George's taxonomic arrangement of Banksia
Encyclopedia
Alex George
Alex George
Alexander Segger George is a Western Australian botanist. He is the authority on the plant genera Banksia and Dryandra...

's taxonomic arrangement of Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

 was the first modern-day arrangement for that genus. First published in 1981 in the classic monograph The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. is a 1981 monograph by Alex George on the taxonomy of the plant genus Banksia. Published by the Western Australian Herbarium as Nuytsia 3, it presented George's taxonomic arrangement of Banksia, the first major taxonomic revision of the genus since George Bentham published...

, it superseded the arrangement of
Bentham's taxonomic arrangement of Banksia
George Bentham's taxonomic arrangement of Banksia was published in 1870, in Volume 5 of Bentham's Flora Australiensis. A substantial improvement on the previous arrangement, it would stand for over a century. It was eventually replaced by Alex George's 1981 arrangement, published in his classic...

 George Bentham
George Bentham
George Bentham CMG FRS was an English botanist, characterized by Duane Isely as "the premier systematic botanist of the nineteenth century".- Formative years :...

, which had stood for over a hundred years. It was overturned in 1996 by Kevin Thiele
Kevin Thiele
Kevin R. Thiele is curator of the Western Australian Herbarium. His research interests include the systematics of the plant families Proteaceae, Rhamnaceae and Violaceae, and the conservation ecology of grassy woodland ecosystems...

 and Pauline Ladiges, but restored by George in 1999. A recent publication by Austin Mast
Austin Mast
Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently an associate professor within the Department of Biological Science at Florida State University , and has been director of FSU's since August 2003.One of his...

 and Kevin Thiele
Kevin Thiele
Kevin R. Thiele is curator of the Western Australian Herbarium. His research interests include the systematics of the plant families Proteaceae, Rhamnaceae and Violaceae, and the conservation ecology of grassy woodland ecosystems...

 suggests that it will soon be overturned again.

Background

Banksia is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of around 80 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 in the plant
Plant
Plants are living organisms belonging to the kingdom Plantae. Precise definitions of the kingdom vary, but as the term is used here, plants include familiar organisms such as trees, flowers, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The group is also called green plants or...

 family Proteaceae
Proteaceae
Proteaceae is a family of flowering plants distributed in the Southern Hemisphere. The family comprises about 80 genera with about 1600 species. Together with the Platanaceae and Nelumbonaceae they make up the order Proteales. Well known genera include Protea, Banksia, Embothrium, Grevillea,...

. An iconic Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

n wildflower and popular garden
Garden
A garden is a planned space, usually outdoors, set aside for the display, cultivation, and enjoyment of plants and other forms of nature. The garden can incorporate both natural and man-made materials. The most common form today is known as a residential garden, but the term garden has...

 plant
Plant
Plants are living organisms belonging to the kingdom Plantae. Precise definitions of the kingdom vary, but as the term is used here, plants include familiar organisms such as trees, flowers, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The group is also called green plants or...

, they are easily recognised by their characteristic flower spikes and fruiting "cones". They grow in forms varying from prostrate woody shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

s to trees up to 35 metres tall, and occur in all but the most arid
Arid
A region is said to be arid when it is characterized by a severe lack of available water, to the extent of hindering or even preventing the growth and development of plant and animal life...

 areas of Australia. As heavy producers of nectar
Nectar (plant)
Nectar is a sugar-rich liquid produced by plants. It is produced in glands called nectaries, either within the flowers, in which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists, which in turn provide anti-herbivore protection...

, they are important sources of food for nectariferous animals such as honeyeater
Honeyeater
The honeyeaters are a large and diverse family of small to medium sized birds most common in Australia and New Guinea, but also found in New Zealand, the Pacific islands as far east as Samoa and Tonga, and the islands to the north and west of New Guinea known as Wallacea...

s and honey possum
Honey Possum
The honey possum or tait, its Native Australian name or noolbenger is a tiny Australian marsupial weighing just seven to eleven grams for the male, and eight to sixteen grams for the female—about half the weight of a mouse. Their physical size ranges from a body length of between 6.5 –...

, and they are of economic importance to the nursery
Nursery (horticulture)
A nursery is a place where plants are propagated and grown to usable size. They include retail nurseries which sell to the general public, wholesale nurseries which sell only to businesses such as other nurseries and to commercial gardeners, and private nurseries which supply the needs of...

 and cut flower
Floristry
Floristry is the general term used to describe production, commerce and trade in flowers. It encompasses flower care and handling, floral design or flower arranging, merchandising, and display and flower delivery. Wholesale florists sell bulk flowers and related supplies to professionals in the trade...

 industries. However they are seriously threatened by a number of processes including land clearing, frequent burning, and disease, and a number of species are rare
Rare species
A rare species is a group of organisms that are very uncommon or scarce. This designation may be applied to either a plant or animal taxon, and may be distinct from the term "endangered" or "threatened species" but not "extinct"....

 and endangered
Endangered species
An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

.

Specimens of Banksia were first collected by Sir Joseph Banks
Joseph Banks
Sir Joseph Banks, 1st Baronet, GCB, PRS was an English naturalist, botanist and patron of the natural sciences. He took part in Captain James Cook's first great voyage . Banks is credited with the introduction to the Western world of eucalyptus, acacia, mimosa and the genus named after him,...

 and Dr Daniel Solander
Daniel Solander
Daniel Carlsson Solander or Daniel Charles Solander was a Swedish naturalist and an apostle of Carl Linnaeus. Solander was the first university educated scientist to set foot on Australian soil.-Biography:...

, naturalists on the Endeavour during Lieutenant (later Captain) James Cook
James Cook
Captain James Cook, FRS, RN was a British explorer, navigator and cartographer who ultimately rose to the rank of captain in the Royal Navy...

's 1770 voyage to the Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

. A number of arrangements were published in the 1800s, culminating in George Bentham's 1870 arrangement
Bentham's taxonomic arrangement of Banksia
George Bentham's taxonomic arrangement of Banksia was published in 1870, in Volume 5 of Bentham's Flora Australiensis. A substantial improvement on the previous arrangement, it would stand for over a century. It was eventually replaced by Alex George's 1981 arrangement, published in his classic...

. This arrangement would stand for over a hundred years.

1981 arrangement

Georges arrangement of Banksia was first published in his 1981 monograph The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. is a 1981 monograph by Alex George on the taxonomy of the plant genus Banksia. Published by the Western Australian Herbarium as Nuytsia 3, it presented George's taxonomic arrangement of Banksia, the first major taxonomic revision of the genus since George Bentham published...

. The arrangement was as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

B. sect. Banksia
Banksia sect. Banksia
Banksia sect. Banksia is one of four sections of Banksia subgenus Banksia. It contains those species of subgenus Banksia with straight or sometimes curved but not hooked styles. These species all have cylindrical inflorescences and usually exhibit a bottom-up sequence of flower anthesis...

B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

B. dentata
Banksia dentata
The Tropical Banksia is a species of small tree in the plant genus Banksia.-Distribution and habitat:It occurs throughout northern Australia, and also extends into New Guinea and the Aru Islands...

B. integrifolia
Banksia integrifolia
Banksia integrifolia, commonly known as Coast Banksia, is a species of tree that grows along the east coast of Australia. One of the most widely distributed Banksia species, it occurs between Victoria and Central Queensland in a broad range of habitats, from coastal dunes to mountains...

B. integrifolia var. integrifolia (now B. integrifolia subsp. integrifolia
Banksia integrifolia subsp. integrifolia
Banksia integrifolia subsp. integrifolia is a subspecies of Banksia integrifolia.-Description:B. integrifolia subsp. integrifolia can be distinguished from the other subspecies of B. integrifolia by its slightly smaller, broader leaves, which are also somewhat duller than those of...

)
B. integrifolia var. compar (now B. integrifolia subsp. compar
Banksia integrifolia subsp. compar
Banksia integrifolia subsp. compar is a subspecies of Banksia integrifolia. It has larger, glossier leaves than other subspecies, and occurs much further north.-Description:...

)
B. integrifolia var. aquilonia (now Banksia aquilonia
Banksia aquilonia
Banksia aquilonia is a species of tall shrub or tree in the plant genus Banksia.-Description:B. aquilonia grows as a tall shrub or small tree up to 8 metres high. It has hard, fissured, grey bark, and elliptic leaves without serrated margins...

)
B. conferta
Banksia conferta
The Glasshouse Banksia is a species of montane shrub or small tree in the plant genus Banksia. It occurs as distinct subspecies in two widely separated locations: B. conferta subsp. conferta in southern Queensland on the Lamington Plateau and further north in the Glass House Mountains; and...

B. conferta var. conferta (now B. conferta subsp. conferta
Banksia conferta subsp. conferta
Banksia conferta subsp. conferta is a subspecies of Banksia conferta. As an autonym, it is defined as encompassing the type material of the species. It grows only in Queensland between the Lamington Plateau and the Glass House Mountains....

)
B. conferta var. penicillata (now B. conferta subsp. penicillata
Banksia conferta subsp. penicillata
The Newnes Plateau Banksia is a plant only described in 1981 though collected in the 1970s...

)
B. marginata
Banksia marginata
Banksia marginata, commonly known as the Silver Banksia, is a species of tree or woody shrub in the plant genus Banksia found throughout much of southeastern Australia. It ranges from the Eyre Peninsula in South Australia, to north of Armidale, New South Wales, and across Tasmania and the islands...

B. canei
Banksia canei
The mountain banksia is a species of shrub in the plant genus Banksia. It occurs in subalpine areas of the Great Dividing Range between Melbourne and Canberra in southeastern Australia...

B. saxicola
Banksia saxicola
Banksia saxicola, commonly known as the Grampians Banksia is a species of tree or shrub in the plant genus Banksia. It occurs in Victoria in two distinct populations, one in The Grampians and the other on Wilsons Promontory....

B. oblongifolia
Banksia oblongifolia
The Fern-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs along the eastern coast of Australia from Wollongong, New South Wales in the south to Rockhampton, Queensland in the north...

B. robur
Banksia robur
Banksia robur, commonly known as Swamp Banksia or, less commonly, Broad-leaved Banksia grows in sand or peaty sand in coastal areas from Cooktown in north Queensland to the Illawarra region on the New South Wales south coast...

B. paludosa
Banksia paludosa
Banksia paludosa, commonly known as the marsh or swamp banksia, is a species of shrub in the plant genus Banksia. It is endemic to New South Wales, where it is found between Sydney and Batemans Bay, with an isolate population further south around Eden...

B. ser. Grandes
Banksia ser. Grandes
Banksia ser. Grandes is a taxonomic series in the genus Banksia. It consists of two closely related species in section Banksia, both endemic to Western Australia. These are B. grandis and B. solandri ....

B. grandis
Banksia grandis
Banksia grandis, commonly known as Bull Banksia, Giant Banksia or Mangite, is a common and distinctive tree in South West Western Australia....

B. solandri
Banksia solandri
Banksia solandri, commonly known as Stirling Range Banksia, is a species of large shrub in the plant genus Banksia. It occurs only within the Stirling Range in southwest Western Australia.-Description:...

B. ser. Quercinae
Banksia ser. Quercinae
Banksia ser. Quercinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

B. quercifolia
Banksia quercifolia
The Oak-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Manypeaks in the east.-Ecology:...

B. oreophila
Banksia oreophila
The Western Mountain Banksia or Mountain Banksia is a species of shrub in the plant genus Banksia. It occurs on the slopes and hilltops of the Stirling and Barren Ranges in southwest Western Australia.-Taxonomy:...

B. baueri
Banksia baueri
The Woolly Banksia is a species of shrub in the plant genus Banksia. It occurs in southwest Western Australia north and east of Albany. It has a distinctively large and hairy looking inflorescence which can be 300 mm or more long and up to 200 mm in diameter.It is placed alone in series...

B. ser. Orthostylis (now B. ser. Banksia
Banksia ser. Banksia
Banksia ser. Banksia is avalid botanic name for a series of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-According to Bentham:Banksia ser. Banksia originated in 1870 as...

)
B. serrata
Banksia serrata
Banksia serrata, commonly known as Old Man Banksia, Saw Banksia, Saw-tooth Banksia and Red Honeysuckle, is a species of woody shrub or tree of the genus Banksia in the Proteaceae family. Native the east coast of Australia, it is found from Queensland through to Victoria with outlying populations on...

B. aemula
Banksia aemula
Banksia aemula, commonly known as the wallum banksia, is a lignotuberous shrub of the Proteaceae family. Found from Bundaberg south to Sydney on the Australian east coast, it is encountered as a shrub or a taller tree to 8 m in coastal heath on deep sandy soil, known as Wallum...

B. ornata
Banksia ornata
The Desert Banksia is a species of shrub in the plant genus Banksia which grows up to 3 m tall. It occurs in western Victoria, and in South Australia, where it is common on the Eyre Peninsula, Kangaroo Island and east of Adelaide, but is apparently absent from Yorke Peninsula.It tends to grow...

B. menziesii
Banksia menziesii
Banksia menziesii, commonly known as firewood banksia, is a species of flowering plant in the genus Banksia. It is a gnarled tree up to 10 m tall, or a lower spreading 1–3 m shrub in the more northern parts of its range. The serrated leaves are dull green with new growth a paler grey...

B. speciosa
Banksia speciosa
The Showy Banksia is a species of large shrub or small tree in the plant genus Banksia. It reaches up to 8 m in height...

B. baxteri
Banksia baxteri
The Baxter's Banksia , also known as Bird's Nest Banksia, is a species of shrub in the plant genus Banksia. It occurs along the south coast of Western Australia between Albany and Esperance....

B. candolleana
Banksia candolleana
The Propeller Banksia is a species of shrub in the plant genus Banksia. It occurs in sandplain country north of Gingin, Western Australia.-Description:...

B. sceptrum
Banksia sceptrum
Banksia sceptrum, the Sceptre Banksia, occurs in Western Australia near the central west coast from Geraldton north through Kalbarri to Hamelin Pool. It extends inland almost to Mullewa....

B. ser. Crocinae
Banksia ser. Crocinae
Banksia ser. Crocinae is a taxonomic series in the genus Banksia. It consists of four closely related species, all of which are endemic to Western Australia; namely B. prionotes , B. burdettii , B. hookeriana and B. victoriae...

B. prionotes
Banksia prionotes
Banksia prionotes, commonly known as Acorn Banksia or Orange Banksia, is a species of shrub or tree of the genus Banksia in the family Proteaceae. It is native to the southwest of Western Australia and can reach up to 10 m in height. It can be much smaller in more exposed areas or in the...

B. victoriae
Banksia victoriae
Banksia victoriae, commonly known as Woolly Orange Banksia, is a species of large shrub or small tree in the plant genus Banksia...

B. hookerana (now spelled B. hookeriana
Banksia hookeriana
The Hooker's Banksia is a species of shrub in the plant genus Banksia. It occurs on sandplain shrubland between Arrowsmith and Eneabba in Western Australia. Seeds do not require any treatment, and take 23 to 49 days to germinate....

)
B. burdettii
Banksia burdettii
Burdett's Banksia is a species of large shrub or small tree in the plant genus Banksia. It occurs on sandplain country north of Gingin, Western Australia....

B. ser. Cyrtostylis
Banksia ser. Cyrtostylis
Banksia ser. Cyrtostylis is a valid botanic name for a taxonomic series within the plant genus Banksia. First published at sectional rank by George Bentham in 1870, it was demoted to a series by Alex George in 1981. The name has had three circumscriptions.-According to Bentham:Bentham published B....

B. media
Banksia media
The Southern Plains Banksia , also known as Golden Stalk Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia between Albany and Israelite Bay, where it is a common plant....

B. praemorsa
Banksia praemorsa
The Cut-leaf Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in a few isolated populations on the south coast of Western Australia between Albany and Cape Riche. Seeds do not require any treatment, and take 30 to 49 days to germinate.-References:...

B. pilostylis
Banksia pilostylis
The Marsh Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia around Esperance. Seeds do not require any treatment, and take 18 to 49 days to germinate.-External links:...

B. attenuata
Banksia attenuata
Banksia attenuata, commonly known as the candlestick banksia or slender banksia, is a species of plant in the proteaceae family. Commonly a tree, it reaches 10 m high, but is often a shrub in dryer areas 0.4 to 2 m high...

B. lindleyana
Banksia lindleyana
Banksia lindleyana, commonly known as the Porcupine Banksia, is a species of woody shrub in the genus Banksia of the family Proteaceae. It generally grows as a small shrub to 1 m high with long narrow serrated leaves, and bright yellow oval or round inflorescences...

B. ashbyi
Banksia ashbyi
The Ashby's Banksia is a species of shrub in the plant genus Banksia. It occurs in heath and spinifex country along the coast of Western Australia between Geraldton and Exmouth.-Description:...

B. benthamiana
Banksia benthamiana
Bentham's Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in scattered populations in the Wheatbelt region of Western Australia, one near Mullewa and the other near Dalwallinu.-Description:...

B. audax
Banksia audax
Banksia audax is a species of shrub in the plant genus Banksia. It occurs over a large area in the central south of Western Australia.-Description:...

B. laevigata
Banksia laevigata
Banksia laevigata is a species of shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland from Southern Cross south to the Fitzgerald River National Park. It is composed of two closely related subspecies, B. laevigata subsp. laevigata and B. laevigata...

B. laevigata subsp. laevigata
Banksia laevigata subsp. laevigata
The Tennis Ball Banksia is a subspecies of small woody shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland. It and the closely related B. laevigata subsp...

B. laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea is a subspecies of Banksia laevigata. It is native to the Southwest Botanical Province of Western Australia....

B. lullfitzii
Banksia lullfitzii
Banksia lullfitzii is a species of shrub in the plant genus Banksia. A many-branched, spreading bush with golden-orange flowers, it occurs in scattered populations over a large area of the eastern goldfields of Western Australia...

B. elderana (now spelled B. elderiana
Banksia elderiana
The Swordfish Banksia , commonly known as the swordfish banksia or palm banksia, is a species of shrub in the plant genus Banksia. It occurs in two disjunct areas in the Goldfields-Esperance region of Western Australia...

)
B. elegans
Banksia elegans
The Elegant Banksia is a species of shrub or small tree in the plant genus Banksia. It occurs only over a 65 square kilometre area north and west of Eneabba, Western Australia.-Ecology:...

B. ser. Prostratae
Banksia ser. Prostratae
Banksia ser. Prostratae is a taxonomic series in the genus Banksia. It consists of six closely related species in section Banksia, all endemic to Western Australia, with a prostrate habit.Banksia ser...

B. goodii
Banksia goodii
Banksia goodii, commonly known as Good's Banksia, is an endangered shrub of Southwest Western Australia.Good's Banksia grows as a low shrub, either prostrate or with stems up to twenty centimetres high. The leaves are dark green with a prominent yellow midrib, and are held erect. They may be up...

B. gardneri
Banksia gardneri
The Prostrate Banksia is a species of prostrate shrub in the plant genus Banksia. It occurs along the south coast of Western Australia.-Scientific name:...

B. gardneri var. gardneri
Banksia gardneri var. gardneri
Banksia gardneri var. gardneri is a variety of Banksia gardneri. As an autonym, it is defined as encompassing the type material of the species. It is native to the Southwest Botanical Province of Western Australia....

B. gardneri var. brevidentata
Banksia gardneri var. brevidentata
Banksia gardneri var. brevidentata is a variety of Banksia gardneri. It is native to the Southwest Botanical Province of Western Australia....

B. gardneri var. hiemalis
Banksia gardneri var. hiemalis
Banksia gardneri var. hiemalis is a variety of Banksia gardneri. It is native to the Southwest Botanical Province of Western Australia. Seeds do not require any treatment, and take around 19 days to germinate....

B. chamaephyton
Banksia chamaephyton
The Fishbone Banksia is a species of prostrate shrub in the plant genus Banksia. It occurs in sandplain country amongst low heath north of Perth, Western Australia.....

B. repens
Banksia repens
Banksia repens, the Creeping Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Ragged in the east....

B. blechnifolia
Banksia blechnifolia
Banksia blechnifolia is a species of prostrate shrub in the plant genus Banksia. Found in sandy soils in the south coastal region of Western Australia in the vicinity of Lake King, it is non-lignotuberous, regenerating by seed after bushfire...

B. petiolaris
Banksia petiolaris
Banksia petiolaris is a species of shrub of the genus Banksia in the Proteaceae family. It is a prostrate banksia, a group of several closely related species all with horizontal stems and thick, leathery upright leaves. No varieties have been subsequently described. It bears yellow inflorescences...

B. ser. Tetragonae
Banksia ser. Tetragonae
Banksia ser. Tetragonae is a taxonomic series in the genus Banksia. It consists of three closely related species of erect shrub with pendulous inflorescences in section Banksia. These are B. lemanniana , B. caleyi and B. aculeata ....

B. lemanniana
Banksia lemanniana
Banksia lemanniana, commonly known as the Yellow lantern Banksia or Lemann's Banksia, is a species of woody shrub in the genus Banksia of the family Proteaceae. It generally grows as an open shrub or small tree to 5 m high with stiff serrated leaves, and unusually hanging inflorescences...

B. caleyi
Banksia caleyi
Banksia caleyi, commonly known as the red lantern banksia or Cayley's banksia, is a species of shrub in the plant genus Banksia. It occurs in southern Western Australia south and east of the Stirling Ranges through to around Jerramungup...

B. aculeata
Banksia aculeata
Banksia aculeata, commonly known as Prickly Banksia, is a species of shrub native to the Stirling Range in southwest Western Australia. One of three closely related species all of which have distinctive upside down lantern-like inflorescences, it bears pinkish blooms in early summer. The leaves...

B. ser. Coccineae
B. coccinea
Banksia coccinea
Banksia coccinea, commonly known as the Scarlet Banksia, Waratah Banksia or Albany Banksia, is an erect shrub or small tree in the plant genus Banksia...

B. sect. Oncostylis
Banksia sect. Oncostylis
Banksia sect. Oncostylis is one of four sections of subgenus Banksia subg. Banksia. It contains those Banksia species with hooked pistils. All of the species in Oncostylis also exhibit a top-down sequence of flower anthesis, except for Banksia nutans which is bottom-up.Banksia sect...

B. ser. Spicigerae
Banksia ser. Spicigerae
Banksia ser. Spicigerae is a taxonomic series in the genus Banksia. It consists of the seven species in section Oncostylis that have cylindrical inflorescences. These range in form from small shrubs to tall trees. The leaves grow in either an alternate or whorled pattern, with various shape forms...

B. spinulosa
Banksia spinulosa
The Hairpin Banksia is a species of woody shrub, of the genus Banksia in the Proteaceae family, native to eastern Australia. Widely distributed, it is found as an understorey plant in open dry forest or heathland from Victoria to northern Queensland, generally on sandstone though sometimes also...

B. spinulosa var. spinulosa
Banksia spinulosa var. spinulosa
Banksia spinulosa var. spinulosa is a shrub that grows along the east coast of Australia, in Queensland and New South Wales.-Description:...

B. spinulosa var. collina
Banksia spinulosa var. collina
Banksia spinulosa var. collina is a shrub that grows along the east coast of Australia, in Queensland and New South Wales. Commonly known as Hill Banksia or Golden Candlesticks, it is a taxonomic variety of B. spinulosa...

B. spinulosa var. cunninghamii
Banksia spinulosa var. cunninghamii
Banksia spinulosa var. cunninghamii, sometimes given species rank as Banksia cunninghamii, is a shrub that grows along the east coast of Australia, in Victoria and New South Wales...

B. ericifolia
Banksia ericifolia
Banksia ericifolia, the Heath-leaved Banksia , is a species of woody shrub of the Proteaceae family native to Australia. It grows in two separate regions of Central and Northern New South Wales east of the Great Dividing Range...

B. ericifolia subsp. ericifolia
Banksia ericifolia subsp. ericifolia
Banksia ericifolia subsp. ericifolia is a subspecies of Banksia ericifolia. As an autonym, it is defined as encompassing the type material of the species. It is native to New South Wales, but has naturalised in small numbers in Victoria....

B. ericifolia subsp. macrantha
Banksia ericifolia subsp. macrantha
Banksia ericifolia subsp. macrantha is a subspecies of Banksia ericifolia. It is native to New South Wales....

B. brownii
Banksia brownii
Banksia brownii, commonly known as Feather-leaved Banksia or Brown's Banksia, is a species of shrub that occurs in southwest Western Australia. An attractive plant with fine feathery leaves and large red-brown flower spikes, it usually grows as an upright bush around two metres high, but can also...

B. occidentalis
Banksia occidentalis
The Red Swamp Banksia or Waterbush is a species of shrub or small tree in the plant genus Banksia. It occurs on the south coast of Western Australia in three disjunct populations: at Augusta, around Albany and in the Esperance area.A 1980 field study at Cheyne beach showed it to be pollinated by...

B. littoralis
Banksia littoralis
Banksia littoralis, commonly known as the Swamp Banksia, Swamp Oak, Pungura and the Western Swamp Banksia, is a tree in the plant genus Banksia. It is found in south west Western Australia from the south eastern metropolitan area of Perth to the Stirling Range and Albany...

B. littoralis var. littoralis
B. littoralis var. seminuda (now B. seminuda
Banksia seminuda
Banksia seminuda, commonly known as the River Banksia, is a tree in the plant genus Banksia. It is found in south west Western Australia from Dwellingup to the Broke Inlet east of Denmark . It is often mistaken for and was originally considered a subspecies of the Banksia littoralis...

)
B. verticillata
Banksia verticillata
Banksia verticillata, commonly known as Granite Banksia or Albany Banksia, is a species of shrub or tree of the genus Banksia in the Proteaceae family. It is native to the southwest of Western Australia and can reach up to 3 m in height. It can grow taller to 5 m in sheltered areas,...

B. tricuspis
Banksia tricuspis
The Lesueur Banksia or Pine Banksia is a species of shrub or tree in the plant genus Banksia. It occurs within a geographic range of just 15 square kilometres near Jurien, Western Australia.-External links:...

B. ser. Dryandroideae
Banksia ser. Dryandroideae
Banksia ser. Dryandroideae is a valid botanic name for a taxonomic series in the plant genus Banksia. First published by Carl Meissner in 1856, the name has had two circumscriptions. As presently circumscribed it is monotypic, containing only B. dryandroides.-According to Meissner:B. ser...

B. dryandroides
Banksia dryandroides
Banksia dryandroides, the Dryandra-leaved Banksia, is a species of small shrub in the plant genus Banksia. It occurs in shrubland, coastal heath and woodland on the south coast of Western Australia between Two Peoples Bay and Cheyne Bay. The species is placed alone in series B. ser...

B. ser. Abietinae
Banksia ser. Abietinae
Banksia ser. Abietinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

B. sphaerocarpa
Banksia sphaerocarpa
Banksia sphaerocarpa, commonly known as the Fox Banksia or Round-fruit Banksia, is a species of shrub or tree in the plant genus Banksia . It is generally encountered as a 1–2 m high shrub, and is usually smaller in the north of its range...

B. sphaerocarpa  var. sphaerocarpa
Banksia sphaerocarpa var. sphaerocarpa
Banksia sphaerocarpa var. sphaerocarpa is a variety of Banksia sphaerocarpa. It is native to the Southwest Botanical Province of Western Australia. As an autonym, it is defined as containing the type specimen of the species....

B. sphaerocarpa  var. caesia
Banksia sphaerocarpa var. caesia
Banksia sphaerocarpa var. caesia is a variety of Banksia sphaerocarpa. It is native to the Southwest Botanical Province of Western Australia....

B. sphaerocarpa  var. dolichostyla
Banksia sphaerocarpa var. dolichostyla
Banksia sphaerocarpa var. dolichostyla, commonly known as Ironcap Banksia, is a plant which is either considered a variety of Banksia sphaerocarpa, or as a species in its own right. It is native to the Southwest Botanical Province of Western Australia...

B. micrantha
Banksia micrantha
Banksia micrantha is a species of shrub in the plant genus Banksia. A small spreading bush with pale yellow flower spikes, it occurs between Eneabba and Cervantes in South west Western Australia...

B. grossa
Banksia grossa
Banksia grossa, commonly known as Coarse Banksia is a species of shrub in the plant family Proteaceae endemic to south west Western Australia. One of fourteen species of banksia with predominantly round or oval inflorescences of the series Abietinae, it was described in 1981 as a distinct species...

B. leptophylla
Banksia leptophylla
The Slender-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs along the west coast of Western Australia from Gingin to Kalbarri. Before Alex George's revision of 1981, it was labelled informally as B. sphaerocarpa var. pinifolia or var...

B. lanata
Banksia lanata
The Coomallo Banksia is a species of shrub in the plant genus Banksia. It occurs within a range of less than 100 square kilometres between Eneabba and Mount Lesueur, Western Australia. It has roughly spherical inflorescences with flowers of cream to orange-brown colour. The leaves are linear and...

B. scabrella
Banksia scabrella
Banksia scabrella, commonly known as the Burma Road Banksia, is a species of woody shrub in the genus Banksia. It is classified in the series Abietinae, a group of several species of shrubs with small round or oval inflorescences...

B. telmatiaea
Banksia telmatiaea
Banksia telmatiaea, commonly known as Swamp Fox Banksia or rarely Marsh Banksia, is a shrub that grows in marshes and swamps along the lower west coast of Australia. It grows as an upright bush up to 2 m tall, with narrow leaves and a pale brown flower spike, which can produce profuse...

B. laricina
Banksia laricina
The Rose-Fruited Banksia is a species of shrub in the plant genus Banksia endemic to southwestern Western Australia. It derives its specific Latin name from larix or larch, which its foliage is said to resemble. The common name comes from the striking fruits which resemble wooden roses...

B. incana
Banksia incana
The Hoary Banksia is a species of small shrub in the plant genus Banksia. It occurs on sandplain heathland between Badgingarra and Eneabba in Western Australia, with outlying populations as far south as Perth. Seeds do not require any treatment, and take around 14 days to germinate....

B. violacea
Banksia violacea
Banksia violacea, commonly known as Violet Banksia, is a species of shrub or tree in the plant genus Banksia . It generally grows as a small shrub to 1.5 m high with fine narrow leaves, and is best known for its unusually coloured dark purple-violet inflorescences...

B. meisneri
Banksia meisneri
The Meisner's Banksia is a species of small shrub in the plant genus Banksia. It occurs in a number of isolated populations throughout southwest Western Australia. Seeds do not require any treatment, and take 28 to 39 days to germinate.-External links:...

B. meisneri subsp. meisneri
Banksia meisneri subsp. meisneri
Banksia meisneri subsp. meisneri is a subspecies of Banksia meisneri. It is native to the Southwest Botanical Province of Western Australia. As an autonym, it is defined as containing the type specimen of the species....

B. meisneri subsp. ascendens
Banksia meisneri subsp. ascendens
Banksia meisneri subsp. ascendens, commonly known as Scott River Banksia, is a subspecies of Banksia meisneri. It is native to the Southwest Botanical Province of Western Australia....

B. pulchella
Banksia pulchella
The Teasel Banksia is a species of small shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from Fitzgerald River National Park east to Israelite Bay....

B. nutans
Banksia nutans
Banksia nutans, commonly known as Nodding Banksia, is a species of shrub native to the south coast of Western Australia in the genus Banksia...

B. nutans var. nutans
Banksia nutans var. nutans
Banksia nutans var. nutans is a variety of Banksia nutans. It is native to the Southwest Botanical Province of Western Australia. As an autonym, it is defined as containing the type specimen of the species....

B. nutans var. cernuella
Banksia nutans var. cernuella
Banksia nutans var. cernuella is a variety of Banksia nutans. It is native to the Southwest Botanical Province of Western Australia. Seeds do not require any treatment, and take around 17 days to germinate....

B. subg. Isostylis
Banksia subg. Isostylis
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser...

B. ilicifolia
Banksia ilicifolia
Banksia ilicifolia, commonly known as Holly-leaved Banksia, is a tree in the plant Proteaceae family. Endemic to southwest Western Australia, it belongs to Banksia subg. Isostylis, a subgenus of three closely related Banksia species with inflorescences that are dome-shaped heads rather than...

B. cuneata
Banksia cuneata
Banksia cuneata, commonly known as Matchstick Banksia or Quairading Banksia, is an endangered species of flowering plants in the Proteaceae family. Endemic to southwest Western Australia, it belongs to Banksia subg...


1999 arrangement

George's arrangement was superseded by Thiele and Ladiges' arrangement
Thiele and Ladiges' taxonomic arrangement of Banksia
Kevin Thiele and Pauline Ladiges' taxonomic arrangement of Banksia, published in 1996, was a novel taxonomic arrangement that was intended to align the taxonomy of Banksia more closely with the phylogeny that they had inferred from their cladistic analysis of the genus...

 in 1996, but in 1999 George largely restored it in his 1999 treatment of the Flora of Australia
Flora of Australia (series)
The Flora of Australia is a 59 volume series describing the vascular plants, bryophytes and lichens present in Australia and its external territories...

 series. George's 1999 treatment included a number of taxa described after 1981, followed Thiele and Ladiges in a few minor rankings, and altered the phyletic order of taxa in places, but it was otherwise identical to his 1981 arrangement. None of Thiele and Ladiges' four promotions to species rank was accepted, and none of the thirteen infrageneric taxa introduced by Thiele and Ladiges was retained.

George's 1999 taxonomic arrangement of Banksia was as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

B. sect. Banksia
Banksia sect. Banksia
Banksia sect. Banksia is one of four sections of Banksia subgenus Banksia. It contains those species of subgenus Banksia with straight or sometimes curved but not hooked styles. These species all have cylindrical inflorescences and usually exhibit a bottom-up sequence of flower anthesis...

B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

B. dentata
Banksia dentata
The Tropical Banksia is a species of small tree in the plant genus Banksia.-Distribution and habitat:It occurs throughout northern Australia, and also extends into New Guinea and the Aru Islands...

B. aquilonia
Banksia aquilonia
Banksia aquilonia is a species of tall shrub or tree in the plant genus Banksia.-Description:B. aquilonia grows as a tall shrub or small tree up to 8 metres high. It has hard, fissured, grey bark, and elliptic leaves without serrated margins...

B. integrifolia
Banksia integrifolia
Banksia integrifolia, commonly known as Coast Banksia, is a species of tree that grows along the east coast of Australia. One of the most widely distributed Banksia species, it occurs between Victoria and Central Queensland in a broad range of habitats, from coastal dunes to mountains...

B. integrifolia subsp. integrifolia
Banksia integrifolia subsp. integrifolia
Banksia integrifolia subsp. integrifolia is a subspecies of Banksia integrifolia.-Description:B. integrifolia subsp. integrifolia can be distinguished from the other subspecies of B. integrifolia by its slightly smaller, broader leaves, which are also somewhat duller than those of...

B. integrifolia subsp. compar
Banksia integrifolia subsp. compar
Banksia integrifolia subsp. compar is a subspecies of Banksia integrifolia. It has larger, glossier leaves than other subspecies, and occurs much further north.-Description:...

B. integrifolia subsp. monticola
Banksia integrifolia subsp. monticola
Banksia integrifolia subsp. monticola, commonly known as White Mountain Banksia, is a subspecies of Banksia integrifolia. Described in 1994, it occurs in the Blue Mountains in northern New South Wales. It contains the largest recorded Banksia trees.-Description:B. integrifolia subsp. monticola...

B. plagiocarpa
Banksia plagiocarpa
Banksia plagiocarpa, commonly known as the Dallachy's banksia or blue banksia, is a species of shrub or tree in the plant genus Banksia. It occurs only on Hinchinbrook Island, Queensland and the immediately adjacent mainland....

B. oblongifolia
Banksia oblongifolia
The Fern-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs along the eastern coast of Australia from Wollongong, New South Wales in the south to Rockhampton, Queensland in the north...

B. robur
Banksia robur
Banksia robur, commonly known as Swamp Banksia or, less commonly, Broad-leaved Banksia grows in sand or peaty sand in coastal areas from Cooktown in north Queensland to the Illawarra region on the New South Wales south coast...

B. conferta
Banksia conferta
The Glasshouse Banksia is a species of montane shrub or small tree in the plant genus Banksia. It occurs as distinct subspecies in two widely separated locations: B. conferta subsp. conferta in southern Queensland on the Lamington Plateau and further north in the Glass House Mountains; and...

B. conferta subsp. conferta
Banksia conferta subsp. conferta
Banksia conferta subsp. conferta is a subspecies of Banksia conferta. As an autonym, it is defined as encompassing the type material of the species. It grows only in Queensland between the Lamington Plateau and the Glass House Mountains....

B. conferta subsp. penicillata
Banksia conferta subsp. penicillata
The Newnes Plateau Banksia is a plant only described in 1981 though collected in the 1970s...

B. paludosa
Banksia paludosa
Banksia paludosa, commonly known as the marsh or swamp banksia, is a species of shrub in the plant genus Banksia. It is endemic to New South Wales, where it is found between Sydney and Batemans Bay, with an isolate population further south around Eden...

B. paludosa subsp. astrolux
B. paludosa subsp. paludosa
B. marginata
Banksia marginata
Banksia marginata, commonly known as the Silver Banksia, is a species of tree or woody shrub in the plant genus Banksia found throughout much of southeastern Australia. It ranges from the Eyre Peninsula in South Australia, to north of Armidale, New South Wales, and across Tasmania and the islands...

B. canei
Banksia canei
The mountain banksia is a species of shrub in the plant genus Banksia. It occurs in subalpine areas of the Great Dividing Range between Melbourne and Canberra in southeastern Australia...

B. saxicola
Banksia saxicola
Banksia saxicola, commonly known as the Grampians Banksia is a species of tree or shrub in the plant genus Banksia. It occurs in Victoria in two distinct populations, one in The Grampians and the other on Wilsons Promontory....

B. ser. Grandes
Banksia ser. Grandes
Banksia ser. Grandes is a taxonomic series in the genus Banksia. It consists of two closely related species in section Banksia, both endemic to Western Australia. These are B. grandis and B. solandri ....

B. grandis
Banksia grandis
Banksia grandis, commonly known as Bull Banksia, Giant Banksia or Mangite, is a common and distinctive tree in South West Western Australia....

B. solandri
Banksia solandri
Banksia solandri, commonly known as Stirling Range Banksia, is a species of large shrub in the plant genus Banksia. It occurs only within the Stirling Range in southwest Western Australia.-Description:...

B. ser. Banksia
Banksia ser. Banksia
Banksia ser. Banksia is avalid botanic name for a series of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-According to Bentham:Banksia ser. Banksia originated in 1870 as...

B. serrata
Banksia serrata
Banksia serrata, commonly known as Old Man Banksia, Saw Banksia, Saw-tooth Banksia and Red Honeysuckle, is a species of woody shrub or tree of the genus Banksia in the Proteaceae family. Native the east coast of Australia, it is found from Queensland through to Victoria with outlying populations on...

B. aemula
Banksia aemula
Banksia aemula, commonly known as the wallum banksia, is a lignotuberous shrub of the Proteaceae family. Found from Bundaberg south to Sydney on the Australian east coast, it is encountered as a shrub or a taller tree to 8 m in coastal heath on deep sandy soil, known as Wallum...

B. ornata
Banksia ornata
The Desert Banksia is a species of shrub in the plant genus Banksia which grows up to 3 m tall. It occurs in western Victoria, and in South Australia, where it is common on the Eyre Peninsula, Kangaroo Island and east of Adelaide, but is apparently absent from Yorke Peninsula.It tends to grow...

B. baxteri
Banksia baxteri
The Baxter's Banksia , also known as Bird's Nest Banksia, is a species of shrub in the plant genus Banksia. It occurs along the south coast of Western Australia between Albany and Esperance....

B. speciosa
Banksia speciosa
The Showy Banksia is a species of large shrub or small tree in the plant genus Banksia. It reaches up to 8 m in height...

B. menziesii
Banksia menziesii
Banksia menziesii, commonly known as firewood banksia, is a species of flowering plant in the genus Banksia. It is a gnarled tree up to 10 m tall, or a lower spreading 1–3 m shrub in the more northern parts of its range. The serrated leaves are dull green with new growth a paler grey...

B. candolleana
Banksia candolleana
The Propeller Banksia is a species of shrub in the plant genus Banksia. It occurs in sandplain country north of Gingin, Western Australia.-Description:...

B. sceptrum
Banksia sceptrum
Banksia sceptrum, the Sceptre Banksia, occurs in Western Australia near the central west coast from Geraldton north through Kalbarri to Hamelin Pool. It extends inland almost to Mullewa....

B. ser. Crocinae
Banksia ser. Crocinae
Banksia ser. Crocinae is a taxonomic series in the genus Banksia. It consists of four closely related species, all of which are endemic to Western Australia; namely B. prionotes , B. burdettii , B. hookeriana and B. victoriae...

B. prionotes
Banksia prionotes
Banksia prionotes, commonly known as Acorn Banksia or Orange Banksia, is a species of shrub or tree of the genus Banksia in the family Proteaceae. It is native to the southwest of Western Australia and can reach up to 10 m in height. It can be much smaller in more exposed areas or in the...

B. burdettii
Banksia burdettii
Burdett's Banksia is a species of large shrub or small tree in the plant genus Banksia. It occurs on sandplain country north of Gingin, Western Australia....

B. hookeriana
Banksia hookeriana
The Hooker's Banksia is a species of shrub in the plant genus Banksia. It occurs on sandplain shrubland between Arrowsmith and Eneabba in Western Australia. Seeds do not require any treatment, and take 23 to 49 days to germinate....

B. victoriae
Banksia victoriae
Banksia victoriae, commonly known as Woolly Orange Banksia, is a species of large shrub or small tree in the plant genus Banksia...

B. ser. Prostratae
Banksia ser. Prostratae
Banksia ser. Prostratae is a taxonomic series in the genus Banksia. It consists of six closely related species in section Banksia, all endemic to Western Australia, with a prostrate habit.Banksia ser...

B. goodii
Banksia goodii
Banksia goodii, commonly known as Good's Banksia, is an endangered shrub of Southwest Western Australia.Good's Banksia grows as a low shrub, either prostrate or with stems up to twenty centimetres high. The leaves are dark green with a prominent yellow midrib, and are held erect. They may be up...

B. gardneri
Banksia gardneri
The Prostrate Banksia is a species of prostrate shrub in the plant genus Banksia. It occurs along the south coast of Western Australia.-Scientific name:...

B. gardneri var. gardneri
Banksia gardneri var. gardneri
Banksia gardneri var. gardneri is a variety of Banksia gardneri. As an autonym, it is defined as encompassing the type material of the species. It is native to the Southwest Botanical Province of Western Australia....

B. gardneri var. brevidentata
Banksia gardneri var. brevidentata
Banksia gardneri var. brevidentata is a variety of Banksia gardneri. It is native to the Southwest Botanical Province of Western Australia....

B. gardneri var. hiemalis
Banksia gardneri var. hiemalis
Banksia gardneri var. hiemalis is a variety of Banksia gardneri. It is native to the Southwest Botanical Province of Western Australia. Seeds do not require any treatment, and take around 19 days to germinate....

B. chamaephyton
Banksia chamaephyton
The Fishbone Banksia is a species of prostrate shrub in the plant genus Banksia. It occurs in sandplain country amongst low heath north of Perth, Western Australia.....

B. blechnifolia
Banksia blechnifolia
Banksia blechnifolia is a species of prostrate shrub in the plant genus Banksia. Found in sandy soils in the south coastal region of Western Australia in the vicinity of Lake King, it is non-lignotuberous, regenerating by seed after bushfire...

B. repens
Banksia repens
Banksia repens, the Creeping Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Ragged in the east....

B. petiolaris
Banksia petiolaris
Banksia petiolaris is a species of shrub of the genus Banksia in the Proteaceae family. It is a prostrate banksia, a group of several closely related species all with horizontal stems and thick, leathery upright leaves. No varieties have been subsequently described. It bears yellow inflorescences...

B. ser. Cyrtostylis
Banksia ser. Cyrtostylis
Banksia ser. Cyrtostylis is a valid botanic name for a taxonomic series within the plant genus Banksia. First published at sectional rank by George Bentham in 1870, it was demoted to a series by Alex George in 1981. The name has had three circumscriptions.-According to Bentham:Bentham published B....

B. media
Banksia media
The Southern Plains Banksia , also known as Golden Stalk Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia between Albany and Israelite Bay, where it is a common plant....

B. praemorsa
Banksia praemorsa
The Cut-leaf Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in a few isolated populations on the south coast of Western Australia between Albany and Cape Riche. Seeds do not require any treatment, and take 30 to 49 days to germinate.-References:...

B. epica
Banksia epica
Banksia epica is a shrub that grows on the south coast of Western Australia. A spreading bush with wedge-shaped serrated leaves and large creamy-yellow flower spikes, it grows up to 3½ metres high. It is known only from two isolated populations in the remote south east of the state, near the...

B. pilostylis
Banksia pilostylis
The Marsh Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia around Esperance. Seeds do not require any treatment, and take 18 to 49 days to germinate.-External links:...

B. attenuata
Banksia attenuata
Banksia attenuata, commonly known as the candlestick banksia or slender banksia, is a species of plant in the proteaceae family. Commonly a tree, it reaches 10 m high, but is often a shrub in dryer areas 0.4 to 2 m high...

B. ashbyi
Banksia ashbyi
The Ashby's Banksia is a species of shrub in the plant genus Banksia. It occurs in heath and spinifex country along the coast of Western Australia between Geraldton and Exmouth.-Description:...

B. benthamiana
Banksia benthamiana
Bentham's Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in scattered populations in the Wheatbelt region of Western Australia, one near Mullewa and the other near Dalwallinu.-Description:...

B. audax
Banksia audax
Banksia audax is a species of shrub in the plant genus Banksia. It occurs over a large area in the central south of Western Australia.-Description:...

B. lullfitzii
Banksia lullfitzii
Banksia lullfitzii is a species of shrub in the plant genus Banksia. A many-branched, spreading bush with golden-orange flowers, it occurs in scattered populations over a large area of the eastern goldfields of Western Australia...

B. elderiana
Banksia elderiana
The Swordfish Banksia , commonly known as the swordfish banksia or palm banksia, is a species of shrub in the plant genus Banksia. It occurs in two disjunct areas in the Goldfields-Esperance region of Western Australia...

B. laevigata
Banksia laevigata
Banksia laevigata is a species of shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland from Southern Cross south to the Fitzgerald River National Park. It is composed of two closely related subspecies, B. laevigata subsp. laevigata and B. laevigata...

B. laevigata subsp. laevigata
Banksia laevigata subsp. laevigata
The Tennis Ball Banksia is a subspecies of small woody shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland. It and the closely related B. laevigata subsp...

B. laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea is a subspecies of Banksia laevigata. It is native to the Southwest Botanical Province of Western Australia....

B. elegans
Banksia elegans
The Elegant Banksia is a species of shrub or small tree in the plant genus Banksia. It occurs only over a 65 square kilometre area north and west of Eneabba, Western Australia.-Ecology:...

B. lindleyana
Banksia lindleyana
Banksia lindleyana, commonly known as the Porcupine Banksia, is a species of woody shrub in the genus Banksia of the family Proteaceae. It generally grows as a small shrub to 1 m high with long narrow serrated leaves, and bright yellow oval or round inflorescences...

B. ser. Tetragonae
Banksia ser. Tetragonae
Banksia ser. Tetragonae is a taxonomic series in the genus Banksia. It consists of three closely related species of erect shrub with pendulous inflorescences in section Banksia. These are B. lemanniana , B. caleyi and B. aculeata ....

B. lemanniana
Banksia lemanniana
Banksia lemanniana, commonly known as the Yellow lantern Banksia or Lemann's Banksia, is a species of woody shrub in the genus Banksia of the family Proteaceae. It generally grows as an open shrub or small tree to 5 m high with stiff serrated leaves, and unusually hanging inflorescences...

B. caleyi
Banksia caleyi
Banksia caleyi, commonly known as the red lantern banksia or Cayley's banksia, is a species of shrub in the plant genus Banksia. It occurs in southern Western Australia south and east of the Stirling Ranges through to around Jerramungup...

B. aculeata
Banksia aculeata
Banksia aculeata, commonly known as Prickly Banksia, is a species of shrub native to the Stirling Range in southwest Western Australia. One of three closely related species all of which have distinctive upside down lantern-like inflorescences, it bears pinkish blooms in early summer. The leaves...

B. ser. Bauerinae
B. baueri
Banksia baueri
The Woolly Banksia is a species of shrub in the plant genus Banksia. It occurs in southwest Western Australia north and east of Albany. It has a distinctively large and hairy looking inflorescence which can be 300 mm or more long and up to 200 mm in diameter.It is placed alone in series...

B. ser. Quercinae
Banksia ser. Quercinae
Banksia ser. Quercinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

B. quercifolia
Banksia quercifolia
The Oak-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Manypeaks in the east.-Ecology:...

B. oreophila
Banksia oreophila
The Western Mountain Banksia or Mountain Banksia is a species of shrub in the plant genus Banksia. It occurs on the slopes and hilltops of the Stirling and Barren Ranges in southwest Western Australia.-Taxonomy:...

B. sect. Coccinea
B. coccinea
Banksia coccinea
Banksia coccinea, commonly known as the Scarlet Banksia, Waratah Banksia or Albany Banksia, is an erect shrub or small tree in the plant genus Banksia...

B. sect. Oncostylis
Banksia sect. Oncostylis
Banksia sect. Oncostylis is one of four sections of subgenus Banksia subg. Banksia. It contains those Banksia species with hooked pistils. All of the species in Oncostylis also exhibit a top-down sequence of flower anthesis, except for Banksia nutans which is bottom-up.Banksia sect...

B. ser. Spicigerae
Banksia ser. Spicigerae
Banksia ser. Spicigerae is a taxonomic series in the genus Banksia. It consists of the seven species in section Oncostylis that have cylindrical inflorescences. These range in form from small shrubs to tall trees. The leaves grow in either an alternate or whorled pattern, with various shape forms...

B. spinulosa
Banksia spinulosa
The Hairpin Banksia is a species of woody shrub, of the genus Banksia in the Proteaceae family, native to eastern Australia. Widely distributed, it is found as an understorey plant in open dry forest or heathland from Victoria to northern Queensland, generally on sandstone though sometimes also...

B. spinulosa var. spinulosa
Banksia spinulosa var. spinulosa
Banksia spinulosa var. spinulosa is a shrub that grows along the east coast of Australia, in Queensland and New South Wales.-Description:...

B. spinulosa var. collina
Banksia spinulosa var. collina
Banksia spinulosa var. collina is a shrub that grows along the east coast of Australia, in Queensland and New South Wales. Commonly known as Hill Banksia or Golden Candlesticks, it is a taxonomic variety of B. spinulosa...

B. spinulosa var. neoanglica
Banksia spinulosa var. neoanglica
Banksia spinulosa var. neoanglica, commonly known as New England Banksia, is a shrub that grows along the east coast of Australia, in Queensland and New South Wales.-Description:...

B. spinulosa var. cunninghamii
Banksia spinulosa var. cunninghamii
Banksia spinulosa var. cunninghamii, sometimes given species rank as Banksia cunninghamii, is a shrub that grows along the east coast of Australia, in Victoria and New South Wales...

B. ericifolia
Banksia ericifolia
Banksia ericifolia, the Heath-leaved Banksia , is a species of woody shrub of the Proteaceae family native to Australia. It grows in two separate regions of Central and Northern New South Wales east of the Great Dividing Range...

B. ericifolia subsp. ericifolia
Banksia ericifolia subsp. ericifolia
Banksia ericifolia subsp. ericifolia is a subspecies of Banksia ericifolia. As an autonym, it is defined as encompassing the type material of the species. It is native to New South Wales, but has naturalised in small numbers in Victoria....

B. ericifolia subsp. macrantha
Banksia ericifolia subsp. macrantha
Banksia ericifolia subsp. macrantha is a subspecies of Banksia ericifolia. It is native to New South Wales....

B. verticillata
Banksia verticillata
Banksia verticillata, commonly known as Granite Banksia or Albany Banksia, is a species of shrub or tree of the genus Banksia in the Proteaceae family. It is native to the southwest of Western Australia and can reach up to 3 m in height. It can grow taller to 5 m in sheltered areas,...

B. seminuda
Banksia seminuda
Banksia seminuda, commonly known as the River Banksia, is a tree in the plant genus Banksia. It is found in south west Western Australia from Dwellingup to the Broke Inlet east of Denmark . It is often mistaken for and was originally considered a subspecies of the Banksia littoralis...

B. littoralis
Banksia littoralis
Banksia littoralis, commonly known as the Swamp Banksia, Swamp Oak, Pungura and the Western Swamp Banksia, is a tree in the plant genus Banksia. It is found in south west Western Australia from the south eastern metropolitan area of Perth to the Stirling Range and Albany...

B. occidentalis
Banksia occidentalis
The Red Swamp Banksia or Waterbush is a species of shrub or small tree in the plant genus Banksia. It occurs on the south coast of Western Australia in three disjunct populations: at Augusta, around Albany and in the Esperance area.A 1980 field study at Cheyne beach showed it to be pollinated by...

B. brownii
Banksia brownii
Banksia brownii, commonly known as Feather-leaved Banksia or Brown's Banksia, is a species of shrub that occurs in southwest Western Australia. An attractive plant with fine feathery leaves and large red-brown flower spikes, it usually grows as an upright bush around two metres high, but can also...

B. ser. Tricuspidae
B. tricuspis
Banksia tricuspis
The Lesueur Banksia or Pine Banksia is a species of shrub or tree in the plant genus Banksia. It occurs within a geographic range of just 15 square kilometres near Jurien, Western Australia.-External links:...

B. ser. Dryandroideae
Banksia ser. Dryandroideae
Banksia ser. Dryandroideae is a valid botanic name for a taxonomic series in the plant genus Banksia. First published by Carl Meissner in 1856, the name has had two circumscriptions. As presently circumscribed it is monotypic, containing only B. dryandroides.-According to Meissner:B. ser...

B. dryandroides
Banksia dryandroides
Banksia dryandroides, the Dryandra-leaved Banksia, is a species of small shrub in the plant genus Banksia. It occurs in shrubland, coastal heath and woodland on the south coast of Western Australia between Two Peoples Bay and Cheyne Bay. The species is placed alone in series B. ser...

B. ser. Abietinae
Banksia ser. Abietinae
Banksia ser. Abietinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

B. sphaerocarpa
Banksia sphaerocarpa
Banksia sphaerocarpa, commonly known as the Fox Banksia or Round-fruit Banksia, is a species of shrub or tree in the plant genus Banksia . It is generally encountered as a 1–2 m high shrub, and is usually smaller in the north of its range...

B. sphaerocarpa  var. sphaerocarpa
Banksia sphaerocarpa var. sphaerocarpa
Banksia sphaerocarpa var. sphaerocarpa is a variety of Banksia sphaerocarpa. It is native to the Southwest Botanical Province of Western Australia. As an autonym, it is defined as containing the type specimen of the species....

B. sphaerocarpa  var. caesia
Banksia sphaerocarpa var. caesia
Banksia sphaerocarpa var. caesia is a variety of Banksia sphaerocarpa. It is native to the Southwest Botanical Province of Western Australia....

B. sphaerocarpa  var. dolichostyla
Banksia sphaerocarpa var. dolichostyla
Banksia sphaerocarpa var. dolichostyla, commonly known as Ironcap Banksia, is a plant which is either considered a variety of Banksia sphaerocarpa, or as a species in its own right. It is native to the Southwest Botanical Province of Western Australia...

B. micrantha
Banksia micrantha
Banksia micrantha is a species of shrub in the plant genus Banksia. A small spreading bush with pale yellow flower spikes, it occurs between Eneabba and Cervantes in South west Western Australia...

B. grossa
Banksia grossa
Banksia grossa, commonly known as Coarse Banksia is a species of shrub in the plant family Proteaceae endemic to south west Western Australia. One of fourteen species of banksia with predominantly round or oval inflorescences of the series Abietinae, it was described in 1981 as a distinct species...

B. telmatiaea
Banksia telmatiaea
Banksia telmatiaea, commonly known as Swamp Fox Banksia or rarely Marsh Banksia, is a shrub that grows in marshes and swamps along the lower west coast of Australia. It grows as an upright bush up to 2 m tall, with narrow leaves and a pale brown flower spike, which can produce profuse...

B. leptophylla
Banksia leptophylla
The Slender-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs along the west coast of Western Australia from Gingin to Kalbarri. Before Alex George's revision of 1981, it was labelled informally as B. sphaerocarpa var. pinifolia or var...

B. leptophylla var. leptophylla
Banksia leptophylla var. leptophylla
Banksia leptophylla var. leptophylla is a variety of Banksia leptophylla. It is native to the Southwest Botanical Province of Western Australia. As an autonym, it is defined as containing the type specimen of the species....

B. leptophylla var. melletica
Banksia leptophylla var. melletica
Banksia leptophylla var. melletica is a variety of Banksia leptophylla. It is native to the Southwest Botanical Province of Western Australia....

B. lanata
Banksia lanata
The Coomallo Banksia is a species of shrub in the plant genus Banksia. It occurs within a range of less than 100 square kilometres between Eneabba and Mount Lesueur, Western Australia. It has roughly spherical inflorescences with flowers of cream to orange-brown colour. The leaves are linear and...

B. scabrella
Banksia scabrella
Banksia scabrella, commonly known as the Burma Road Banksia, is a species of woody shrub in the genus Banksia. It is classified in the series Abietinae, a group of several species of shrubs with small round or oval inflorescences...

B. violacea
Banksia violacea
Banksia violacea, commonly known as Violet Banksia, is a species of shrub or tree in the plant genus Banksia . It generally grows as a small shrub to 1.5 m high with fine narrow leaves, and is best known for its unusually coloured dark purple-violet inflorescences...

B. incana
Banksia incana
The Hoary Banksia is a species of small shrub in the plant genus Banksia. It occurs on sandplain heathland between Badgingarra and Eneabba in Western Australia, with outlying populations as far south as Perth. Seeds do not require any treatment, and take around 14 days to germinate....

B. laricina
Banksia laricina
The Rose-Fruited Banksia is a species of shrub in the plant genus Banksia endemic to southwestern Western Australia. It derives its specific Latin name from larix or larch, which its foliage is said to resemble. The common name comes from the striking fruits which resemble wooden roses...

B. pulchella
Banksia pulchella
The Teasel Banksia is a species of small shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from Fitzgerald River National Park east to Israelite Bay....

B. meisneri
Banksia meisneri
The Meisner's Banksia is a species of small shrub in the plant genus Banksia. It occurs in a number of isolated populations throughout southwest Western Australia. Seeds do not require any treatment, and take 28 to 39 days to germinate.-External links:...

B. meisneri subsp. meisneri
Banksia meisneri subsp. meisneri
Banksia meisneri subsp. meisneri is a subspecies of Banksia meisneri. It is native to the Southwest Botanical Province of Western Australia. As an autonym, it is defined as containing the type specimen of the species....

B. meisneri subsp. ascendens
Banksia meisneri subsp. ascendens
Banksia meisneri subsp. ascendens, commonly known as Scott River Banksia, is a subspecies of Banksia meisneri. It is native to the Southwest Botanical Province of Western Australia....

B. nutans
Banksia nutans
Banksia nutans, commonly known as Nodding Banksia, is a species of shrub native to the south coast of Western Australia in the genus Banksia...

B. nutans var. nutans
Banksia nutans var. nutans
Banksia nutans var. nutans is a variety of Banksia nutans. It is native to the Southwest Botanical Province of Western Australia. As an autonym, it is defined as containing the type specimen of the species....

B. nutans var. cernuella
Banksia nutans var. cernuella
Banksia nutans var. cernuella is a variety of Banksia nutans. It is native to the Southwest Botanical Province of Western Australia. Seeds do not require any treatment, and take around 17 days to germinate....

B. subg. Isostylis
Banksia subg. Isostylis
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser...

B. ilicifolia
Banksia ilicifolia
Banksia ilicifolia, commonly known as Holly-leaved Banksia, is a tree in the plant Proteaceae family. Endemic to southwest Western Australia, it belongs to Banksia subg. Isostylis, a subgenus of three closely related Banksia species with inflorescences that are dome-shaped heads rather than...

B. oligantha
Banksia oligantha
Banksia oligantha, commonly known as Wagin Banksia, is an endangered species in the plant family Proteaceae endemic to south west Western Australia. It belongs to Banksia subg. Isostylis, a subgenus of three closely related Banksia species with dome-shaped heads as inflorescences, rather than...

B. cuneata
Banksia cuneata
Banksia cuneata, commonly known as Matchstick Banksia or Quairading Banksia, is an endangered species of flowering plants in the Proteaceae family. Endemic to southwest Western Australia, it belongs to Banksia subg...


Legacy

George's arrangement was undisputed as the accepted arrangement between 1981 and 1996. It was replaced by the arrangement of Thiele and Ladiges in 1996, but this lasted only until 1999, when George published a revised version of his 1981 arrangement. This revised version was not universally accepted, however: some Australian herbaria
Herbarium
In botany, a herbarium – sometimes known by the Anglicized term herbar – is a collection of preserved plant specimens. These specimens may be whole plants or plant parts: these will usually be in a dried form, mounted on a sheet, but depending upon the material may also be kept in...

 continued to follow Thiele and Ladiges on some points, for example by recognising the four species that they promoted.

In 2005, Austin Mast, Eric Jones and Shawn Havery published the results of their cladistic analyses of DNA
DNA
Deoxyribonucleic acid is a nucleic acid that contains the genetic instructions used in the development and functioning of all known living organisms . The DNA segments that carry this genetic information are called genes, but other DNA sequences have structural purposes, or are involved in...

 sequence data for Banksia. They inferred a phylogeny very greatly different from George's taxonomic arrangement, including finding Banksia to be paraphyletic
Paraphyly
A group of taxa is said to be paraphyletic if the group consists of all the descendants of a hypothetical closest common ancestor minus one or more monophyletic groups of descendants...

 with respect to Dryandra. A new taxonomic arrangement was not published at the time, but early in 2007 Mast and Thiele initiated a rearrangement by transferring Dryandra to Banksia, and publishing B. subg. Spathulatae
Banksia subg. Spathulatae
Banksia subg. Spathulatae is a valid botanic name for a subgenus of Banksia. It was published in 2007 by Austin Mast and Kevin Thiele, and defined as containing all those Banksia species having spathulate cotyledons...

 for the species having spoon-shaped cotyledon
Cotyledon
A cotyledon , is a significant part of the embryo within the seed of a plant. Upon germination, the cotyledon may become the embryonic first leaves of a seedling. The number of cotyledons present is one characteristic used by botanists to classify the flowering plants...

s. They foreshadowed publishing a full arrangement once DNA sampling of Dryandra was complete.
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