Uria
Encyclopedia
Uria is a genus of seabird
Seabird
Seabirds are birds that have adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding niches have resulted in similar adaptations...

s in the auk
Auk
An auk is a bird of the family Alcidae in the order Charadriiformes. Auks are superficially similar to penguins due to their black-and-white colours, their upright posture and some of their habits...

 family known in Britain
United Kingdom
The United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages...

 as guillemot
Guillemot
Guillemots is the common name for several species of seabird in the auk family . In British use, the term comprises two genera: Uria and Cepphus. In North America the Uria species are called "murres" and only the Cepphus species are called "guillemots"...

s
, in most of North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 as murres, and in Newfoundland and Labrador
Newfoundland and Labrador
Newfoundland and Labrador is the easternmost province of Canada. Situated in the country's Atlantic region, it incorporates the island of Newfoundland and mainland Labrador with a combined area of . As of April 2011, the province's estimated population is 508,400...

 as turr. These are medium-sized birds with mainly brown or black plumage in the breeding season. They breed on the coasts of the northern Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 and Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

s.

There are two species:
  • Common Murre or Common Guillemot, Uria aalge
  • Thick-billed Murre or Brünnich's Guillemot, Uria lomvia


Uria auks are relatives of the Razorbill
Razorbill
The Razorbill is colonial seabird that will only come to land in order to breed. It is the largest living member of the Auk family. This agile bird will choose only one partner for life and females will lay one egg per year. Razorbills will nest along coastal cliffs in enclosed or slightly exposed...

, Little Auk
Little Auk
The Little Auk, or Dovekie , is a small auk, the only member of the genus Alle. It breeds on islands in the high Arctic. There are two subspecies: A. a. alle breeds in Greenland, Iceland, Novaya Zemlya and Spitsbergen, and A. a...

 and the extinct Great Auk
Great Auk
The Great Auk, Pinguinus impennis, formerly of the genus Alca, was a large, flightless alcid that became extinct in the mid-19th century. It was the only modern species in the genus Pinguinus, a group of birds that formerly included one other species of flightless giant auk from the Atlantic Ocean...

 and together make up the tribe
Tribe (biology)
In biology, a tribe is a taxonomic rank between family and genus. It is sometimes subdivided into subtribes.Some examples include the tribes: Canini, Acalypheae, Hominini, Bombini, and Antidesmeae.-See also:* Biological classification* Rank...

 Alcini. Despite the similar British common names, they are not so closely related to the Cepphus
Cepphus
Cepphus is a genus of seabirds in the auk family also referred to as true guillemots or, in North America, simply as guillemots. These are medium-sized birds with mainly black plumage in the breeding season, thin dark bills and red legs and feet. Two species have white wing patches, the third has...

guillemots, which form the tribe Cepphini.

These birds breed in large colonies
Bird colony
A bird colony is a large congregation of individuals of one or more species of bird that nest or roost in close proximity at a particular location. Many kinds of birds are known to congregate in groups of varying size; a congregation of nesting birds is called a breeding colony...

 on coastal cliffs, laying single elongated conical egg
Egg (biology)
An egg is an organic vessel in which an embryo first begins to develop. In most birds, reptiles, insects, molluscs, fish, and monotremes, an egg is the zygote, resulting from fertilization of the ovum, which is expelled from the body and permitted to develop outside the body until the developing...

s directly on cliff ledges. They move
Bird migration
Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. Sometimes, journeys are not termed "true migration" because they are irregular or in only one direction...

 south in winter to keep in ice-free waters.

They dive for food from the surface, swimming underwater and being among the deepest divers of all birds, using their stubby wings to routinely dive to more than 100 meters, and feeding on fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

 and crustaceans, also some molluscs, insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s and plant material.

Adult birds are black or brown on the head, neck, back and wings with white underparts. The bill is long and pointed. They have a small rounded black tail. The lower face becomes white in winter.

The flight is strong and direct, and these species have fast wing beats due to the short wings.

Uria guillemots produces a variety of harsh cackling calls at the breeding colonies, but are silent at sea.

Some prehistoric Uria species are also known:
  • Uria bordkorbi (Monterey
    Monterey Formation
    The Monterey Formation is an extensive Miocene oil-rich geological sedimentary formation in California, with massive outcroppings of the formation in areas of the California Coast Ranges, Peninsular Ranges, and on some of California's off-shore islands...

     or Sisquoc
    Sisquoc Formation
    The Sisquoc Formation is a sedimentary geologic unit widespread in Southern California, both on the coast and in mountains near the coast. Overlying the Monterey Formation, it is of upper Miocene and lower Pliocene age...

    , Late Miocene
    Miocene
    The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene...

     of Lompoc, USA)
  • Uria affinis (Late Pleistocene
    Pleistocene
    The Pleistocene is the epoch from 2,588,000 to 11,700 years BP that spans the world's recent period of repeated glaciations. The name pleistocene is derived from the Greek and ....

     of E USA) - possibly a subspecies of U. lomvia
  • Uria paleohesperis (Late Miocene of W USA)


U. brodkorbi is interesting insofar as it is the only known occurrence of the Alcini tribe in the temperate to subtropical Pacific, except for the very fringe of the range of U. aalge. It suggests that the Uria species, which are the sister taxon to all the other Alcini, and like them are usually believed to have evolved in the Atlantic, may have evolved in the Caribbean
Caribbean
The Caribbean is a crescent-shaped group of islands more than 2,000 miles long separating the Gulf of Mexico and the Caribbean Sea, to the west and south, from the Atlantic Ocean, to the east and north...

 or possibly close to the Isthmus of Panama
Isthmus of Panama
The Isthmus of Panama, also historically known as the Isthmus of Darien, is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country of Panama and the Panama Canal...

. The modern Pacific distribution would then be part of a later arctic
Arctic
The Arctic is a region located at the northern-most part of the Earth. The Arctic consists of the Arctic Ocean and parts of Canada, Russia, Greenland, the United States, Norway, Sweden, Finland, and Iceland. The Arctic region consists of a vast, ice-covered ocean, surrounded by treeless permafrost...

 expansion, whereas most other auk lineages form clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

s with a continuous range in the Pacific, from arctic to subtropical waters.

Origin of names

The name "guillemot" is of French
French language
French is a Romance language spoken as a first language in France, the Romandy region in Switzerland, Wallonia and Brussels in Belgium, Monaco, the regions of Quebec and Acadia in Canada, and by various communities elsewhere. Second-language speakers of French are distributed throughout many parts...

 origin, a diminutive
Diminutive
In language structure, a diminutive, or diminutive form , is a formation of a word used to convey a slight degree of the root meaning, smallness of the object or quality named, encapsulation, intimacy, or endearment...

 of the personal name Guillaume (William). Earlier common names such as "willock" and "wilkie" have the same root.

The name "murre" comes from the Cornish
Cornish language
Cornish is a Brythonic Celtic language and a recognised minority language of the United Kingdom. Along with Welsh and Breton, it is directly descended from the ancient British language spoken throughout much of Britain before the English language came to dominate...

 word for the Razorbill
Razorbill
The Razorbill is colonial seabird that will only come to land in order to breed. It is the largest living member of the Auk family. This agile bird will choose only one partner for life and females will lay one egg per year. Razorbills will nest along coastal cliffs in enclosed or slightly exposed...

.
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