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Woolly Rhinoceros

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Woolly Rhinoceros



 
 
The woolly rhinoceros (Coelodonta antiquitatis) is an extinct species of rhinoceros
Rhinoceros

Rhinoceros , often colloquially abbreviated rhino, is a name used to group five extant species of odd-toed ungulates in the family Rhinocerotidae....
 native to the northern steppes of Eurasia
Eurasia

Eurasia is a large landmass covering about 53,990,000 km? or about 10.6% of the Earth's surface . Often considered a single continent, Eurasia comprises the traditional continents of Europe and Asia, concepts which date back to classical antiquity and the borders for which are somewhat arbitrary....
 that lived during the Pleistocene
Pleistocene

The Pleistocene is the epoch from 1.8 million to 10,000 years Before Present covering the world's recent period of repeated glaciations. The name pleistocene is derived from the Greek and ....
 epoch
Epoch

Periodization* Epoch - A defining moment in the beginning of, or characteristic of, a distinctive historical period or era.* On the geologic time scale, a span of time smaller than a "period" and larger than an "age"....
 and survived the last glacial period. The woolly rhinoceros are members of the Pleistocene megafauna
Pleistocene megafauna

Pleistocene megafauna is the set of species of large animals — mammals, birds and reptiles — that lived on Earth during the Pleistocene epoch and went extinct in a Quaternary extinction event....
.

dult wooly rhinoceros was 3.7 metres (12 feet) in length. Two horns on the skull were made of keratin
Keratin

Keratins are a family of fibrous protein; tough and insoluble, they form the hard but mineral structures found in reptiles, birds, amphibians and mammals....
, the anterior horn being 1 metre (3 feet) in length, with a smaller horn between its eyes.






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The woolly rhinoceros (Coelodonta antiquitatis) is an extinct species of rhinoceros
Rhinoceros

Rhinoceros , often colloquially abbreviated rhino, is a name used to group five extant species of odd-toed ungulates in the family Rhinocerotidae....
 native to the northern steppes of Eurasia
Eurasia

Eurasia is a large landmass covering about 53,990,000 km? or about 10.6% of the Earth's surface . Often considered a single continent, Eurasia comprises the traditional continents of Europe and Asia, concepts which date back to classical antiquity and the borders for which are somewhat arbitrary....
 that lived during the Pleistocene
Pleistocene

The Pleistocene is the epoch from 1.8 million to 10,000 years Before Present covering the world's recent period of repeated glaciations. The name pleistocene is derived from the Greek and ....
 epoch
Epoch

Periodization* Epoch - A defining moment in the beginning of, or characteristic of, a distinctive historical period or era.* On the geologic time scale, a span of time smaller than a "period" and larger than an "age"....
 and survived the last glacial period. The woolly rhinoceros are members of the Pleistocene megafauna
Pleistocene megafauna

Pleistocene megafauna is the set of species of large animals — mammals, birds and reptiles — that lived on Earth during the Pleistocene epoch and went extinct in a Quaternary extinction event....
.

Appearance

An adult wooly rhinoceros was 3.7 metres (12 feet) in length. Two horns on the skull were made of keratin
Keratin

Keratins are a family of fibrous protein; tough and insoluble, they form the hard but mineral structures found in reptiles, birds, amphibians and mammals....
, the anterior horn being 1 metre (3 feet) in length, with a smaller horn between its eyes. It had thick, long fur
Fur

Fur is a Hair of any non-human mammal, also known as the pelage. It may consist of short ground hair, long guard hair, and, in some cases, medium awn hair....
, small ears, short, thick legs, and a stocky body. Cave paintings suggest a wide dark band between the front and hind legs, but it is not universal and identification of rhinoceros as woolly rhinoceros is uncertain. The woolly rhinoceros used its horns to sweep snow away from vegetation so it could eat in the winter, and is also thought to have used its horns for defensive purposes and to attract mates.

As the last and most derived member of the Pleistocene
Pleistocene

The Pleistocene is the epoch from 1.8 million to 10,000 years Before Present covering the world's recent period of repeated glaciations. The name pleistocene is derived from the Greek and ....
 rhinoceros lineage
Lineage

Lineage may refer to:In science:* Lineage , descent group that can demonstrate their common descent from an apical ancestor* Lineage , group composed of species, taxa, or individuals related by descent from a common ancestor...
, the woolly rhinoceros was supremely well adapted to its environment. Stocky limbs and thick woolly pelage
Pelage

In mammals, pelage is the hair, fur, or wool that covers the animal. In many mammals, the pelage is made up of more than one type of hair. Some of the most prominent types of hair that make up the pelage include guard hairs , bristles , and the underfur, which traps in air to maintain temperature....
 made it well suited to the steppe-tundra environment prevalent across the Palearctic ecozone during the Pleistocene glaciation
Glacier

A glacier is a large, slow-moving mass of ice, formed from compacted layers of snow, that slowly deforms and flows in response to gravity and high pressure....
s. Its geographical range expanded and contracted with the alternating cold and warm cycles, forcing populations to migrate as glaciers receded. Like the vast majority of rhinoceroses, the body plan of the woolly rhinoceros adhered to the conservative morphology
Morphology (biology)

The term morphology in biology refers to form, structure and configuration of an organism. This includes aspects of the outward appearance as well as the form and structure of the internal parts like bones and organs....
, like the first rhinoceroses seen in the late Eocene. A close relative, the Giant Unicorn
Elasmotherium

Elasmotherium was a genus of giant rhinoceros which stood, on average, high and long, with a single two-meter-long horn in the forehead. The animal may have weighed up to ....
 (Elasmotherium), had a more southern range.

Diet

Controversy has long surrounded the precise dietary preference of Coelodonta as past investigations have found both grazing
Grazing

Grazing generally describes a type of predation in which a herbivore feeds on plants , or more broadly on a multicellular autotrophs . Grazing differs from true predation because the organism being eaten is not death, and it differs from parasitism as the two organisms do not symbiosis, nor is the grazer necessarily so limited in what it can...
 and browsing
Browsing (predation)

Browsing is a type of predation in which a herbivore feeds on high growing plants such as leaves, soft shoots, or fruits. This is contrasted with grazing predation, usually associated with animals feeding on grass or other low vegetation....
 modes of life to be plausible. The palaeodiet of the woolly rhinoceros has been reconstructed using several lines of evidence. Climatic reconstructions indicate the preferred environment to have been cold and arid steppe-tundra, with large herbivores forming an important part of the feedback
Feedback

Feedback describes the situation when output from an event or phenomenon in the past will influence the same event/phenomenon in the present or future....
 cycle. Pollen analysis
Pollen analysis

Analysis of the distribution of pollen grains of various species contained in surface layer deposits, especially peat bogs and lake sediments, from which a record of Paleoclimatology may be inferred....
 shows a prevalence of grasses and sedge
Cyperaceae

The family Cyperaceae, or the sedges, is a taxon of monocotyledon flowering plants that superficially resemble Poaceae or Juncaceae. The family is large, with some 4,000 species described in about 70 genera....
s within a more complicated vegetation mosaic. A strain vector biomechanical
Biomechanics

Biomechanics is the application of mechanical principles to living organisms. This includes bioengineering, the research and analysis of the mechanics of living organisms and the application of engineering principles to and from biological systems....
 investigation of the skull
Skull

The skull is a bone structure found in the head of many animals. The skull supports the structures of the face and protects the head against injury....
, mandible
Mandible

The mandible or inferior maxillary bone forms the lower jaw and holds the lower tooth in place. It also refers to both the upper and lower sections of the beaks of birds....
 and teeth
Tooth

Teeth are small whitish structures found in the jaws of many vertebrates that are used to tear, scrape, and chew food. Some animals, particularly carnivores, also use teeth for hunting or defense....
 of a well-preserved last cold stage individual recovered from Whitemoor Haye
Whitemoor Haye

Whitemoor Haye is a mainly agriculture area in the floodplain of the River Tame, West Midlands, near Alrewas in Staffordshire, England.It is very popular with Birdwatching, as several rare species are either resident or seasonal visitors, notably Corn Bunting....
, Staffordshire
Staffordshire

Staffordshire is a landlocked Counties of England in the West Midlands region of England. The county town is Stafford. Part of the National Forest, England lies within its borders....
, revealed musculature
MUSCLE

MUSCLE is public domain, multiple sequence alignment software for protein and nucleotide sequences.MUSCLE is integrated into UGENE bioinformatics tool as a plugin....
 and dental characteristics that support a grazing feeding preference. In particular, the enlargement of the temporalis
Temporalis muscle

The temporalis muscle is one of the Mastication#Muscles of mastication....
 and neck muscles is consistent with that required to resist the large tugging forces generated when taking large mouthfuls of fodder
Fodder

In agriculture, fodder or animal feed is any foodstuff that is used specifically to feed domesticated livestock, such as cattle, goats, sheep, horses, chickens and pigs....
 from the ground. The presence of a large diastema
Dentistry

Dentistry is the known evaluation, diagnosis, prevention, and treatment of diseases, disorders and conditions of the mouth, maxillofacial area and the adjacent and associated structures and their impact on the human body....
 supports this theory.

Comparisons with extant perissodactyls confirm that Coelodonta was a hindgut
Hindgut

The hindgut is the posterior part of the alimentary canal. It includes the distal third of the transverse colon and the splenic flexure, the descending colon, sigmoid colon, rectum and upper part of the anal canal....
 fermentor with a single stomach, and as such would have grazed upon cellulose
Cellulose

File:Cellulose Sessel.svgCellulose is an organic compound with the chemical formula , a polysaccharide consisting of a linear chain of several hundred to over ten thousand ? linked D-glucose units....
-rich, protein
Protein

Proteins are organic compounds made of amino acids arranged in a linear chain and joined together by peptide bonds between the carboxyl and amino groups of adjacent amino acid Residue ....
-poor fodder. This method of digestion would have required a large throughput of food and thus links the large mouthful size to the low nutritive content of the chosen grasses and sedges.

Extinction

Wooly Rhinoceros
Many species of Pleistocene megafauna
Megafauna

The term megafauna has two distinct meanings in the biological sciences. The less commonly found meaning is of any animal which can be seen with the unaided eye, in contrast to microfauna....
, like the wooly rhinoceros, became extinct around the same time period. Human and Neanderthal
Neanderthal

The Neanderthal , or Neandertal, is an extinct member of the Homo genus that is known from Pleistocene specimens found in Europe and parts of western and central Asia....
 hunting
Hunting

Hunting is the practice of pursuing living animals for food, recreation, or trade. In present-day use, the term refers to lawful hunting, as distinguished from poaching, which is the killing, trapping or capture of the hunted species contrary to law....
 is often cited as one cause. Other theories for the cause of the extinctions are climate change associated with the receding Ice age
Ice age

The general term "ice age" or, more precisely, "glacial age" denotes a geological period of long-term reduction in the temperature of the Earth's surface and atmosphere, resulting in an expansion of continental ice sheets, polar ice sheets and alpine glaciers....
 and the hyperdisease hypothesis.

Its shape was known only from prehistoric cave drawings until a completely preserved specimen (missing only the fur and hooves) was discovered in a tar pit
Tar pit

A tar pit, or more accurately known as an asphalt pit, is a geology occurrence where subterranean bitumen leaks to the surface, creating a large puddle, pit, or lake of asphalt....
 in Starunia, Poland
Poland

Poland , officially the Republic of Poland , is a country in Central Europe. Poland is bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian Enclave and exclave, to the north....
. The specimen, an adult female, is now on display in the Polish Academy of Sciences
Polish Academy of Sciences

The Polish Academy of Sciences, headquartered in Warsaw, is one of two Polish institutions having the nature of an academy of sciences....
's Museum of Natural History in Kraków
Kraków

Krak?w , in English also spelled Krakow or Cracow , is one of the largest and oldest cities in Poland, with a population of 756,336 in 2007 ....
. The woolly rhinoceros roamed much of Northern Europe
Northern Europe

Northern Europe is the northern part or region of Europe. The United Nations defines Northern Europe as including the following countries and dependent regions:...
 and was common in the then cold, arid desert that is southern England
Southern England

Southern England is an imprecise term used to refer to the southern counties of England. Differing usages apply the term with varying geographic extents....
 and the North Sea
North Sea

The North Sea is a marginal sea, epeiric sea on the European continental shelf. The Dover Strait and the English Channel in the south and the Norwegian Sea in the north connect it to the Atlantic Ocean....
 today. During Greenland Stadial 2 (The Last Glacial Maximum
Last Glacial Maximum

The Last Glacial Maximum refers to the time of maximum extent of the ice sheets during the last glaciation , approximately 20,000 years ago. This extreme persisted for several thousand years....
) the North Sea did not exist as sea levels were up to lower than today.

The woolly rhinoceros co-existed with woolly mammoth
Woolly mammoth

The woolly mammoth , also called the tundra mammoth, is an extinct species of mammoth. This animal is known from bones and frozen carcasses from northern North America and northern Eurasia with the best preserved carcasses in Siberia....
s and several other extinct larger mammals. No specimens have been dated in the U.K. after 15,000 14C
Radiocarbon dating

Radiocarbon dating, or carbon dating, is a radiometric dating method that uses the naturally occurring radioisotope carbon-14 to determine the age of carbonaceous materials up to about 60,000 years....
 years B.P. Recent radiocarbon dating
Radiocarbon dating

Radiocarbon dating, or carbon dating, is a radiometric dating method that uses the naturally occurring radioisotope carbon-14 to determine the age of carbonaceous materials up to about 60,000 years....
 indicates that populations survived as recently as 8,000 B.C.
8th millennium BC

In the 8th millennium BC, agriculture becomes widely practiced in the Fertile Crescent and Anatolia.Pottery becomes widespread and animal husbandry spreads to Africa and Eurasia....
 in Western Siberia
Siberia

Siberia , is the name given to the vast region constituting almost all of North Asia and for the most part currently serving as the massive central and eastern portion of the Russian Federation, having served in the same capacity previously for the Soviet Union from its beginning, and the Russian Empire beginning in the 16th century....
. However, the accuracy of this date is uncertain as several radiocarbon plateaus exist around this time. The extinction does not coincide with the end of the last ice age but does coincide however, with a minor yet severe climatic reversal that lasted for about 1,000–1,250 years, the Younger Dryas
Younger Dryas

The Younger Dryas stadial, named after the alpine/tundra wildflower Dryas octopetala, and also referred to as the Big Freeze, was a brief cold climate period following the B?lling/Aller?d Oscillation interstadial at the end of the Pleistocene between approximately 12,800 to 11,500 years Before Present, and preceding the Boreal of t...
 (GS1 - Greenland Stadial 1), characterized by glacial readvances and severe cooling globally, a brief interlude in the continuing warming subsequent to the termination of the last major ice age (GS2), thought to have been due to a shutdown of the thermohaline circulation
Thermohaline circulation

The term thermohaline circulation refers to the part of the large-scale ocean circulation that is driven by global Density gradient created by surface heat and freshwater Flux....
 in the ocean due to huge influxes of cold, fresh water from the preceding sustained glacial melting during the warmer Interstadial (GI1 - Greenland Interstadial 1 - ca. 16,000 - 11,450 14C years B.P.).

A relative, the hairy sumatran rhinoceros
Sumatran Rhinoceros

The Sumatran Rhinoceros is a member of the family Rhinocerotidae and one of five extant rhinoceroses. It is the smallest rhinoceros, standing about 120?145 centimetres high at the shoulder, with a body length of and weight of 500?800 kilograms ....
 (Dicerorhinus sumatrensis), still lives in Southeast Asia
Southeast Asia

Southeast Asia or Southeastern Asia is a subregion of Asia, consisting of the countries that are geographically south of China, east of India and north of Australia....
, as an endangered species
Endangered species

An endangered species is a population of an organism which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters....
.

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