Banksia ser. Quercinae
Encyclopedia
Banksia ser. Quercinae is a
valid botanic name for a series
Series (botany)
Series is a low-level taxonomic rank below that of section but above that of species.In botany, a series is a subdivision of a genus...

 of Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

. First published by Carl Meissner
Carl Meissner
Carl Daniel Friedrich Meissner was a Swiss botanist.Born in Bern, Switzerland on 1 November 1800, he was christened Meisner but later changed the spelling of his name to Meissner. For most of his 40 year career he was Professor of Botany at University of Basel...

 in 1856, the name has had three circumscriptions.

According to Meissner

B. ser. Quercinae was first published in 1856, in Carl Meissner
Carl Meissner
Carl Daniel Friedrich Meissner was a Swiss botanist.Born in Bern, Switzerland on 1 November 1800, he was christened Meisner but later changed the spelling of his name to Meissner. For most of his 40 year career he was Professor of Botany at University of Basel...

's chapter on the Proteaceae
Proteaceae
Proteaceae is a family of flowering plants distributed in the Southern Hemisphere. The family comprises about 80 genera with about 1600 species. Together with the Platanaceae and Nelumbonaceae they make up the order Proteales. Well known genera include Protea, Banksia, Embothrium, Grevillea,...

 in A. P. de Candolle
A. P. de Candolle
Augustin Pyramus de Candolle also spelled Augustin Pyrame de Candolle was a Swiss botanist. René Louiche Desfontaines launched Candolle's botanical career by recommending him at an herbarium...

's Prodromus systematis naturalis regni vegetabilis
Prodromus Systematis Naturalis Regni Vegetabilis
Prodromus Systematis Naturalis Regni Vegetabilis, also known by its standard botanical abbreviation Prodr. , is a 17-volume treatise on botany initiated by A. P. de Candolle. De Candolle intended it as a summary of all known seed plants, encompassing taxonomy, ecology, evolution and biogeography....

. It was one of four series into which the subgenus Eubanksia was divided. These four series were defined in terms of leaf characters, with series Quercinae containing the species with strongly dentate, cuneate to obovate leaves. As they were defined on leaf characters alone, all of Meissner's series were highly heterogeneous.

The placement and circumscription of B. ser. Quercinae in Meissner's arrangement
Meissner's taxonomic arrangement of Banksia
Carl Meissner's taxonomic arrangement of Banksia was published in 1856, as part of his chapter on the Proteaceae in A. P. de Candolle's Prodromus systematis naturalis regni vegetabilis. It was the first attempt to provide an infrageneric classification for the genus, aside from Robert Brown's...

 may be summarised as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. sect. Eubanksia
Banksia sect. Eubanksia
Banksia sect. Eubanksia is an obsolete section of Banksia. There have been two circumscriptions, one of which is synonymous with the recently abandoned B. subg. Banksia sensu Alex George, the other having no modern equivalent....

B. ser. Abietinae
Banksia ser. Abietinae
Banksia ser. Abietinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (8 species, 1 variety)
B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (23 species, 8 varieties)
B. ser. Quercinae
B. coccinea
Banksia coccinea
Banksia coccinea, commonly known as the Scarlet Banksia, Waratah Banksia or Albany Banksia, is an erect shrub or small tree in the plant genus Banksia...

B. sceptrum
Banksia sceptrum
Banksia sceptrum, the Sceptre Banksia, occurs in Western Australia near the central west coast from Geraldton north through Kalbarri to Hamelin Pool. It extends inland almost to Mullewa....

B. Baueri
Banksia baueri
The Woolly Banksia is a species of shrub in the plant genus Banksia. It occurs in southwest Western Australia north and east of Albany. It has a distinctively large and hairy looking inflorescence which can be 300 mm or more long and up to 200 mm in diameter.It is placed alone in series...

B. ornata
Banksia ornata
The Desert Banksia is a species of shrub in the plant genus Banksia which grows up to 3 m tall. It occurs in western Victoria, and in South Australia, where it is common on the Eyre Peninsula, Kangaroo Island and east of Adelaide, but is apparently absent from Yorke Peninsula.It tends to grow...

B. latifolia (now B. robur
Banksia robur
Banksia robur, commonly known as Swamp Banksia or, less commonly, Broad-leaved Banksia grows in sand or peaty sand in coastal areas from Cooktown in north Queensland to the Illawarra region on the New South Wales south coast...

)
B. marcescens (now B. praemorsa
Banksia praemorsa
The Cut-leaf Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in a few isolated populations on the south coast of Western Australia between Albany and Cape Riche. Seeds do not require any treatment, and take 30 to 49 days to germinate.-References:...

)
B. oblongifolia
Banksia oblongifolia
The Fern-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs along the eastern coast of Australia from Wollongong, New South Wales in the south to Rockhampton, Queensland in the north...

B. serrata
Banksia serrata
Banksia serrata, commonly known as Old Man Banksia, Saw Banksia, Saw-tooth Banksia and Red Honeysuckle, is a species of woody shrub or tree of the genus Banksia in the Proteaceae family. Native the east coast of Australia, it is found from Queensland through to Victoria with outlying populations on...

B. æmula
Banksia aemula
Banksia aemula, commonly known as the wallum banksia, is a lignotuberous shrub of the Proteaceae family. Found from Bundaberg south to Sydney on the Australian east coast, it is encountered as a shrub or a taller tree to 8 m in coastal heath on deep sandy soil, known as Wallum...

B. Caleyi
Banksia caleyi
Banksia caleyi, commonly known as the red lantern banksia or Cayley's banksia, is a species of shrub in the plant genus Banksia. It occurs in southern Western Australia south and east of the Stirling Ranges through to around Jerramungup...

B. caleyi var. sinuosa (now B. caleyi
Banksia caleyi
Banksia caleyi, commonly known as the red lantern banksia or Cayley's banksia, is a species of shrub in the plant genus Banksia. It occurs in southern Western Australia south and east of the Stirling Ranges through to around Jerramungup...

)
B. Lemanniana
Banksia lemanniana
Banksia lemanniana, commonly known as the Yellow lantern Banksia or Lemann's Banksia, is a species of woody shrub in the genus Banksia of the family Proteaceae. It generally grows as an open shrub or small tree to 5 m high with stiff serrated leaves, and unusually hanging inflorescences...

B. quercifolia
Banksia quercifolia
The Oak-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Manypeaks in the east.-Ecology:...

B. dentata
Banksia dentata
The Tropical Banksia is a species of small tree in the plant genus Banksia.-Distribution and habitat:It occurs throughout northern Australia, and also extends into New Guinea and the Aru Islands...

B. prostrata (now B. gardneri
Banksia gardneri
The Prostrate Banksia is a species of prostrate shrub in the plant genus Banksia. It occurs along the south coast of Western Australia.-Scientific name:...

)
B. Goodii
Banksia goodii
Banksia goodii, commonly known as Good's Banksia, is an endangered shrub of Southwest Western Australia.Good's Banksia grows as a low shrub, either prostrate or with stems up to twenty centimetres high. The leaves are dark green with a prominent yellow midrib, and are held erect. They may be up...

B. barbigera
B. repens
Banksia repens
Banksia repens, the Creeping Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Ragged in the east....

B. Solandri
Banksia solandri
Banksia solandri, commonly known as Stirling Range Banksia, is a species of large shrub in the plant genus Banksia. It occurs only within the Stirling Range in southwest Western Australia.-Description:...

B. solandri var. major (now B. solandri
Banksia solandri
Banksia solandri, commonly known as Stirling Range Banksia, is a species of large shrub in the plant genus Banksia. It occurs only within the Stirling Range in southwest Western Australia.-Description:...

)
B. ser. Dryandroideae
Banksia ser. Dryandroideae
Banksia ser. Dryandroideae is a valid botanic name for a taxonomic series in the plant genus Banksia. First published by Carl Meissner in 1856, the name has had two circumscriptions. As presently circumscribed it is monotypic, containing only B. dryandroides.-According to Meissner:B. ser...

 (8 species)
B. sect.Isostylis (1 species)


Meissner's arrangement was current until 1870, when George Bentham
George Bentham
George Bentham CMG FRS was an English botanist, characterized by Duane Isely as "the premier systematic botanist of the nineteenth century".- Formative years :...

 published his arrangement
Bentham's taxonomic arrangement of Banksia
George Bentham's taxonomic arrangement of Banksia was published in 1870, in Volume 5 of Bentham's Flora Australiensis. A substantial improvement on the previous arrangement, it would stand for over a century. It was eventually replaced by Alex George's 1981 arrangement, published in his classic...

, discarding all four of Meissner's series.

According to George

In 1981, Alex George
Alex George
Alexander Segger George is a Western Australian botanist. He is the authority on the plant genera Banksia and Dryandra...

 published a thorough revision of Banksia in his classic monograph The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. is a 1981 monograph by Alex George on the taxonomy of the plant genus Banksia. Published by the Western Australian Herbarium as Nuytsia 3, it presented George's taxonomic arrangement of Banksia, the first major taxonomic revision of the genus since George Bentham published...

. He reinstated B. ser. Quercinae, placing it within B. sect. Banksia
Banksia sect. Banksia
Banksia sect. Banksia is one of four sections of Banksia subgenus Banksia. It contains those species of subgenus Banksia with straight or sometimes curved but not hooked styles. These species all have cylindrical inflorescences and usually exhibit a bottom-up sequence of flower anthesis...

, and redefining it as containing those species with awned perianth
Perianth
The term perianth has two similar but separate meanings in botany:* In flowering plants, the perianth are the outer, sterile whorls of a flower...

s, and beaked follicle
Follicle (fruit)
In botany, a follicle is a dry unilocular many-seeded fruit formed from one carpel and dehiscing by the ventral suture in order to release seeds, such as in larkspur, magnolia, banksia, peony and milkweed....

s. Initially, the series contained three species, but in 1988 George moved B. baueri
Banksia baueri
The Woolly Banksia is a species of shrub in the plant genus Banksia. It occurs in southwest Western Australia north and east of Albany. It has a distinctively large and hairy looking inflorescence which can be 300 mm or more long and up to 200 mm in diameter.It is placed alone in series...

 (Woolly Orange Banksia) into its own series, B. ser. Bauerinae, leaving just B. quercifolia
Banksia quercifolia
The Oak-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Manypeaks in the east.-Ecology:...

 (Oak-leaved Banksia) and B. oreophila
Banksia oreophila
The Western Mountain Banksia or Mountain Banksia is a species of shrub in the plant genus Banksia. It occurs on the slopes and hilltops of the Stirling and Barren Ranges in southwest Western Australia.-Taxonomy:...

 (Western Mountain Banksia) in B. ser. Quercinae.

The placement and circumscription of B. ser. Quercinae in George's taxonomic arrangement of Banksia
George's taxonomic arrangement of Banksia
Alex George's taxonomic arrangement of Banksia was the first modern-day arrangement for that genus. First published in 1981 in the classic monograph The genus Banksia L.f. , it superseded the arrangement of George Bentham, which had stood for over a hundred years. It was overturned in 1996 by Kevin...

 may be summarised as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

B. sect. Banksia
Banksia sect. Banksia
Banksia sect. Banksia is one of four sections of Banksia subgenus Banksia. It contains those species of subgenus Banksia with straight or sometimes curved but not hooked styles. These species all have cylindrical inflorescences and usually exhibit a bottom-up sequence of flower anthesis...

B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (11 species, 7 subspecies)
B. ser. Grandes
Banksia ser. Grandes
Banksia ser. Grandes is a taxonomic series in the genus Banksia. It consists of two closely related species in section Banksia, both endemic to Western Australia. These are B. grandis and B. solandri ....

 (2 species)
B. ser. Banksia
Banksia ser. Banksia
Banksia ser. Banksia is avalid botanic name for a series of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-According to Bentham:Banksia ser. Banksia originated in 1870 as...

 (8 species)
B. ser. Crocinae
Banksia ser. Crocinae
Banksia ser. Crocinae is a taxonomic series in the genus Banksia. It consists of four closely related species, all of which are endemic to Western Australia; namely B. prionotes , B. burdettii , B. hookeriana and B. victoriae...

 (4 species)
B. ser. Prostratae
Banksia ser. Prostratae
Banksia ser. Prostratae is a taxonomic series in the genus Banksia. It consists of six closely related species in section Banksia, all endemic to Western Australia, with a prostrate habit.Banksia ser...

 (6 species, 3 varieties)
B. ser. Cyrtostylis
Banksia ser. Cyrtostylis
Banksia ser. Cyrtostylis is a valid botanic name for a taxonomic series within the plant genus Banksia. First published at sectional rank by George Bentham in 1870, it was demoted to a series by Alex George in 1981. The name has had three circumscriptions.-According to Bentham:Bentham published B....

 (13 species, 2 subspecies)
B. ser. Tetragonae
Banksia ser. Tetragonae
Banksia ser. Tetragonae is a taxonomic series in the genus Banksia. It consists of three closely related species of erect shrub with pendulous inflorescences in section Banksia. These are B. lemanniana , B. caleyi and B. aculeata ....

 (3 species)
B. ser. Bauerinae (1 species)
B. ser. Quercinae
B. quercifolia
Banksia quercifolia
The Oak-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Manypeaks in the east.-Ecology:...

B. oreophila
Banksia oreophila
The Western Mountain Banksia or Mountain Banksia is a species of shrub in the plant genus Banksia. It occurs on the slopes and hilltops of the Stirling and Barren Ranges in southwest Western Australia.-Taxonomy:...

B. sect. Coccinea (1 species)
B. sect. Oncostylis
Banksia sect. Oncostylis
Banksia sect. Oncostylis is one of four sections of subgenus Banksia subg. Banksia. It contains those Banksia species with hooked pistils. All of the species in Oncostylis also exhibit a top-down sequence of flower anthesis, except for Banksia nutans which is bottom-up.Banksia sect...

 (3 series, 22 species, 4 subspecies, 11 varieties)
B. subg. Isostylis
Banksia subg. Isostylis
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser...

 (3 species)

According to Thiele and Ladiges

In 1996, Kevin Thiele
Kevin Thiele
Kevin R. Thiele is curator of the Western Australian Herbarium. His research interests include the systematics of the plant families Proteaceae, Rhamnaceae and Violaceae, and the conservation ecology of grassy woodland ecosystems...

 and Pauline Ladiges undertook a cladistic
Cladistics
Cladistics is a method of classifying species of organisms into groups called clades, which consist of an ancestor organism and all its descendants . For example, birds, dinosaurs, crocodiles, and all descendants of their most recent common ancestor form a clade...

 analysis of morphological
Morphology (biology)
In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....

 characters of Banksia, which yielded a phylogeny somewhat at odds with George's taxonomic arrangement. B. ser. Quercinae was found to be monophyletic
Monophyly
In common cladistic usage, a monophyletic group is a taxon which forms a clade, meaning that it contains all the descendants of the possibly hypothetical closest common ancestor of the members of the group. The term is synonymous with the uncommon term holophyly...

, though quite closely related to B. ser. Spicigerae
Banksia ser. Spicigerae
Banksia ser. Spicigerae is a taxonomic series in the genus Banksia. It consists of the seven species in section Oncostylis that have cylindrical inflorescences. These range in form from small shrubs to tall trees. The leaves grow in either an alternate or whorled pattern, with various shape forms...

. They therefore retained the series, placing it next to B. ser. Spicigerae in their arrangement.

The placement and circumscription of B. ser. Quercinae in Thiele and Ladiges' arrangement
Thiele and Ladiges' taxonomic arrangement of Banksia
Kevin Thiele and Pauline Ladiges' taxonomic arrangement of Banksia, published in 1996, was a novel taxonomic arrangement that was intended to align the taxonomy of Banksia more closely with the phylogeny that they had inferred from their cladistic analysis of the genus...

 may be summarised as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. subg. Isostylis
Banksia subg. Isostylis
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser...

 (3 species)
B. elegans
Banksia elegans
The Elegant Banksia is a species of shrub or small tree in the plant genus Banksia. It occurs only over a 65 square kilometre area north and west of Eneabba, Western Australia.-Ecology:...

 (incertae sedis
Incertae sedis
, is a term used to define a taxonomic group where its broader relationships are unknown or undefined. Uncertainty at specific taxonomic levels is attributed by , , and similar terms.-Examples:*The fossil plant Paradinandra suecica could not be assigned to any...

)
B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

B. ser. Tetragonae
Banksia ser. Tetragonae
Banksia ser. Tetragonae is a taxonomic series in the genus Banksia. It consists of three closely related species of erect shrub with pendulous inflorescences in section Banksia. These are B. lemanniana , B. caleyi and B. aculeata ....

 (4 species)
B. ser. Lindleyanae (1 species)
B. ser. Banksia
Banksia ser. Banksia
Banksia ser. Banksia is avalid botanic name for a series of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-According to Bentham:Banksia ser. Banksia originated in 1870 as...

 (2 subseries, 12 species)
B. baueri
Banksia baueri
The Woolly Banksia is a species of shrub in the plant genus Banksia. It occurs in southwest Western Australia north and east of Albany. It has a distinctively large and hairy looking inflorescence which can be 300 mm or more long and up to 200 mm in diameter.It is placed alone in series...

 (incertae sedis)
B. lullfitzii
Banksia lullfitzii
Banksia lullfitzii is a species of shrub in the plant genus Banksia. A many-branched, spreading bush with golden-orange flowers, it occurs in scattered populations over a large area of the eastern goldfields of Western Australia...

 (incertae sedis)
B. attenuata
Banksia attenuata
Banksia attenuata, commonly known as the candlestick banksia or slender banksia, is a species of plant in the proteaceae family. Commonly a tree, it reaches 10 m high, but is often a shrub in dryer areas 0.4 to 2 m high...

 (incertae sedis)
B. ashbyi
Banksia ashbyi
The Ashby's Banksia is a species of shrub in the plant genus Banksia. It occurs in heath and spinifex country along the coast of Western Australia between Geraldton and Exmouth.-Description:...

 (incertae sedis)
B. coccinea
Banksia coccinea
Banksia coccinea, commonly known as the Scarlet Banksia, Waratah Banksia or Albany Banksia, is an erect shrub or small tree in the plant genus Banksia...

 (incertae sedis)
B. ser. Prostratae
Banksia ser. Prostratae
Banksia ser. Prostratae is a taxonomic series in the genus Banksia. It consists of six closely related species in section Banksia, all endemic to Western Australia, with a prostrate habit.Banksia ser...

 (8 species)
B. ser. Cyrtostylis
Banksia ser. Cyrtostylis
Banksia ser. Cyrtostylis is a valid botanic name for a taxonomic series within the plant genus Banksia. First published at sectional rank by George Bentham in 1870, it was demoted to a series by Alex George in 1981. The name has had three circumscriptions.-According to Bentham:Bentham published B....

 (4 species)
B. ser. Ochraceae
Banksia ser. Ochraceae
Banksia ser. Ochraceae is a valid botanic name for a taxonomic series within the plant genus Banksia. It was published by Kevin Thiele in 1996, but discarded by Alex George in 1999.-Cladistics:...

 (3 species, 2 subspecies)
B. ser. Grandes
Banksia ser. Grandes
Banksia ser. Grandes is a taxonomic series in the genus Banksia. It consists of two closely related species in section Banksia, both endemic to Western Australia. These are B. grandis and B. solandri ....

 (2 species)
B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (2 subseries, 11 species, 4 subspecies)
B. ser. Spicigerae
Banksia ser. Spicigerae
Banksia ser. Spicigerae is a taxonomic series in the genus Banksia. It consists of the seven species in section Oncostylis that have cylindrical inflorescences. These range in form from small shrubs to tall trees. The leaves grow in either an alternate or whorled pattern, with various shape forms...

 (3 subseries, 7 species, 6 varieties)
B. ser. Quercinae
B. quercifolia
Banksia quercifolia
The Oak-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Manypeaks in the east.-Ecology:...

B. oreophila
Banksia oreophila
The Western Mountain Banksia or Mountain Banksia is a species of shrub in the plant genus Banksia. It occurs on the slopes and hilltops of the Stirling and Barren Ranges in southwest Western Australia.-Taxonomy:...

B. ser. Dryandroideae
Banksia ser. Dryandroideae
Banksia ser. Dryandroideae is a valid botanic name for a taxonomic series in the plant genus Banksia. First published by Carl Meissner in 1856, the name has had two circumscriptions. As presently circumscribed it is monotypic, containing only B. dryandroides.-According to Meissner:B. ser...

 (1 species)
B. ser. Abietinae
Banksia ser. Abietinae
Banksia ser. Abietinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (4 subseries, 15 species, 8 varieties)


Thiele and Ladiges' arrangement remained current only until 1999, when George's treatment of the genus for the Flora of Australia
Flora of Australia (series)
The Flora of Australia is a 59 volume series describing the vascular plants, bryophytes and lichens present in Australia and its external territories...

 series of monographs was published. This was essentially a revision of George's 1981 arrangement, which took into account some of Thiele and Ladiges' data, but rejected their overall arrangement. With respect to B. ser. Quercinae, George's 1999 arrangement was no different to that of 1988.

Recent developments

Since 1998, Austin Mast
Austin Mast
Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently an associate professor within the Department of Biological Science at Florida State University , and has been director of FSU's since August 2003.One of his...

 has been publishing results of ongoing cladistic analyses of DNA sequence
DNA sequence
The sequence or primary structure of a nucleic acid is the composition of atoms that make up the nucleic acid and the chemical bonds that bond those atoms. Because nucleic acids, such as DNA and RNA, are unbranched polymers, this specification is equivalent to specifying the sequence of...

 data for the subtribe Banksiinae. His analyses suggest a phylogeny that is very greatly different to George's taxonomic arrangement, including finding Banksia to be paraphyletic
Paraphyly
A group of taxa is said to be paraphyletic if the group consists of all the descendants of a hypothetical closest common ancestor minus one or more monophyletic groups of descendants...

 with respect to Dryandra
Dryandra
Banksia ser. Dryandra is a series of 94 species of shrub to small tree in the plant genus Banksia. It was considered a separate genus named Dryandra until early 2007, when it was merged into Banksia on the basis of extensive molecular and morphological evidence that Banksia was paraphyletic with...

. With respect to B. ser. Quercinae, however, Mast's analysis concurs with previous arrangements, placing its two species alone in a clade that is fairly widely separated from other clades.

Early in 2007 Mast and Thiele initiated a rearrangement of Banksia by transferring Dryandra into it, and publishing B. subg. Spathulatae
Banksia subg. Spathulatae
Banksia subg. Spathulatae is a valid botanic name for a subgenus of Banksia. It was published in 2007 by Austin Mast and Kevin Thiele, and defined as containing all those Banksia species having spathulate cotyledons...

 for the species having spoon-shaped cotyledon
Cotyledon
A cotyledon , is a significant part of the embryo within the seed of a plant. Upon germination, the cotyledon may become the embryonic first leaves of a seedling. The number of cotyledons present is one characteristic used by botanists to classify the flowering plants...

s. The members of B. ser. Quercinae fall within B. subg. Spathulatae, but no further details have been proffered. Mast and Thiele have foreshadowed publishing a full arrangement once DNA sampling of Dryandra is complete.
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