flood plains...	United States	The most basal known ankylosaurid.
Citipati Citipati Citipati is a genus of oviraptorid theropod dinosaur from the Late Cretaceous Period of what is now Mongolia . It is one of the best-known oviraptorids, thanks to a number of well-preserved skeletons, including several specimens found in brooding positions atop nests of eggs...	Valid taxon	James M. Clark Mark Norell Rinchen Barsbold			Djadokhta Formation	Mongolia	A relatively large oviraptorid known to brood its nests.
Draconyx Draconyx Draconyx is a genus of dinosaur from the Late Jurassic. It was an ornithopod which lived in what is now Portugal. It was found in Lourinhã, in 1991, and described by Octávio Mateus and Miguel Telles Antunes in 2001.-Etymology:...	Valid taxon	Octávio Mateus Octávio Mateus Octávio Mateus is a Portuguese dinosaur paleontologist and biologist . He received his PhD at Universidade Nova de Lisboa in 2005... Antunes			Lourinhã Formation Lourinhã Formation The Lourinhã Formation is a geological formation in West Portugal, named for the municipality of Lourinhã, from which a wide array of fossils comes...	Portugal	A relative of Camptosaurus Camptosaurus Camptosaurus is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America. The name means 'flexible lizard', .... .
Eotyrannus Eotyrannus Eotyrannus was a genus of tyrannosauroid theropod dinosaur hailing from the Early Cretaceous Wessex Formation beds, included in Wealden Group, located in the southwest coast of the Isle of Wight, United Kingdom...	Valid taxon	Hutt Naish et al.			Wessex Formation Wessex Formation The Wessex Formation is an English fossil site and geological formation that dates to the Barremian stage of the Early Cretaceous. It is part of the Wealden Group and underlies the younger Vectis Formation.-Invertebrates:...		A twenty foot tyrannosauroid.
Eshanosaurus Eshanosaurus Eshanosaurus is a possible genus of therizinosaurian dinosaur from the early Jurassic Period. It is known only from a partial lower jawbone. The fossil was found in China...	Valid taxon	Xu Xing Zhao Xijin Zhao Xijin Zhao Xijin is a Chinese paleontologist. He is notable for having named numerous dinosaurs. He is currently a professor at Beijing's Institute of Vertebrate Paleontology and Paleoanthropology.-Career:... James M. Clark			Lower Lufeng Formation	Mainland China	A therizinosauroid and possibly the earliest known coelurosaur.
Gobisaurus Gobisaurus Gobisaurus is an extinct genus of basal ankylosaurid ankylosaur from the Upper Cretaceous Ulansuhai Formation of China. The holotype consists of a skull and as yet undescribed postcranial remains. It was first named by Matthew K. Vickaryous, Anthony P. Russell, Philip J...	Valid taxon	Matthew K. Vickaryous Anthony P. Russell et al.			Ulansuhai Formation Ulansuhai Formation The Ulansuhai Formation is a geological formation in Inner Mongolia, north China. Dinosaur remains are among the fossils that have been recovered from the formation....	Mainland China	An ankylosaurid that resembled Shamosaurus Shamosaurus Shamosaurus is an extinct genus of basal ankylosaurid ankylosaur from Early Cretaceous deposits of Höövör, Mongolia. Shamosaurus is known from the holotype PIN N 3779/2 complete skull and jaw and the paratypes include partial skeleton remains and an armor. It was collected from the... .
“Hanwulosaurus Hanwulosaurus "Hanwulosaurus" is the informal name given to an as-yet undescribed genus of dinosaur from the Late Cretaceous. It was an ankylosaur around 9 meters long, which is long for an ankylosaur. Its fossils were found in Inner Mongolia, China... ”	Nomen nudum Nomen nudum The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...	Anonymous Anonymity Anonymity is derived from the Greek word ἀνωνυμία, anonymia, meaning "without a name" or "namelessness". In colloquial use, anonymity typically refers to the state of an individual's personal identity, or personally identifiable information, being publicly unknown.There are many reasons why a...				Mainland China	A thirty foot ankylosaur.
"Heilongjiangosaurus Heilongjiangosaurus "Heilongjiangosaurus" is the informal name given to an as-yet undescribed genus of duckbilled dinosaur from the Late Cretaceous. It possibly was a lambeosaurine, and may in fact be the same animal as Charonosaurus. The fossils were found in Maastrichtian-age rocks in Heilongjiang, China... "	Nomen nudum Nomen nudum The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...	Li W. Jin J.				Mainland China	A hadrosaur, possibly synonymous with Charonosaurus Charonosaurus Charonosaurus |Charon]]'s lizard") is the name of a genus of dinosaur whose fossils were discovered by Godefroit, Zan & Jin in 2000 on the south bank of the Amur River, dividing China from Russia.-Description:... .
Hesperosaurus Hesperosaurus Hesperosaurus was a herbivorous dinosaur from the Kimmeridgian to Tithonian epochs of the Jurassic period , whose fossils are found in the state of Wyoming in the United States of America...	Valid taxon	Kenneth Carpenter Kenneth Carpenter Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life... Miles Cloward			Morrison Formation Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone and limestone and is light grey, greenish...	United States	A stegosaurid slightly older and more primitive than Stegosaurus Stegosaurus Stegosaurus is a genus of armored stegosaurid dinosaur. They lived during the Late Jurassic period , some 155 to 150 million years ago in what is now western North America. In 2006, a specimen of Stegosaurus was announced from Portugal, showing that they were present in Europe as well... , although the genera may be synonymous.
Jiangshanosaurus Jiangshanosaurus Jiangshanosaurus was a titanosaurian sauropod that lived approximately 105 million years ago, during the Albian stage of the Early Cretaceous. Its remains were discovered in Lixian Village, Jiangshan county, in the eastern Chinese province of Zhejiang. The type and only known species,...	Valid taxon	Tang F. Kang et al.				Mainland China	A titanosaur known only from a partial skeleton found near Lixian Village, China.
Jinzhousaurus Jinzhousaurus Jinzhousaurus is a genus of hadrosauroid dinosaur of the Early Cretaceous of China.The type species is Jinzhousaurus yangi. The generic name refers to the town Jinzhou. The specific name honours Yang Zhongjian as the founder of Chinese paleontology...	Valid taxon	Wang X. Xu Xing			Yixian Formation Yixian Formation The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...	Mainland China	A hadrosauroid known from a nearly complete skeleton.
Khaan Khaan Khaan was an oviraptorid dinosaur that was found in the Djadochta Formation of Mongolia and lived in the Late Cretaceous Period , 75 million years ago....	Valid taxon	James M. Clark Mark Norell Rinchen Barsbold			Djadochta Formation Djadochta Formation The Djadochta Formation is situated in central Asia and dates from the Late Cretaceous Period. Laid down in the early Campanian, possibly starting in the latest Santonian, it is dated somewhat uncertainly at about 84-75 mya...	Mongolia	A fairly typical oviraptorid once misidentified as Ingenia Ingenia "Ingenia" is a genus of oviraptorid theropod dinosaurs, with one known species, "Ingenia" yanshini. Fossils have been found in several Late Cretaceous-age formations of Mongolia, most prominently in the Bugin Tsav beds of the Nemegt Formation, which has also yielded the holotype of Mononykus... .
"Kittysaurus"	Junior synonym of Eotyrannus Eotyrannus Eotyrannus was a genus of tyrannosauroid theropod dinosaur hailing from the Early Cretaceous Wessex Formation beds, included in Wealden Group, located in the southwest coast of the Isle of Wight, United Kingdom...	Hargreaves			Wessex Formation Wessex Formation The Wessex Formation is an English fossil site and geological formation that dates to the Barremian stage of the Early Cretaceous. It is part of the Wealden Group and underlies the younger Vectis Formation.-Invertebrates:...
Liaoningosaurus Liaoningosaurus Liaoningosaurus is a very unusual ankylosaur from the Lower Cretaceous. The holotype IVPP V12560 , an essentially complete juvenile skeleton compressed on a slab, was collected from the Yixian Formation of Liaoxi, Yixian County, Liaoning Province, People's Republic of China.The type species...	Valid taxon	Xu Xing Wang X. You			Yixian Formation Yixian Formation The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...	Mainland China	A bizarre ankylosaur of uncertain classification. Known from the complete specimen of a juvenile 34 cm long, it's the smallest known ankylosaur to date.
Losillasaurus Losillasaurus Losillasaurus is a genus of sauropod dinosaur from the Jurassic-Cretaceous boundary in the southeast of Spain. The type species of the turiasaurian Losillasaurus giganteus was discovered in the Los Serranos basin in Valencia and formally described by Casanovas, Santafé and Sanz in 2001...	Valid taxon	Maria Lourdes Casanovas-Cladellas Santafe Sanz				Spain	A large turiasaur known from a partial subadult skeleton.
Masiakasaurus Masiakasaurus Masiakasaurus was a small predatory theropod dinosaur from the Late Cretaceous of Madagascar. It was named in 2001 by Scott D. Sampson, Matthew Carrano, and Catherine A. Forster. Unlike most theropods, the front teeth of Masiakasaurus projected forward instead of straight down...	Valid taxon	Sampson Carrano Forster			Maevarano Formation Maevarano Formation The Maevarano Formation is an Upper Cretaceous sedimentary rock formation found in the Mahajanga Province of northwestern Madagascar. It is most likely Maastrichtian in age, and records a seasonal, semiarid environment with rivers that had greatly varying discharges...	Madagascar	A 2 meter (about 6–7 feet) noasaurid with unusual forward-pointing teeth.
Megapnosaurus Megapnosaurus Megapnosaurus was a dinosaur of the theropod family Coelophysidae, formerly called Syntarsus , living during the Early Jurassic. It was renamed by American entomologist Dr. Michael Ivie , Polish Australian Dr...	Same as Syntarsus Syntarsus *Syntarsus is a beetle in the Colydiinae subfamily.*Megapnosaurus, a theropod dinosaur, was formerly called Syntarsus, and is occasionally still referred to as such.... .	Ivie Slipinski Wegrzynowicz					Formerly known as Syntarsus Syntarsus *Syntarsus is a beetle in the Colydiinae subfamily.*Megapnosaurus, a theropod dinosaur, was formerly called Syntarsus, and is occasionally still referred to as such.... , this new name was erected by entomologists due to synonymy between the dinosaur and a genus of beetle.
Neimongosaurus Neimongosaurus Neimongosaurus is a genus of herbivorous therizinosaur theropod dinosaur known from the Upper Cretaceous of Nei Mongol, China....	Valid taxon	Zhang X. H. Xu Xing et al.			Iren Dabasu Formation Iren Dabasu Formation The Iren Dabasu Formation is a Mesozoic geologic formation. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation.-Ornithischians:-Saurischians:-References:...	Mainland China	A therizinosaur about 2.3 meters in length.
Nothronychus Nothronychus Nothronychus is a genus of theropod dinosaur classified in the group Therizinosauria, from the Cretaceous of North America.The type species of this dinosaur, Nothronychus mckinleyi, was described by James Kirkland and Douglas G. Wolfe in 2001...	Valid taxon	James Kirkland Wolfe			Moreno Hill Formation Moreno Hill Formation The Moreno Hill Formation is a geological formation in New Mexico whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.-Dinosaurs:-References:...	United States	A therizinosaur.
Paralititan Paralititan Paralititan was a giant titanosaurian sauropod dinosaur genus discovered in coastal deposits in the Upper Cretaceous Bahariya Formation of Egypt. The fossil represents the first tetrapod reported from the Bahariya Formation since 1935. Its 1.69 meter long humerus is longer than that of any...	Valid taxon	J. B. Smith Lamanna et al.			Bahariya Formation Bahariya Formation The Bahariya Formation is a fossil bearing geologic formation dating back to the Cenomanian, geographically located in Marsa Matruh, central Egypt.-Lepidosaurs:-Sauropods:Indeterminate sauropod remains.-Theropods:...	Egypt	A titanosaur and of the most massive dinosaurs ever discovered, with an estimated weight of 59 tonnes (65 short tons) and length of around 26 meters (85 ft).
Planicoxa Planicoxa Planicoxa is a genus of advanced iguanodontian dinosaur from the Early Cretaceous of North America. It is known from the partial skeletons of several individual specimens. Its fossils were discovered in Utah and South Dakota, United States....	Valid taxon	DiCroce Kenneth Carpenter Kenneth Carpenter Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life...			Cedar Mountain Formation Cedar Mountain Formation The Cedar Mountain Formation is the name given to distinctive sedimentary rocks in eastern Utah that occur between the underlying Morrison Formation and overlying Naturita Formation . It is composed of non-marine sediments, that is, sediments deposited in rivers, lakes and on flood plains...	United States	An advanced iguanodontian.
Pukyongosaurus Pukyongosaurus Pukyongosaurus is a genus of titanosauriform dinosaur that lived in Korea during the Early Cretaceous period. It was closely related to Euhelopus, and is known from a series of vertebrae in the neck and back.-References:...	Valid taxon	Dong Zhiming Dong Zhiming Dong Zhiming , from the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing, is one of China's leading paleontologists. He began working at the IVPP in 1962, learning from Yang Zhongjian who was director at the time... Paik Kim H. J.			Hasandong Formation Hasandong Formation The Hasandong Formation is a Mesozoic geologic formation. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation.-Paleofauna:* Chiayusaurus asianensis - "Tooth."...	South Korea	A titanosauriform related to Euhelopus Euhelopus Euhelopus is a genus of sauropod dinosaur that lived during the Early Cretaceous, between 130 and 112 million years ago . It lived in what is now Shandong Province in China. A large herbivore, it weighed approximately 15-20 tons and attained an adult length of 15m... .
Quilmesaurus Quilmesaurus Quilmesaurus is a genus of theropod dinosaur from the Patagonian Upper Cretaceous , a time when South American theropod assemblages were dominated by abelisaurids and carcharodontosaurs. However, the morphology of Quilmesaurus suggests it may have been a tetanuran...	Valid taxon	Rodolfo Coria Rodolfo Coria Dr. Rodolfo Coria is an Argentine paleontologist.He is best known for having directed the field study and co-naming of the Argentinosaurus in 1993, and the Giganotosaurus , in 1996 among other landmark South American dinosaurs...			Allen Formation Allen Formation The Allen Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous The Allen Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous The Allen Formation is a geological formation in Argentina whose strata date back to the...	Argentina	A 5–6 meter (16–20 feet) theropod known from a partial leg.
Rapetosaurus Rapetosaurus Rapetosaurus is a genus of sauropod dinosaur that lived in Madagascar from 70 to 65 million years ago, at the end of the Cretaceous Period. Only one species Rapetosaurus is a genus of sauropod dinosaur that lived in Madagascar from 70 to 65 million years ago, at the end of the Cretaceous Period....	Valid taxon	Kristina Curry-Rogers Catherine A. Forster			Maevarano Formation Maevarano Formation The Maevarano Formation is an Upper Cretaceous sedimentary rock formation found in the Mahajanga Province of northwestern Madagascar. It is most likely Maastrichtian in age, and records a seasonal, semiarid environment with rivers that had greatly varying discharges...	Madagascar	A 15 metres (49 ft) titanosaur.
Ruehleia Ruehleia Ruehleia is a genus of prosauropod dinosaur from the Late Triassic period of Germany. The type species is R. bedheimensis, described by Galton in 2001, and is named for the German paleontologist Hugo Ruehle von Lilienstern...	Valid taxon	Peter Galton Peter Galton Peter M. Galton is a British vertebrate paleontologist working in America, who has to date written or co-written about a hundred papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosauropod dinosaurs.With Robert Bakker in a joint article... .			Knollenmergel Knollenmergel The Knollenmergel is a geological formation in Germany and Switzerland. It dates back to the late Norian.-Dinosaurs:-References:* Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka : The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. ISBN 0-520-24209-2....	Germany	A prosauropod named for Hugo Ruehle von Lilienstern.
"Szechuanoraptor Szechuanoraptor "Szechuanoraptor" is the informal name given to an as yet undescribed genus of dinosaur from the Middle Jurassic . It was a theropod which lived in what is now China... "	Nomen ex dissertatione	Daniel Chure			Dashanpu Formation Dashanpu Formation The Dashanpu Formation is a Mid to Late Jurassic rock formation in China, most notable for the wealth of dinosaurs that have been excavated from the area...	Mainland China	A theropod not yet formally described. The name was coined by Daniel Chure in 2000.
Wellnhoferia Wellnhoferia Wellnhoferia is a genus of early prehistoric bird closely related to Archaeopteryx. It lived in what is now Germany, during the Late Jurassic. While Wellnhoferia was similar to Archaeopteryx, it had a shorter tail and its fourth toe was shorter than in Archaeopteryx...	Valid non-dinosaurian Bird Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...  taxon.	Andrzej Elżanowski Andrzej Elżanowski Andrzej Elżanowski is a Polish paleontologist and vertebrate zoologist specializing in bird phylogeny. Together with Peter Wellnhofer he described a coelurosaur theropod Archaeornithoides in 1992...			Solnhofen Limestone Solnhofen limestone The Solnhofen Plattenkalk is a Jurassic Konservat-Lagerstätte that preserves a rare assemblage of fossilized organisms, including highly detailed imprints of soft bodied organisms such as sea jellies...	Germany	Very similar to Archaeopteryx Archaeopteryx Archaeopteryx , sometimes referred to by its German name Urvogel , is a genus of theropod dinosaur that is closely related to birds. The name derives from the Ancient Greek meaning "ancient", and , meaning "feather" or "wing"... , which may be its senior synonym.
Venenosaurus Venenosaurus Venenosaurus named after the Poison Strip Member of the Cedar Mountain Formation in Utah, where the fossils were discovered by a Denver Museum of Natural History volunteer Tony DiCroce in 1998. Venenosaurus dicrocei was first described as a new species in 2001 by Virginia Tidwell, Kenneth...	Valid taxon	Tidwell Kenneth Carpenter Kenneth Carpenter Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life... S. Meyer			Cedar Mountain Formation Cedar Mountain Formation The Cedar Mountain Formation is the name given to distinctive sedimentary rocks in eastern Utah that occur between the underlying Morrison Formation and overlying Naturita Formation . It is composed of non-marine sediments, that is, sediments deposited in rivers, lakes and on flood plains...	United States	A relatively small (probably around 10 m (33 ft) long) titanosauriform sauropod, known from an incomplete skeleton of an adult and a juvenile. Its tail vertebrae articulated in a unique fashion that may be of evolutionary significance.
"Yibinosaurus Yibinosaurus "Yibinosaurus" is the informal name given to an as yet undescribed genus of herbivorous dinosaur from the Early Jurassic. It was a sauropod which lived in what is now Sichuan, China. The suggested "type species", "Yibinosaurus zhoui", has not been formally described yet, but the formal publication... "	Nomen nudum Nomen nudum The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...	Ouyang vide: Anonymous Anonymity Anonymity is derived from the Greek word ἀνωνυμία, anonymia, meaning "without a name" or "namelessness". In colloquial use, anonymity typically refers to the state of an individual's personal identity, or personally identifiable information, being publicly unknown.There are many reasons why a...				Mainland China	A saropod from Szechuan slated to be described by Ouyang Hui.

Laelaps trihedrodon synonymized with Allosaurus

Laelaps trihedrodon was one of the many species described by Edward Drinker Cope

Edward Drinker Cope

Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

 that he referred to his preoccupied genus Laelaps, later renamed Dryptosaurus

Dryptosaurus

Dryptosaurus was a genus of primitive tyrannosaur that lived in Eastern North America during the middle Maastrichtian stage of the Late Cretaceous period. Although largely unknown now outside of academic circles, a famous painting of the genus by Charles R...

. Although the type specimen included a partial dentary, all material except for a collection of five damaged partial tooth crowns (AMNH 5780) has been lost. The rediscovered material had lately been mistaken by Mcintosh for the type specimen in 1998

1998 in paleontology

-Amphibians:A. C. Henrici. 1998. A new pipoid anuran from the Late Jurassic Morrison Formation at Dinosaur National Monument, Utah. Journal of Vertebrate Paleontology 18:321-332-Dinosaurs:...

. However this identification was impossible because the type's teeth were described by Cope as "smooth" and having a "fine silky luster," while the AMNH 5780 teeth were thoroughly worn and had matrix

Matrix (biology)

In biology, matrix is the material between animal or plant cells, in which more specialized structures are embedded, and a specific part of the mitochondrion that is the site of oxidation of organic molecules. The internal structure of connective tissues is an extracellular matrix...

 clinging to them. Most of the type specimen's teeth were successional, but all the AMNH 5780 teeth were functional. AMNH 5780 has many features in common with Allosaurus and is probably referrable to that genus. However some of the Allosaurus-like characters of the tooth are primitive to theropods as a whole and may have been present in the less studied or poorly preserved Morrison theropod species. Consequently the synonymization

Synonym (taxonomy)

In scientific nomenclature, a synonym is a scientific name that is or was used for a taxon of organisms that also goes by a different scientific name. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies...

 of L. trihedrodon with Allosaurus is tentative, despite its high likelihood.

Carcharodontosaurus saharicus brain and inner ear examined

Hans C. E. Larsson published a description of the inner ear

Inner ear

The inner ear is the innermost part of the vertebrate ear. In mammals, it consists of the bony labyrinth, a hollow cavity in the temporal bone of the skull with a system of passages comprising two main functional parts:...

 and endocranium

Endocranium

For internal cast of the cranium, see Endocast.The endocranium in comparative anatomy is a part of the skull base in vertebrates and represent the basal, inner part of the cranium. The term is also applied to the outer layer of the dura mater in human anatomy.-Basic structure:Structurally, the...

 of Carcharodontosaurus saharicus. The C. saharicus braincase "completely encloses the endocranial region." This "high degree of ossification

Ossification

Ossification is the process of laying down new bone material by cells called osteoblasts. It is synonymous with bone tissue formation...

" made detailed analysis of its anatomy significantly easier. The C. saharicus endocast is similar to that of a related dinosaur, Allosaurus fragilis. Larsson describes the olfactory bulbs and peduncles

Cerebral peduncle

Mainly, the three common areas that give rise to the cerebral peduncles are the cortex, the spinal cord and the cerebellum. The cerebral peduncle, by most classifications, is everything in the mesencephalon except the tectum. The region includes the midbrain tegmentum, crus cerebri and pretectum...

 as lying "on approximately the same horizontal plane as the forebrain." The cephalic flexure

Cephalic flexure

The mesencephalic flexure or cephalic flexure is the first flexure, or bend, of the embryonic brain; it appears in the region of the mid-brain...

, the bend between the fore- and midbrain, has an angle of 45 degrees. The pontine flexure

Pontine flexure

The pontine flexure is a bend in the axis of the embryological central nervous system . This flexure marks the junction between the metencephalon and the myelencephalon. The division of the rhombencephalon into the metencephalon and the myelencephalon occurs at the 7th week of development...

, the bend between the mid- and hindbrain has an angle of about 40 degrees. Carcharodontosaurus

Carcharodontosaurus

Carcharodontosaurus was a gigantic carnivorous carcharodontosaurid dinosaur that lived around 100 to 93 million years ago, during the late Albian to early Cenomanian stages of the mid-Cretaceous Period...

had a large optic (II) nerve. The C. saharicus vena capitis dorsalis" drains the anterior neck muscles through a pair of long canals on the posterior surface of the endocast." Allosaurus

Allosaurus

Allosaurus is a genus of large theropod dinosaur that lived 155 to 150 million years ago during the late Jurassic period . The name Allosaurus means "different lizard". It is derived from the Greek /allos and /sauros...

, Dromaeosaurus albertensis, although in C. saharicus and Troodon

Troodon

Troodon is a genus of relatively small, bird-like dinosaur from the Late Cretaceous period . Discovered in 1855, it was among the first dinosaurs found in North America...

"the transverse sinus probably drained into a middle cerebral vein

Middle cerebral vein

The superficial middle cerebral vein begins on the lateral surface of the hemisphere, and, running along the lateral cerebral sulcus, ends in the cavernous or the sphenoparietal sinus.-Relations:It is connected:...

 that exited the brain in the ridge present on the dorsal edge of the trigeminal foramen."

The three semicircular canals of the inner ear of Carcharodontosaurus saharicus, when viewed from the side, had a subtriangular outline. This subtriangular inner ear configuration is present in Allosaurus

Allosaurus

Allosaurus is a genus of large theropod dinosaur that lived 155 to 150 million years ago during the late Jurassic period . The name Allosaurus means "different lizard". It is derived from the Greek /allos and /sauros...

, lizards, turtles, but not in birds. The pointed apex "at the junction of the anterior and posterior semicircular canals" is caused by the near linearity of the canals and closely resembles the condition of modern crocodiles. The subtriangular configuration may be the basal condition of archosauromorphs. A recess which would have held the floccular lobe of the brain projects into the area surrounded by the semicircular canals. This condition is also present in other non-avian theropods, birds, and pterosaurs. The orientation of the lagena of C. saharicus resembles the condition in crocodillians and some birds. The extent of its perilymphatic duct resembled those of Varanus, crocodillians, and birds. The crista which would have supported the secondary tympanic membrane in C. saharicus was either absent, or not preserved. This contrasts with Troodon, whose crista were ossified at least in their dorsal and ventral regions and their remaining portions either cartilaginous or too delicate to be preserved. The metotic strut of C. saharicus is reduced and medial compared to the "laterally hypertrophied" condition of non-avian maniraptors like Dromaeosaurus and Troodon, as well as primitive birds like Archaeopteryx

Archaeopteryx

Archaeopteryx , sometimes referred to by its German name Urvogel , is a genus of theropod dinosaur that is closely related to birds. The name derives from the Ancient Greek meaning "ancient", and , meaning "feather" or "wing"...

and Hesperornis

Hesperornis

Hesperornis is a genus of flightless aquatic birds that spanned the first half of the Campanian age of the Late Cretaceous period . One of the lesser-known discoveries of the paleontologist O. C. Marsh in the late 19th century Bone Wars, it was an important early find in the history of avian...

.

Newly named birds

Name	Status	Authors	Age	Unit	Location	Notes	Images
Apsaravis Apsaravis Apsaravis is a Mesozoic bird genus from the Late Cretaceous. The single known species, Apsaravis ukhaana, lived about 78 million years ago, in the Campanian age of the Cretaceous period. Its fossilized remains were found in the Camel's Humps sublocality of the Djadokhta Formation, at Ukhaa...	Valid	Norell Clarke	Late Cretaceous	Djadokhta Formation	Mongolia	The type species is Apsaravis ukhaana.
Limenavis Limenavis Limenavis is a prehistoric bird genus from the Late Cretaceous. It lived about 70 million years ago, around the Campanian-Maastrichtian boundary...	Valid	Clarke Chiappe	Late Cretaceous	Allen Formation Allen Formation The Allen Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous The Allen Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous The Allen Formation is a geological formation in Argentina whose strata date back to the...	Argentina	The type species is Limenavis patagonica.
Longipteryx Longipteryx Longipteryx is a genus of prehistoric bird which lived during the Early Cretaceous . It contains a single species, Longipteryx chaoyangensis. Its remains have been recovered from the Jiufotang Formation at Chaoyang in Liaoning Province, People's Republic of China...	Valid	Zhang Zhou Hou Gu	Early Cretaceous	Jiufotang Formation Jiufotang Formation The Jiufotang Formation is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms. . It is a member of the Jehol group. The exact age of the Jiufotang has been debated for years, with...	Mainland China	The type species is Longipteryx chaoyangensis.
Yanornis Yanornis Yanornis is an extinct genus of omnivorous Early Cretaceous birds, thought to be closely related to the common ancestor of all modern birds. One species, Yanornis martini,...	Valid	Zhou Zhang	Early Createcous	Jiufotang Formation Jiufotang Formation The Jiufotang Formation is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms. . It is a member of the Jehol group. The exact age of the Jiufotang has been debated for years, with...	Mainland China	The type species is Yanornis martini.
Yixianornis Yixianornis Yixianornis is a bird genus from the early Cretaceous period. Its remains have been found in the Jiufotang Formation at Chaoyang dated to the early Aptian age, around 120 million years ago. Only one species, Yixianornis grabaui, is known at present...	Valid	Zhou Zhang	Early Cretaceous	Jiufotang Formation Jiufotang Formation The Jiufotang Formation is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms. . It is a member of the Jehol group. The exact age of the Jiufotang has been debated for years, with...	Mainland China	The type species is Yixianornis grabaui.

Newly named pterosaurs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Haopterus Haopterus Haopterus was a genus of ornithocheirid pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China....	Valid	Wang Lü	Early Cretaceou	Yixian Formation Yixian Formation The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...	Mainland China	The type species is Haopterus gracilis.
Istiodactylus Istiodactylus Istiodactylus was a medium sized pterosaur from the Early Cretaceous period.-History and species:In 1887 Harry Govier Seeley named the remains of a fossil pelvis discovered on the Isle of Wight, Ornithodesmus cluniculus, thinking it was a bird. In 1901 he considered that it might have been a...	Valid	Howse Milner Martill	Early Cretaceous	Vectis Formation Vectis Formation The Vectis Formation is a geological formation on the Isle of Wight, England whose strata date back to the Early Cretaceous . The Vectis Formation is composed of three geological members: the Shepherds Chine member, the Barnes High Sandstone member, and the Cowleaze Chine member...	United Kingdom	A new genus for Ornithodesmus latidens.

Non-mammalian

Name	Status	Authors	Age	Unit	Location	Notes	Images
Delectosaurus Delectosaurus Delectosaurus is a genus of dicynodont from Late Permian of Russia....	Valid	Kurkin	Late Permian	Vyazniki Assemblage	Russia	A dicynodont Dicynodont Dicynodontia is a taxon of anomodont therapsids or mammal-like reptiles. Dicynodonts were small to large herbivorous animals with two tusks, hence their name, which means 'two dog tooth'... . Two species are described Delectosaurus arefjevi and D. berezhanensis.
Interpresosaurus Interpresosaurus Interpresosaurus is a genus of dicynodont from Late Permian of Russia....	Valid	Kurkin	Late Permian	Vyazniki Assemblage	Russia	A dicynodont. The type species Type species In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...  is Interpresosaurus blomi.
Lumkuia Lumkuia Lumkuia is an extinct genus of probainognathian cynodont. It is the earliest and most basal known member of Probainognathia, with fossils being found from the Lystrosaurus Assemblage Zone of the Beaufort Group in the South African Karoo Basin that date back to the early Middle Triassic...	Valid	Hopson Kitching	Middle Triassic	Burgersdorp Formation	South Africa	A cynodont Cynodont Cynodontia or cynodonts are a taxon of therapsids which first appeared in the Late Permian and were eventually distributed throughout all seven continents by the Early Triassic . This clade includes modern mammals and their extinct close relatives. They were one of the most diverse groups of... . The type species is Lumkuia fuzzi.
Malasaurus Malasaurus Malasaurus is an extinct genus of therocephalian therapsid which existed in Russia. The type species is Malasaurus germanus....	Valid	Tatarinov	Late Triassic	Vyazniki Assemblage	Russia	A therocephalian. The type species is Malasaurus germanus.
Mitredon Mitredon Mitredon is an extinct genus of cynodont which existed in Greenland during the upper Triassic period. The type species is Mitredon cromptoni....	Valid	Shapiro Jenkins	Late Triassic	Fleming Fjord Formation Fleming Fjord Formation The Fleming Fjord Formation is a geological formation in eastern Greenland.-Vertebrate fauna:...	Greenland	A cynodont. The type species is Mitredon cromptoni.
Prozostrodon Prozostrodon Prozostrodon was an advanced cynodont that was closely related to the ancestors of mammals. Its relatives gave rise to the first mammaliformes. Prozostrodon were found in the Geopark of Paleorrota, Santa Maria Formation, Brazil. This deposited in Geosciences of UFRGS....	Valid	Bonaparte Barberena	Late Triassic	Santa Maria Formation Santa Maria Formation The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It has a late Ladinian – early Carnian age , and is notable for its fossils of early dinosaurs, including the herrerasaur Staurikosaurus, the basal saurischian Teyuwasu, and the basal sauropodomorph...	Brazil	New genus for "Thrinaxodon" brasiliensis Barberena, Bonaparte & Sá Teixeira, 1987
Riograndia Riograndia Riograndia is a genus of tritheledontid cynodont from the Late Triassic of South America. The type species is R. guaibensis. Riograndia guaibensis was estimated to have been about 15 cm long and 30 g in weight. Remains have been found in the Caturrita Formation of the geopark of Paleorrota...	Valid	Bonaparte Ferigolo Ribeiro	Late Triassic	Caturrita Formation Caturrita Formation The Caturrita Formation is a rock formation found in Rio Grande do Sul, Brazil. Its sediments were deposited in the Paraná Basin. The formation has a late Carnian to early Norian age and forms part of the Santa Maria Supersequence in the upper section of the Rosário do Sul Group.-Etymology:The...	Brazil	A cynodont. The type species is Riograndia guaibensis.

Pteridophyta

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Wessiea Wessiea Wessiea is an extinct monotypic genus of fern in the Dryopteridaceae family with the sole species Wessiea yakimaensis. Wessia is known from Langhian age Miocene fossils found in Central Washington....	gen et sp nov	Valid	Pigg & Rothwell	Langhian Langhian The Langhian is, in the ICS geologic timescale, an age or stage in the middle Miocene epoch/series. It spans the time between 15.97 ± 0.05 Ma and 13.65 ± 0.05 Ma . The Langhian was a continuing warming period defined by Lorenzo Pareto in 1864, it was originally established in the Langhe area north...  (Middle Miocene)	"Ho ho" site, Grande Ronde Basalt, Columbia River Basalt Group Columbia River Basalt Group The Columbia River Basalt Group is a large igneous province that lies across parts of the Western United States. It is found in the U.S. states of Washington, Oregon, Idaho, Nevada, and California... .	United States

Conifers

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Metasequoia foxii Metasequoia foxii Metasequoia foxii is an extinct redwood species in the family Cupressaceae described from numerous fossils of varying growth stage. The species is solely known from the Paleocene sediments exposed in central Alberta, Canada...	sp nov	Valid	Stockey, Rothwell, & Falder	late Tiffanian Tiffanian The Tiffanian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology , typically set from 60,200,000 to 56,800,000 years BP lasting . It is usually considered to overlap the Selandian and Thanetian within the...	Munce’s Hill and Gao mine sites, Paskapoo Formation Paskapoo Formation The Paskapoo Formation is a stratigraphical unit of Paleocene age in the Western Canadian Sedimentary Basin.It takes the name from the Blindman River , and was first described in outcrops along the river, north of Red Deer by J. B...	Canada

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Trochodendron nastae Trochodendron nastae Trochodendron nastae is an extinct species of flowering plant in the family Trochodendraceae known from fossil leaves found in the early Eocene Ypresian stage Klondike Mountain Formation deposits of Northern Washington State. T. nastae is one of the oldest members of the genus Trochodendron, which...	sp nov	Valid	Pigg, Wehr Wesley C. Wehr Wesley Conrad Wehr was an American paleontologist and artist best known for his studies of Tertiary fossil floras in western North America, the Stonerose Interpretive Center, and as a part of the Northwest School of art.-Early life:... , & Ickert-Bond	Ypresian Ypresian In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between and , is preceded by the Thanetian age and is followed by the Eocene Lutetian age....	Tom Thumb Tuff, Klondike Mountain Formation Klondike Mountain Formation The Klondike Mountain Formation is an early Eocene geologic formation that records the sedimentation in the southern most of a string of highland subtropical/Temperate lakes in Washington state and British Columbia. The formation is best known for exceptionally well preserved plant and insect fossils...	United States

Complete author list

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.

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