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Hesperornis

 

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Hesperornis



 
 
Hesperornis is an extinct genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 of flightless aquatic bird
Bird

Birds are wing, Bipedalismal, endothermic , vertebrate animals that lay egg . There are around 10,000 living species, making them the most numerous tetrapod vertebrates....
s that lived during the Santonian
Santonian

The Santonian is a faunal stage of the Late Cretaceous epoch . It spans the time between 85.8 ? 0.7 mya and 83.5 ? 0.7 mya.According to the International Commission on Stratigraphy, its start is defied by the appearance of the Inoceramidae bivalve Cladoceramus undulatoplicatus....
 to Campanian
Campanian

The Campanian is a faunal stage on the geologic time scale occurring from 83.5 ? 0.7 annum to 70.6 ? 0.6 Ma .It is the middle stage of the Late Cretaceous epoch ....
 sub-epochs of the Late Cretaceous
Late Cretaceous

Late Cretaceous refers to the second half of the Cretaceous Period , named after the famous white chalk cliffs of southern England, which date from this time....
 (89-65 mya
Mya (unit)

In astronomy, geology, and paleontology, mya or "m.y.a." is an abbreviation for "million years ago". Like the related unit bya, mya is traditionally written in lower case....
). One of the lesser known discoveries of Marsh
Othniel Charles Marsh

Othniel Charles Marsh was one of the pre-eminent paleontologists of the 19th century, who discovered and named many fossils found in the American West....
 in the late 19th century Bone Wars
Bone Wars

The Bone Wars, also known as the "Great Dinosaur Rush", refers to a period of intense fossil speculation and discovery during the Gilded Age of American history, marked by a heated rivalry between Edward Drinker Cope and Othniel Charles Marsh ....
, it was an important early find in the history of avian paleontology. Famous locations for Hesperornis are the Late Cretaceous marine limestone
Limestone

File:Limestone Formation In Waitomo.jpgLimestone is a sedimentary rock composed largely of the mineral calcite . The deposition of limestone strata is often a by-product and indicator of biological activity in the geology record....
s from Kansas
Kansas

The State of Kansas is a Midwestern U.S. state in the Central United States of the United States of America, an area often referred to as the United States "Heartland"....
 and the marine shales from Canada
Canada

Canada is a country occupying most of northern North America, extending from the Atlantic Ocean in the east to the Pacific Ocean in the west and northward into the Arctic Ocean....
, but the genus had probably a Holarctic
Holarctic

The Holarctic ecozone refers to the habitats found throughout the northern continents of the world as a whole. This region is divided into the Palearctic, consisting of Northern Africa and all of Eurasia, with the exception of Southeast Asia and the Indian subcontinent, and the Nearctic, consisting of North America south to northern Mexico....
 distribution.

esperornis were very large birds, reaching up to 2 meters (6.5 feet) in length.






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Encyclopedia


Hesperornis is an extinct genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 of flightless aquatic bird
Bird

Birds are wing, Bipedalismal, endothermic , vertebrate animals that lay egg . There are around 10,000 living species, making them the most numerous tetrapod vertebrates....
s that lived during the Santonian
Santonian

The Santonian is a faunal stage of the Late Cretaceous epoch . It spans the time between 85.8 ? 0.7 mya and 83.5 ? 0.7 mya.According to the International Commission on Stratigraphy, its start is defied by the appearance of the Inoceramidae bivalve Cladoceramus undulatoplicatus....
 to Campanian
Campanian

The Campanian is a faunal stage on the geologic time scale occurring from 83.5 ? 0.7 annum to 70.6 ? 0.6 Ma .It is the middle stage of the Late Cretaceous epoch ....
 sub-epochs of the Late Cretaceous
Late Cretaceous

Late Cretaceous refers to the second half of the Cretaceous Period , named after the famous white chalk cliffs of southern England, which date from this time....
 (89-65 mya
Mya (unit)

In astronomy, geology, and paleontology, mya or "m.y.a." is an abbreviation for "million years ago". Like the related unit bya, mya is traditionally written in lower case....
). One of the lesser known discoveries of Marsh
Othniel Charles Marsh

Othniel Charles Marsh was one of the pre-eminent paleontologists of the 19th century, who discovered and named many fossils found in the American West....
 in the late 19th century Bone Wars
Bone Wars

The Bone Wars, also known as the "Great Dinosaur Rush", refers to a period of intense fossil speculation and discovery during the Gilded Age of American history, marked by a heated rivalry between Edward Drinker Cope and Othniel Charles Marsh ....
, it was an important early find in the history of avian paleontology. Famous locations for Hesperornis are the Late Cretaceous marine limestone
Limestone

File:Limestone Formation In Waitomo.jpgLimestone is a sedimentary rock composed largely of the mineral calcite . The deposition of limestone strata is often a by-product and indicator of biological activity in the geology record....
s from Kansas
Kansas

The State of Kansas is a Midwestern U.S. state in the Central United States of the United States of America, an area often referred to as the United States "Heartland"....
 and the marine shales from Canada
Canada

Canada is a country occupying most of northern North America, extending from the Atlantic Ocean in the east to the Pacific Ocean in the west and northward into the Arctic Ocean....
, but the genus had probably a Holarctic
Holarctic

The Holarctic ecozone refers to the habitats found throughout the northern continents of the world as a whole. This region is divided into the Palearctic, consisting of Northern Africa and all of Eurasia, with the exception of Southeast Asia and the Indian subcontinent, and the Nearctic, consisting of North America south to northern Mexico....
 distribution.

Description

Hesperornis were very large birds, reaching up to 2 meters (6.5 feet) in length. They had virtually no wing
Wing

A wing is a surface used to produce Lift for flight through the Earth's atmosphere or another gaseous or fluid medium. The wing shape is usually an airfoil....
s, swimming with powerful hind legs. Their toes were probably lobed rather than being webbed, as in today's grebe
Grebe

Grebes are members of the Podicipediformes order , a widely distributed order of freshwater diving Avess, some of which visit the sea when Bird migration and in winter....
s; like in these, the toes could rotate well, which is necessary to decrease drag in lobed feet but not in webbed ones such as in loon
Loon

The loons or divers are a group of aquatic birds found in many parts of North America and northern Eurasia. All living species of loons are members of one genus, Gavia, family , Gaviidae, and order Gaviiformes all of their own....
s, where the toes are simply folded together.

Like many other Mesozoic
Mesozoic

The Mesozoic Era is one of three Geologic time scale of the Phanerozoic eon . The division of time into eras dates back to Giovanni Arduino, in the 18th century, although his original name for the era now called the 'Mesozoic' was 'Secondary' ....
 birds such as Ichthyornis
Ichthyornis

Ichthyornis is a genus of seabird from the Late Cretaceous of North America. Its fossil remains are known from the chalks of Alberta, Alabama, Kansas, New Mexico, Saskatchewan, and Texas, in strata that were laid down in the Western Interior Seaway; some fossils from other locations like Argentina and Central Asia are sometimes referred...
, Hesperornis had teeth in its beak
Beak

The beak, bill or rostrum is an external anatomical structure of birds which, in addition to eating, is used for Personal grooming#In animals, manipulating objects, killing prey, probing for food, Courtship#Courtship in the animal kingdom and feeding their young....
 which were used to hold prey (most likely fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
). In the hesperornithiform lineage they were of a different arrangement than in any other known bird (or in non-avian theropod dinosaurs), with the teeth sitting in a longitudinal groove rather than in individual sockets, in a notable case of convergent evolution
Convergent evolution

Convergent evolution describes the acquisition of the same biological trait in unrelated lineages.The wing is a classic example of convergent evolution in action....
 with mosasaur
Mosasaur

Mosasaurs were serpentine marine reptiles. The first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1778. These ferocious marine predators are now considered to be the closest relatives of snakes, due to cladistic analysis of symptomatic similarities in jaw and skull anatomies....
s.

Of the many species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
 described in this genus - such as the famous H. regalis or the huge H. rossicus -, not all may be valid. Some, such as H. macdonaldi, are known from very few or even a single bone and cannot be properly compared against the more plentiful (but also incomplete) remains of other similar-sized and contemporary taxa. Coniornis altus (which includes H. montana) may also belong into this genus. In addition, there are some unassigned remains, such as SGU 3442 Ve02 and LO
Lund University

Lund University , located in Lund in southernmost Sweden, is one of Sweden's most prestigious universities and one of Scandinavia's largest institutions for education and research, frequently ranked among the world's top 100 universities....
 9067t and bones of an undetermined species from Tzimlyanskoe Reservoir near Rostov
Rostov

Rostov is one of the oldest types of inhabited localities in Russia in Russia and an important tourist centre of the so called Golden ring. It is located on the shores of Lake Nero in Yaroslavl Oblast....
. The small form, initially called H. gracilis, was later moved into a monotypic
Monotypic

In biology, a monotype is a alpha taxonomy group with only one biological type:In botany, a monotype is a taxon that has only one species: Ginkgo is a monotypic genus, while Ginkgoaceae is a monotypic family ....
 genus Hargeria, and ultimately placed in Parahesperornis
Parahesperornis

Parahesperornis is a prehistoric flightless bird genus from the Late Cretaceous. Its range in space and time may have been extensive, but its remains are rather few and far between, at least compared with its contemporary relatives in Hesperornis....
.

Paleobiology

Hesperornis Regalis   Project Gutenberg Etext 16474
Hesperornis hunted in the waters of such contemporary shelf seas as the North American Inland Sea, the Turgai Strait and the prehistoric North Sea
North Sea

The North Sea is a marginal sea, epeiric sea on the European continental shelf. The Dover Strait and the English Channel in the south and the Norwegian Sea in the north connect it to the Atlantic Ocean....
, which then were subtropical to tropical waters, much warmer than today. They probably fed mainly on fish, maybe also crustacean
Crustacean

Crustaceans are a large group of arthropods, comprising almost 52,000 described species , and are usually treated as a subphylum . They include various familiar animals, such as crabs, lobsters, crayfish, shrimp, krill and barnacles....
s, cephalopod
Cephalopod

The cephalopods are the mollusc class Cephalopoda characterized by bilateral symmetry, a prominent head, and a modification of the mollusk foot, a muscular hydrostat, into the form of cephalopod arms or tentacles....
s and mollusks as do the diving seabirds of today. Their teeth were helpful in dealing with slippery or hard-shelled prey.

On land, Hesperornis may or may not have been able to walk. They certainly were not able to stand upright like penguin
Penguin

Penguins are a group of Aquatic animal, flightless bird birds living almost exclusively in the Southern Hemisphere. Highly adapted for life in the water, penguins have countershading dark and white plumage, and their wings have become Flipper ....
s as in the early reconstructions. Their legs attached far at the back and sideways, with even the lower leg being tightly attached to the body (see photo of skeleton). Thus, they were limited to a clumsy hobble at best on land and would indeed have been more nimble if they moved by sliding on their belly or galumphing. Indeed, the leg skeleton of the hesperornithids was so much adapted to diving that their mode of locomotion while ashore, as well as where it laid its eggs and how it cared for its young is a matter of much speculation.

Some have even pointed out that it cannot be completely ruled out that these birds were ovoviviparous instead of incubating their eggs. In any case, young Hesperornis grew fairly quickly and continuously to adulthood, as is the case in modern birds, but not Enantiornithes
Enantiornithes

Enantiornithes is an extinct group of primitive birds. They were the most abundant and diverse Avialae of the Mesozoic. Almost all retained teeth and clawed hands, like other primitive birds....
. More young birds are known from the fossil record of the more northernly sites than from locations further south. This suggests that at least some species were migratory
Bird migration

Bird migration refers to the regular seasonal journeys undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather....
 like today's penguins which swim polewards in the summer.

Hesperornis were preyed upon by large marine carnivores. Tylosaurus proriger
Tylosaurus

Tylosaurus was a mosasaur, a large, predatory marine lizard closely related to modern monitor lizards and to snakes. Along with plesiosaurs, sharks, fish, and other Genus of mosasurs, it was a dominant predator of the Western Interior Seaway during the Late Cretaceous....
 specimen SDSMT
South Dakota School of Mines and Technology

The South Dakota School of Mines and Technology is a public institution of higher learning in Rapid City, South Dakota governed by the South Dakota Board of Regents....
 10439 contains the bones of a Hesperornis in its gut, for example.

In popular culture

Hesperornis appeared in the Episode 3 of the science fiction TV series Primeval. Here, a housewife in west London finds water spreading through the basement. She then calls a plumber. When he comes, the plumber is then killed by a frightened Hesperornis. Later a flock of these birds were shown swimming around in a tropical sea and a Hesperornis rookery was seen on the rocky shoreline. Here Hesperornis was depicted as having scales when in real life it was almost certainly covered in feathers, albeit perhaps more scale-like ones as in penguin
Penguin

Penguins are a group of Aquatic animal, flightless bird birds living almost exclusively in the Southern Hemisphere. Highly adapted for life in the water, penguins have countershading dark and white plumage, and their wings have become Flipper ....
s and loon
Loon

The loons or divers are a group of aquatic birds found in many parts of North America and northern Eurasia. All living species of loons are members of one genus, Gavia, family , Gaviidae, and order Gaviiformes all of their own....
s. It is also shown standing upright, which it wouldn't have been capable of.

Hesperornis was also shown in the last episode of the Walking with...
Walking with...

The Walking with... series is a collection of shows that are produced by the BBC and are made by Impossible Pictures. The aim of the series and specials, along with books, is to recreate extinct animals and presents them as a wildlife documentary....
 series spin-off Sea Monsters
Sea Monsters

Sea Monsters was a BBC television program which used computer-generated imagery to show past life in Earth's seas. It was made by Impossible Pictures, the creators of Walking with Dinosaurs, Walking With Beasts and Walking With Monsters....
, being frequently attacked and devoured by larger predators such as Xiphactinus
Xiphactinus

Xiphactinus was a large, 4.5 to 5 m long predatory Teleost that lived in the Western Interior Sea, over what is now the middle of North America, during the Late Cretaceous....
, Squalicorax
Squalicorax

Squalicorax is a genus of extinct Lamniformes shark known to have lived during the Cretaceous period. A fully articulated 1.9 m long fossil skeleton of Squalicorax has been found in Kansas, evidence of its presence in the Western Interior Seaway....
 and Halisaurus
Halisaurus

Halisaurus is an extinct genus of marine lizard belonging to the mosasaur family. With a length of 3-4 m it was small compared to most other mosasaurs....
.

Footnotes


External links

  • Eastern Kentucky University
    Eastern Kentucky University

    Eastern Kentucky University, commonly referred to as Eastern or by the acronym EKU by local residents, is an undergraduate and graduate teaching and research institution located in Richmond, Kentucky, United States of America....
    : . Retrieved 2007-NOV-04.
  • savageancientseas.com: . Note relalistic position of legs and toe rotation. Retrieved 2007-NOV-04.
  • UC Davis: . Digitized from KUVP PU17193. Requires Java, IFC or 3DC plugin. Retrieved 2007-NOV-04.