Hide beetle
Encyclopedia
The or hide beetles are a family of beetle
s with a distinctive warty or bumpy appearance. Found worldwide, the family includes about 300 species contained in three genera.
Trogids range in length from 2.5 to 20.0 mm. Their shape is oblong to oval, with a generally flat abdomen. Their color ranges from brown to gray or black, often obscured with a dirt encrusting. Otherwise they resemble scarab beetle
s, with heavy limbs and spurs.
They are scavengers, being among the last to visit and feed on the dried-out remains of dead animals; both adults and larvae will eat feathers, fur, and skin. They may also be found in bird and mammal nests. Details of species' life histories is usually poorly known, since many are specialized to particular types of nests. Between their covering of dirt and a habit of becoming motionless when disturbed, they are often overlooked, both by predators and by collectors.
The taxonomic position of these beetles is somewhat unsettled, with many authorities placing them as a subfamily Troginae of the Scarabaeidae
. The common name "skin beetle" is sometimes used, but that name is usually used of the Dermestidae.
, Omorgus
, and Polynoncus
. These three genera are declared as the only known genera in the ‘New World
’; with Trox described by Fabricius
in 1775, Omorgus described by Erichson
in 1847, and Polynoncus described by Burmeister
in 1876.
The family of Trogidae favors dry environments over moist environments and therefore is often found within temperate and plains areas. Each particular genus is found in different regions of the world, such as the Trox genus is found in the Holarctic
/Ethiopian area; the Omorgus genus is typically more within the southern continents; and the Polynoncus genus is found in South America.
Specifically the beetles are found within the pellet
of any variety of animal, surrounding carrion or other decaying dry matter, and around birds’ and mammals’ nests and feathers as well as aging bones.
The origins and classifications for the family of Trogidae are very controversial but recent NA literature depicts Trogidae as its own family rather than a subfamily of Scarabaeidae
. One major reason for the dispute between classifications is the possible evolution of the ommatidium
in the eyes. The different environments, predators, etc. probably led to the adaptation of ommatidium structures within this family. For example the more advanced and numerous the ommatidium the more present the larger the ability of the insect to escape and elude predators. Due to these similarities many call the family of Scarabaeidae a ‘superfamily’ to the family of Trogidae. It is believed that Trogidae hails from Australia.
cranium that is almost black in color. The abdominal segments have at least one or more transverse rows of setae.
The Trogidae family is predaceous in addition to being scavengers. The pupae, larval, and adult stages of life have all been documented to be cannibalistic. Because of this tendency, the adult beetles will postpone, or delay pupation in an effort to maintain their safety and find a safe pupation site. The adult hide beetle produces a pheromone
in its feces that leads other adults and larvae to food sources. Studies have shown that this species is not cannibalistic due to over-crowding of populations, but just a food source preference.
Beetles of the Trogidae family have also been found to feed off of carcasses in the wild that have died and are decomposing. In one lab experiment done in 1998 by the Department of Zoology at the University of Melbourne, the hide beetle ate all tissues on a sheep carcass and left the bones.
When carcasses are not available their diet consists of eating the dry remains of dead mammals and birds in later to last stages of decomposition. Since they are usually the last at the scene, they can be found eating feathers, fur, skin, feces and anything else they can scavenge. Trogidae Omorgus candidus or any beetle in the family Trogidae is a scavenger type beetle that all have the same diet and predators for all in this family are the same.
. As the last succession of insects to appear on the carcass, both larvae and adults can be found feeding on the dry remains. At the site of the carcass, an impregnated female will dig small, vertical columns underneath the carcass to lay her eggs allowing the larvae to locate food after hatching.
Upon maturation of the larvae (approximately 6–8 weeks), females and males will mate. The male will sense odor cues from the female about her fertility. The male will mount the female and begin copulation. Copulation will be ended by the male. Males will tend to mate 6-7 times in their life span. Competition between the males for the female occurs occasionally as well as intra-sexual copulation between males. Males and females are polygamous.
There is little known about the life cycle of the Trogidae specifically. Their life cycles are very similar to the other genera of Scarabaeoidea (i.e. Passalidae and Lucanidae). Trogidae are holometabolous and usually have 3-5 instar
s. After impregnation of the female by the male, the female will lay the eggs and the larvae will hatch after an unknown amount of time. The larvae will usually molt twice growing and maturing until pupation. After pupation, the exoskeleton
is finally formed and further growth will cease.
estimates. The adults lay their eggs in vertical burrows in the soil beneath carcasses.
Various species of Trogidae have been used by museums to clean up skeletons by eating any remaining dried material left on the skeletons leaving them clean for display. This method of bone-stripping has been used by some museums for many years as it is the most effective method.
of Trogidae. There is a current study involving advancements at the Chinese Academy of Sciences
in efforts to further our knowledge on the classification of this family of beetles.
African Trogidae are being studied through the University of Pretoria
on the forensic importance of these beetles, as well as other carrion-associated beetles. Their article discusses how the presence of beetles on carrion effects the infestation of other arthropods in Africa.
Beetle
Coleoptera is an order of insects commonly called beetles. The word "coleoptera" is from the Greek , koleos, "sheath"; and , pteron, "wing", thus "sheathed wing". Coleoptera contains more species than any other order, constituting almost 25% of all known life-forms...
s with a distinctive warty or bumpy appearance. Found worldwide, the family includes about 300 species contained in three genera.
Trogids range in length from 2.5 to 20.0 mm. Their shape is oblong to oval, with a generally flat abdomen. Their color ranges from brown to gray or black, often obscured with a dirt encrusting. Otherwise they resemble scarab beetle
Scarab beetle
Scarab beetle may refer to these species: * A beetle of the family Scarabaeidae* Scarabaeus sacer, the dung beetle species worshipped by the ancient Egyptians as an embodiment of the god Khepri...
s, with heavy limbs and spurs.
They are scavengers, being among the last to visit and feed on the dried-out remains of dead animals; both adults and larvae will eat feathers, fur, and skin. They may also be found in bird and mammal nests. Details of species' life histories is usually poorly known, since many are specialized to particular types of nests. Between their covering of dirt and a habit of becoming motionless when disturbed, they are often overlooked, both by predators and by collectors.
The taxonomic position of these beetles is somewhat unsettled, with many authorities placing them as a subfamily Troginae of the Scarabaeidae
Scarabaeidae
The family Scarabaeidae as currently defined consists of over 30,000 species of beetles worldwide. The species in this large family are often called scarabs or scarab beetles. The classification of this family is fairly unstable, with numerous competing theories, and new proposals appearing quite...
. The common name "skin beetle" is sometimes used, but that name is usually used of the Dermestidae.
Origins
Trogidae, otherwise known as hide beetles, are found worldwide. The family of Trogidae has approximately three hundred species in three different genera that are worldwide in distribution: TroxTrox
Trox is a genus of beetles of the Family Trogidae.Its species occur almost worldwide.-Species:Trox is a genus of beetles of the Family Trogidae.Its species occur almost worldwide.-Species:...
, Omorgus
Omorgus
Omorgus is a genus of beetles of the Family Trogidae.Its species occur mostly in Africa and South America, although some species are found north up to Canada, others in Australia or Asia....
, and Polynoncus
Polynoncus
Polynoncus is a genus of beetles of the Family Trogidae.-Species:Polynoncus is a genus of beetles of the Family Trogidae.-Species:Polynoncus is a genus of beetles of the Family Trogidae.-Species:...
. These three genera are declared as the only known genera in the ‘New World
New World
The New World is one of the names used for the Western Hemisphere, specifically America and sometimes Oceania . The term originated in the late 15th century, when America had been recently discovered by European explorers, expanding the geographical horizon of the people of the European middle...
’; with Trox described by Fabricius
Johan Christian Fabricius
Johan Christian Fabricius was a Danish zoologist, specialising in "Insecta", which at that time included all arthropods: insects, arachnids, crustaceans and others...
in 1775, Omorgus described by Erichson
Wilhelm Ferdinand Erichson
Wilhelm Ferdinand Erichson was a German entomologist.He was the author of many articles about insects mainly in Archiv für Naturgeschichte.-Works:*Genera Dytiscorum. Berlin...
in 1847, and Polynoncus described by Burmeister
Hermann Burmeister
Karl Hermann Konrad Burmeister was a German zoologist, entomologist, and herpetologist.Burmeister was born in Stralsund and became a professor of Zoology at the Martin Luther University of Halle-Wittenberg from 1837 to 1861...
in 1876.
The family of Trogidae favors dry environments over moist environments and therefore is often found within temperate and plains areas. Each particular genus is found in different regions of the world, such as the Trox genus is found in the Holarctic
Holarctic
The Holarctic ecozone refers to the habitats found throughout the northern continents of the world as a whole. This region is divided into the Palearctic, consisting of Northern Africa and all of Eurasia, with the exception of Southeast Asia and the Indian subcontinent, and the Nearctic,...
/Ethiopian area; the Omorgus genus is typically more within the southern continents; and the Polynoncus genus is found in South America.
Specifically the beetles are found within the pellet
Pellet (ornithology)
A pellet, in ornithology, is the mass of undigested parts of a bird's food that some bird species occasionally regurgitate. The contents of a bird's pellet depend on its diet, but can include the exoskeletons of insects, indigestible plant matter, bones, fur, feathers, bills, claws, and teeth...
of any variety of animal, surrounding carrion or other decaying dry matter, and around birds’ and mammals’ nests and feathers as well as aging bones.
The origins and classifications for the family of Trogidae are very controversial but recent NA literature depicts Trogidae as its own family rather than a subfamily of Scarabaeidae
Scarabaeidae
The family Scarabaeidae as currently defined consists of over 30,000 species of beetles worldwide. The species in this large family are often called scarabs or scarab beetles. The classification of this family is fairly unstable, with numerous competing theories, and new proposals appearing quite...
. One major reason for the dispute between classifications is the possible evolution of the ommatidium
Ommatidium
The compound eyes of insects, mantis shrimp and millipedes are composed of units called ommatidia . An ommatidium contains a cluster of photoreceptor cells surrounded by support cells and pigment cells. The outer part of the ommatidium is overlaid with a transparent cornea...
in the eyes. The different environments, predators, etc. probably led to the adaptation of ommatidium structures within this family. For example the more advanced and numerous the ommatidium the more present the larger the ability of the insect to escape and elude predators. Due to these similarities many call the family of Scarabaeidae a ‘superfamily’ to the family of Trogidae. It is believed that Trogidae hails from Australia.
Anatomy
Trogidae are characterized by their distinct dirt-encrusted, warty or bumpy appearance. They are usually brown to gray/black in color and are covered with short, fairly dense setae. Their body shape is oblong to oval with a flat abdomen and their length varies from 2 to 20 millimeters depending on species. The antenna of hide beetles are usually fairly short and clubbed. The hardened elytra of Trogidae, which are generally covered with small knobs giving the beetle their rough appearance, meet along the midline of the body and cover the entire abdomen and well-developed wings. Their head is bent down and covered by the pronotum. They also have heavy limbs and spurs resembling those of scarab beetles. Trogidae larvae are a creamy yellow/white in color, except at their caudal end which darkens as it accumulates with feces. They have a heavily sclerotizedSclerosis (medicine)
In medicine, sclerosis refers to the stiffening of a structure, usually caused by a replacement of the normal organ-specific tissue with connective tissue.Types include:...
cranium that is almost black in color. The abdominal segments have at least one or more transverse rows of setae.
Diet and Habitat
Predators are rare due to Trogidae's habit of being covered with dirt and debris, and that when they are disturbed they have a habit of being motionless or faking death to avoid detection or being eaten. Their most common predators are birds since they tend to invade nests among other things.The Trogidae family is predaceous in addition to being scavengers. The pupae, larval, and adult stages of life have all been documented to be cannibalistic. Because of this tendency, the adult beetles will postpone, or delay pupation in an effort to maintain their safety and find a safe pupation site. The adult hide beetle produces a pheromone
Pheromone
A pheromone is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting outside the body of the secreting individual to impact the behavior of the receiving individual...
in its feces that leads other adults and larvae to food sources. Studies have shown that this species is not cannibalistic due to over-crowding of populations, but just a food source preference.
Beetles of the Trogidae family have also been found to feed off of carcasses in the wild that have died and are decomposing. In one lab experiment done in 1998 by the Department of Zoology at the University of Melbourne, the hide beetle ate all tissues on a sheep carcass and left the bones.
When carcasses are not available their diet consists of eating the dry remains of dead mammals and birds in later to last stages of decomposition. Since they are usually the last at the scene, they can be found eating feathers, fur, skin, feces and anything else they can scavenge. Trogidae Omorgus candidus or any beetle in the family Trogidae is a scavenger type beetle that all have the same diet and predators for all in this family are the same.
Mating Habits and Life Cycle
During decomposition of a carcass, the beetles will leave their nest to feed on the carrionCarrion
Carrion refers to the carcass of a dead animal. Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters include vultures, hawks, eagles, hyenas, Virginia Opossum, Tasmanian Devils, coyotes, Komodo dragons, and burying beetles...
. As the last succession of insects to appear on the carcass, both larvae and adults can be found feeding on the dry remains. At the site of the carcass, an impregnated female will dig small, vertical columns underneath the carcass to lay her eggs allowing the larvae to locate food after hatching.
Upon maturation of the larvae (approximately 6–8 weeks), females and males will mate. The male will sense odor cues from the female about her fertility. The male will mount the female and begin copulation. Copulation will be ended by the male. Males will tend to mate 6-7 times in their life span. Competition between the males for the female occurs occasionally as well as intra-sexual copulation between males. Males and females are polygamous.
There is little known about the life cycle of the Trogidae specifically. Their life cycles are very similar to the other genera of Scarabaeoidea (i.e. Passalidae and Lucanidae). Trogidae are holometabolous and usually have 3-5 instar
Instar
An instar is a developmental stage of arthropods, such as insects, between each molt , until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or assume a new form. Differences between instars can often be seen in altered body proportions, colors, patterns, or...
s. After impregnation of the female by the male, the female will lay the eggs and the larvae will hatch after an unknown amount of time. The larvae will usually molt twice growing and maturing until pupation. After pupation, the exoskeleton
Exoskeleton
An exoskeleton is the external skeleton that supports and protects an animal's body, in contrast to the internal skeleton of, for example, a human. In popular usage, some of the larger kinds of exoskeletons are known as "shells". Examples of exoskeleton animals include insects such as grasshoppers...
is finally formed and further growth will cease.
Forensic Importance
Trogidae's use in forensic entomology is unknown at this time. They are scavengers and can be found in carcasses or bird or mammal nests. They typically arrive last in the order of succession and feed on dried feathers, fur, and skin, but could be the first in succession if a body was first burned and charred. After the burned skin is eaten away by the Trogids, the corpse (with now-exposed, "fresher" surfaces) allows for viable colonization by other forensically important insects that help determine accurate Post mortem intervalPost mortem interval
Post-mortem interval is the time that has elapsed since a person has died. If the time in question is not known, a number of medical/scientific techniques are used to determine it. This also can refer to the stage of decompostion the person is in....
estimates. The adults lay their eggs in vertical burrows in the soil beneath carcasses.
Various species of Trogidae have been used by museums to clean up skeletons by eating any remaining dried material left on the skeletons leaving them clean for display. This method of bone-stripping has been used by some museums for many years as it is the most effective method.
Current and Future Research
The Chinese have been especially interested in the taxonomyTaxonomy
Taxonomy is the science of identifying and naming species, and arranging them into a classification. The field of taxonomy, sometimes referred to as "biological taxonomy", revolves around the description and use of taxonomic units, known as taxa...
of Trogidae. There is a current study involving advancements at the Chinese Academy of Sciences
Chinese Academy of Sciences
The Chinese Academy of Sciences , formerly known as Academia Sinica, is the national academy for the natural sciences of the People's Republic of China. It is an institution of the State Council of China. It is headquartered in Beijing, with institutes all over the People's Republic of China...
in efforts to further our knowledge on the classification of this family of beetles.
African Trogidae are being studied through the University of Pretoria
University of Pretoria
The University of Pretoria is a multi campus public research university located in Pretoria, the administrative and de facto capital of South Africa...
on the forensic importance of these beetles, as well as other carrion-associated beetles. Their article discusses how the presence of beetles on carrion effects the infestation of other arthropods in Africa.
Omorgus
- Omorgus acinus Scholtz, 1980 (TanzaniaTanzaniaThe United Republic of Tanzania is a country in East Africa bordered by Kenya and Uganda to the north, Rwanda, Burundi, and the Democratic Republic of the Congo to the west, and Zambia, Malawi, and Mozambique to the south. The country's eastern borders lie on the Indian Ocean.Tanzania is a state...
) - Omorgus alternansOmorgus alternansOmorgus alternans is a beetle of the Family Trogidae. It occurs in Australia....
(MacLeay, 1827) (Australia) - Omorgus amitinus Kolbe, 1904 (KenyaKenyaKenya , officially known as the Republic of Kenya, is a country in East Africa that lies on the equator, with the Indian Ocean to its south-east...
) - Omorgus asperOmorgus asperOmorgus asper is a beetle of the Family Trogidae. It occurs in the southern USA and Mexico....
LeConteJohn Lawrence LeConteJohn Lawrence LeConte was the most important American entomologist of the 19th century, responsible for naming and describing approximately half of the insect taxa known in the United States during his lifetime, including some 5,000 species of beetles...
, 1854 (southern USA, Mexico) - Omorgus asperulatus Harold, 1872 (Southern Africa)
- Omorgus australasiae (Erichson, 1842) (Australia)
- Omorgus baccatus GerstaeckerCarl Eduard Adolph GerstaeckerCarl Eduard Adolph Gerstaecker was a German zoologist and entomologist. After studying medicine and natural sciences in Berlin he was, from 1857 to 1876, Curator of the Zoological Museum of Humboldt University...
, 1867 (Kenya, Tanzania) - Omorgus badeni (Harold, 1872) (Brazil, Colombia)
- Omorgus batesi (Harold, 1872) (Argentina, Brazil)
- Omorgus birmanicus Arrow, 1927 (Southeast Asia)
- Omorgus borgognoi Marchand, 1902 (Mauretania, Mali, Chad)
- Omorgus borrei (Harold, 1872) (Uruguay, Argentina)
- Omorgus brucki Harold, 1872 (Australia)
- Omorgus candezei Harold, 1872 (Argentina)
- Omorgus capillaceus Scholtz, 1990 (Colombia)
- Omorgus carinatusOmorgus carinatusOmorgus carinatus is a beetle of the Family Trogidae....
Loomis, 1922 (Southern USA to Mexico) - Omorgus ciliatus (BlanchardÉmile BlanchardCharles Émile Blanchard was a French zoologist and entomologist.Blanchard was born in Paris. His father was an artist and naturalist and Émile began natural history very early in life. When he was 14 years old, Jean Victoire Audouin , allowed him access to the laboratory of the Muséum national...
, 1846) - Omorgus consanguineus Peringuey, 1901 (DR CongoDemocratic Republic of the CongoThe Democratic Republic of the Congo is a state located in Central Africa. It is the second largest country in Africa by area and the eleventh largest in the world...
, ZimbabweZimbabweZimbabwe is a landlocked country located in the southern part of the African continent, between the Zambezi and Limpopo rivers. It is bordered by South Africa to the south, Botswana to the southwest, Zambia and a tip of Namibia to the northwest and Mozambique to the east. Zimbabwe has three...
, NamibiaNamibiaNamibia, officially the Republic of Namibia , is a country in southern Africa whose western border is the Atlantic Ocean. It shares land borders with Angola and Zambia to the north, Botswana to the east and South Africa to the south and east. It gained independence from South Africa on 21 March...
) - Omorgus costatusOmorgus costatusOmorgus costatus is a species of beetle of the Family Trogidae that occurs in Australia, Tasmania, the Solomon Islands, New Guinea, Vietnam, Java, India and China....
(WiedemannChristian Rudolph Wilhelm WiedemannChristian Rudolph Wilhelm Wiedemann , was a German physician, historian, naturalist and entomologist...
, 1823) (Australia to India and China) - Omorgus crotchi Harold, 1871 (Australia)
- Omorgus denticulatus (OlivierGuillaume-Antoine OlivierGuillaume-Antoine Olivier was a French entomologist.He was the author of Entomologie, ou histoire naturelle des Insectes and Le Voyage dans l'Empire Othoman, l'Égypte et la Perse . He was a close friend of Johan Christian Fabricius and a patron of Pierre André Latreille.-References:...
, 1789) (Africa) - Omorgus desertorum Harold, 1872 (Madagascar, Egypt, Arabia)
- Omorgus discedens Haaf, 1954 (Somalia, Tanzania)
- Omorgus elevatus Harold, 1872 (Angola, Namibia)
- Omorgus endroedyi Scholtz, 1979 (Namibia, Angola)
- Omorgus expansus Arrow, 1900 (Somalia)
- Omorgus eyrensis BlackburnThomas Blackburn (entomologist)Thomas Blackburn was an English-born Australian entomologist who specialized in the study of beetles.Born near Liverpool, England, Blackburn became interested in entomology in his youth...
, 1904 (Australia) - Omorgus foveolatus BohemanCarl Henrik BohemanCarl Henrik Boheman was a Swedish entomologist.Boheman studied at Lund University and trained as an officer, participating in the invasion of Norway in 1814...
, 1860 (Madagascar, Namibia) - Omorgus freyi Haaf, 1954 (Southern Africa)
- Omorgus fuliginosusOmorgus fuliginosusOmorgus fuliginosus is a beetle of the Family Trogidae. It occurs in Texas, Mexico, El Salvador and Costa Rica.-References:* Catalogue of Life: 2007 Annual Checklist -...
Robinson, 1941 (Costa Rica to Texas) - Omorgus funestus Lansberge, 1886 (Angola)
- Omorgus gemmatus (Olivier, 1789) (Africa, Arabia)
- Omorgus glaber Scholtz, 1980 (Tanzania) (= Afromorgus lindemannae)
- Omorgus granulatusOmorgus granulatusOmorgus granulatus is a beetle of the Family Trogidae. It occurs in India and Sri Lanka....
(HerbstJohann Friedrich Wilhelm HerbstJohann Friedrich Wilhelm Herbst was a German naturalist and entomologist from Petershagen, Minden-Ravensberg....
, 1783) (India, Sri Lanka) - Omorgus guttalis Haaf, 1954 (Africa)
- Omorgus inclusus Walker, 1858 (Sri Lanka to China)
- Omorgus indicus Harold, 1872 (India, Thailand, China)
- Omorgus indigenus Scholtz, 1990 (Galapagos:Española IslandEspañola IslandEspañola Island is part of the Galápagos Islands. The English named it Hood Island after Viscount Samuel Hood. It is located in the extreme southeast of the archipelago and is considered, along with Santa Fe, one of the oldest, at approximately four million years...
) - Omorgus inflatusOmorgus inflatusOmorgus inflatus is a beetle of the Family Trogidae....
Loomis, 1922 (Arizona, Texas, Mexico) - Omorgus insignicollis Blackburn, 1896 (Australia)
- Omorgus insignis Haaf, 1954 (Namibia, Angola)
- Omorgus italicus Reiche, 1853 (Italy, India, China)
- Omorgus litigiosusOmorgus litigiosusOmorgus litigiosus is a beetle of the Family Trogidae....
- Omorgus lobicollis Arrow, 1927 (southern BurmaTanintharyi DivisionTanintharyi Region , is an administrative region of Myanmar, covering the long narrow southern part of the country on the Kra Isthmus. It borders with the Andaman Sea to the west and with the Tenasserim Hills, beyond which lies Thailand, to the east. To the north is the Mon State. The capital of...
) - Omorgus loxusOmorgus loxusOmorgus loxus is a beetle of the Family Trogidae....
Vaurie, 1955 (Brazil to Mexico) - Omorgus lugubris Haaf, 1954 (Kenya, Tanzania)
- Omorgus melancholicus (FahraeusOlof Immanuel von FåhraeusOlof Immanuel Fåhraeus , was a Swedish politician and entomologist mainly interested in Coleoptera....
, 1857) (Madagascar, Africa) - Omorgus mentitor Blackburn, 1896 (Australia)
- Omorgus mictlensis Deloya, 1995 (Mexico)
- Omorgus mollis Arrow, 1927 (Indonesia, Malaysia)
- Omorgus monachusOmorgus monachusOmorgus monachus is a beetle of the Family Trogidae....
(Herbst, 1790) (Mexico, Southern USA) - Omorgus mutabilis Haaf, 1954 (Africa)
- Omorgus nanningensis Pittino, 2005 (China)
- Omorgus niloticus Harold, 1872 (Africa)
- Omorgus nocheles Scholtz, 1990 (Argentina)
- Omorgus nodicollis Macleay, 1888 (Western Australia)
- Omorgus nodosusOmorgus nodosusOmorgus nodosus is a beetle of the Family Trogidae....
(Robinson, 1940) (Texas) - Omorgus nomadicus Scholtz, 1980 (Saudi Arabia)
- Omorgus obesus Scholtz, 1980 (Africa)
- Omorgus omacanthus Harold, 1872 (India)
- Omorgus pauliani Haaf, 1954 (Laos, Vietnam)
- Omorgus persuberosus Vaurie, 1962 (South America)
- Omorgus peruanus Erichson, 1847 (South America) (=Polynoncus peruanus)
- Omorgus ponderosus Peringuey, 1901 (Africa)
- Omorgus principalis Haaf, 1954 (Africa)
- Omorgus procerus Harold, 1872 (Africa, Arabia)
- Omorgus punctatusOmorgus punctatusOmorgus punctatus is a beetle of the Family Trogidae....
(GermarErnst Friedrich GermarErnst Friedrich Germar was a German professor and director of the Mineralogical Museum at Halle. As well as being a mineralogist he was interested in entomology and particularly in the Coleoptera and Hemiptera. He monographed the heteropteran family Scutelleridae.Amongst Germar’s publications...
, 1824) (Mexico to Southern USA) - Omorgus quadridensOmorgus quadridensOmorgus quadridens is a beetle of the Family Trogidae....
- Omorgus radula (Erichson, 1843) (Africa)
- Omorgus rodriguezae Deloya, 2005 (Mexico)
- Omorgus rubricansOmorgus rubricansOmorgus rubricans is a beetle of the Family Trogidae....
(Robinson, 1946) (Texas, Mexico) - Omorgus rusticus Fahraeus, 1857 (Africa)
- Omorgus scabrosusOmorgus scabrosusOmorgus scabrosus is a beetle of the Family Trogidae....
(Palisot de BeauvoisPalisot de BeauvoisAmbroise Marie François Joseph Palisot, Baron de Beauvois 27 July 1752 Arras - 21 January 1820 Paris, was a French naturalist.Palisot collected insects in Oware, Benin, Saint Domingue, and the United States, during the period 1786 – 1797. Trained as a botanist, Palisot published a significant...
, 1818) (Canada to southern USA) - Omorgus scutellarisOmorgus scutellarisOmorgus scutellaris is a beetle of the Family Trogidae....
(SayThomas SayThomas Say was an American naturalist, entomologist, malacologist, herpetologist and carcinologist. A taxonomist, he is often considered to be the father of descriptive entomology in the United States. He described more than 1,000 new species of beetles and over 400 species of insects of other...
, 1823) (Southern USA to Mexico) - Omorgus senegalensis Scholtz, 1983 (Senegal)
- Omorgus spatulatus Vaurie, 1962 (Argentina)
- Omorgus squalidusOmorgus squalidusOmorgus squalidus is a beetle of the Family Trogidae....
(Africa, Madagascar, Saudi Arabia) - Omorgus squamosusOmorgus squamosusOmorgus squamosus is a beetle of the Family Trogidae....
- Omorgus stellatusOmorgus stellatusOmorgus stellatus is a beetle of the Family Trogidae....
- Omorgus subcarinatusOmorgus subcarinatusOmorgus subcarinatus is a beetle of the Family Trogidae....
(MacLeay, 1864) (Australia, New Guinea) - Omorgus suberosusOmorgus suberosusOmorgus suberosus is a beetle of the Family Trogidae....
(FabriciusJohan Christian FabriciusJohan Christian Fabricius was a Danish zoologist, specialising in "Insecta", which at that time included all arthropods: insects, arachnids, crustaceans and others...
, 1775) (Spain, southern USA to South America, Australia) - Omorgus tessellatusOmorgus tessellatusOmorgus tessellatus is a beetle of the Family Trogidae....
LeConte, 1854 (Mexico) - Omorgus testudo Arrow, 1927 (southern BurmaTanintharyi DivisionTanintharyi Region , is an administrative region of Myanmar, covering the long narrow southern part of the country on the Kra Isthmus. It borders with the Andaman Sea to the west and with the Tenasserim Hills, beyond which lies Thailand, to the east. To the north is the Mon State. The capital of...
) - Omorgus texanusOmorgus texanusOmorgus texanus is a beetle of the Family Trogidae....
LeConte, 1854 (Texas) - Omorgus tomentosusOmorgus tomentosusOmorgus tomentosus is a beetle of the Family Trogidae....
(Robinson, 1941) (Mexico) - Omorgus tuberosus KlugJohann Christoph Friedrich KlugJohann Christoph Friedrich Klug was a German entomologist born 5 May 1775 in Berlin and died 3 February 1856 in the same city.He described the butterflies and some other insects of Upper Egypt and Arabia in Christian Gottfried Ehrenberg and Wilhelm Friedrich Hemprich's Symbolæ Physicæ...
, 1855 (Africa) - Omorgus tytusOmorgus tytusOmorgus tytus is a beetle of the Family Trogidae....
(Robinson, 1941) (USA) - Omorgus umbonatusOmorgus umbonatusOmorgus umbonatus is a beetle of the Family Trogidae....
LeConte, 1854 (Texas) - Omorgus unguicularis Haaf, 1954 (Africa)
- Omorgus varicosus (Erichson, 1843) (Angola)
- Omorgus villosusOmorgus villosusOmorgus villosus is a beetle of the Family Trogidae....
- Omorgus wittei Haaf, 1955 (Africa)
- Omorgus zumpti Haaf, 1957 (Africa)
Polynoncus
- Polynoncus aeger (Guerin-MenevilleFélix Édouard Guérin-MénevilleFélix Édouard Guérin-Méneville was a French entomologist.Guérin-Méneville changed his surname from Guérin in 1836. He was the author of the illustrated work Iconographie du Règne Animal de G. Cuvier 1829–1844, a complement to the work of Georges Cuvier and Pierre André Latreille, which lacked...
, 1844) (South America) - Polynoncus aricensis (Gutierrez, 1950) (South America)
- Polynoncus bifurcatus (Vaurie, 1962) (South America)
- Polynoncus brasiliensis (Vaurie, 1962) (South America)
- Polynoncus brevicollis (EschscholtzJohann Friedrich von EschscholtzJohann Friedrich Eschscholtz was a Livonian physician, botanist, zoologist and entomologist.Eschscholtz was born in Dorpat , Governorate of Livonia in the Russian Empire...
, 1822) (South America) - Polynoncus bullatus (CurtisJohn Curtis (entomologist)John Curtis was an English entomologist and illustrator.-Biography:Curtis was born in Norwich and learned his engraving skills in the workshop of his father, Charles Morgan Curtis...
, 1845) (Chile, Argentina) - Polynoncus burmeisteri Pittino, 1987 (Argentina)
- Polynoncus chilensis (Harold, 1872) (Chile, Argentina)
- Polynoncus diffluens (Vaurie, 1962) (Chile)
- Polynoncus ecuadorensis Vaurie, 1962 (Ecuador)
- Polynoncus erugatus Scholtz, 1990 (Argentina)
- Polynoncus galapagoensis (Van Dyke, 1953) (Galapagos Islands)
- Polynoncus gemmifer (Blanchard, 1846) (South America)
- Polynoncus gemmingeri (Harold, 1872) (Panama to Argentina)
- Polynoncus gibberosus Scholtz, 1990 (Chile)
- Polynoncus gordoni (Steiner, 1981) (Peru)
- Polynoncus guttifer (Harold, 1868) (South America)
- Polynoncus haafi Vaurie, 1962 (Argentina)
- Polynoncus hemisphaericus (BurmeisterHermann BurmeisterKarl Hermann Konrad Burmeister was a German zoologist, entomologist, and herpetologist.Burmeister was born in Stralsund and became a professor of Zoology at the Martin Luther University of Halle-Wittenberg from 1837 to 1861...
, 1876) (Argentina, Chile) - Polynoncus juglans (Ratcliffe, 1978) (Brazil, Guyana)
- Polynoncus longitarsis (Harold, 1872) (Argentina, Chile)
- Polynoncus mirabilis Pittino, 1987 (Chile, Argentina)
- Polynoncus neuquen (Vaurie, 1962) (Chile, Argentina)
- Polynoncus parafurcatus (Pittino, 1987) (Argentina, Brazil)
- Polynoncus patagonicus (Blanchard, 1846) (Argentina)
- Polynoncus patriciae Pittino, 1987 (Argentina, Uruguay)
- Polynoncus pedestris (Harold, 1872) (Argentina)
- Polynoncus peruanus (Erichson, 1847) (South America)
- Polynoncus pilularius (Germar, 1824) (South America)
- Polynoncus sallei (Harold, 1872) (Madagascar?, Ecuador, Peru)
- Polynoncus seymourensis (Mutchler, 1925) (Galapagos Islands)
- Polynoncus tenebrosus (Harold, 1872) (Ecuador)
Trox
- Trox acanthinus Harold, 1872 (Mexico)
- Trox aculeatus Harold, 1872 (South Africa)
- Trox aequalisTrox aequalisTrox aequalis is a beetle of the Family Trogidae....
Say, 1831 (Canada to Mexico) - Trox affinisTrox affinisTrox affinis is a beetle of the Family Trogidae....
Robinson, 1940 (USA) - Trox alatus Macleay, 1888 (Australia)
- Trox alius Scholtz, 1986 (Western Australia)
- Trox amictus Haaf, 1954 (Australia)
- Trox antiquus Wickham, 1909 (fossil:oligoceneOligoceneThe Oligocene is a geologic epoch of the Paleogene Period and extends from about 34 million to 23 million years before the present . As with other older geologic periods, the rock beds that define the period are well identified but the exact dates of the start and end of the period are slightly...
, FlorissantFlorissant Fossil Beds National MonumentFlorissant Fossil Beds National Monument is a United States National Monument in Teller County, Colorado, that is noted for its fossils. It is located in a mountain valley just west of Pikes Peak and holds spectacular remnants of prehistoric life...
, USA) - Trox aphanocephalus Scholtz, 1986 (Australia)
- Trox arcuatus Haaf, 1953 (South Africa)
- Trox atroxTrox atroxTrox atrox is a beetle of the Family Trogidae....
LeConte, 1854 (USA) - Trox augustae Blackburn, 1892 (Australia)
- Trox boucomonti Paulian, 1933 (China, Vietnam)
- Trox braacki Scholtz, 1980 (South Africa)
- Trox brahminus Pittino, 1985 (India to Vietnam)
- Trox brincki Haaf, 1958 (Lesotho)
- Trox cadaverinus Illiger, 1801 (Europe to China)
- Trox caffer Harold, 1872 (South Africa)
- Trox cambeforti Pittino, 1985 (China)
- Trox cambodjanus Pittino, 1985 (Cambodia, Laos)
- Trox candidus Harold, 1872 (Australia)
- Trox capensis Scholtz, 1979 (South Africa)
- Trox capillarisTrox capillarisTrox capillaris is a beetle of the Family Trogidae....
Say, 1823 (Canada to southern USA) - Trox carinicollis Scholtz, 1986 (Western Australia)
- Trox ciliatus Blanchard, 1846 (Argentina, Bolivia)
- Trox clathratus (ReicheLouis Jérôme ReicheLouis Jérôme Reiche , was a French merchant, manufacturer and entomologist.Reiche travelled widely in Europe making a large insect collection principally of beetles...
, 1861) (CorsicaCorsicaCorsica is an island in the Mediterranean Sea. It is located west of Italy, southeast of the French mainland, and north of the island of Sardinia....
) - Trox conjunctus Petrovitz, 1975 (China)
- Trox consimilis Haaf, 1953 (Southern Africa)
- Trox contractusTrox contractusTrox contractus is a beetle of the Family Trogidae....
Robinson, 1940 (Texas) - Trox coracinus GmelinJohann Friedrich GmelinJohann Friedrich Gmelin was a German naturalist, botanist, entomologist, herpetologist and malacologist.- Education :Johann Friedrich Gmelin was born as the eldest son of Philipp Friedrich Gmelin in 1748 in Tübingen...
, 1788 (unknown distribution) - Trox cotodognanensis Compte, 1986 (Spain)
- Trox cribrum GenéGiuseppe GenéCarlo Giuseppe Géné was an Italian naturalist and author.Géné was born at Turbigo in Lombardy and studied at the University of Pavia. He published a number of papers on natural history, particularly entomology...
, 1836 (France, SardiniaSardiniaSardinia is the second-largest island in the Mediterranean Sea . It is an autonomous region of Italy, and the nearest land masses are the French island of Corsica, the Italian Peninsula, Sicily, Tunisia and the Spanish Balearic Islands.The name Sardinia is from the pre-Roman noun *sard[],...
) - Trox cricetulus Ádám, 1994 (CroatiaCroatiaCroatia , officially the Republic of Croatia , is a unitary democratic parliamentary republic in Europe at the crossroads of the Mitteleuropa, the Balkans, and the Mediterranean. Its capital and largest city is Zagreb. The country is divided into 20 counties and the city of Zagreb. Croatia covers ...
, Spain) - Trox curvipes Harold, 1872 (Australia)
- Trox cyrtus Haaf, 1953 (South Africa)
- Trox demarzi Haaf, 1958 (Australia)
- Trox dhaulagiri Paulus, 1972 (Nepal)
- Trox dilaticollis Macleay, 1888 (Australia)
- Trox dohrni Harold, 1871 (Western Australia)
- Trox doiinthanonensis Masumoto, 1996 (Thailand)
- Trox elderi Blackburn, 1892 (South Australia)
- Trox elongatus Haaf, 1954 (Northern Australia)
- Trox erinaceus LeConte, 1854 (South Carolina)
- Trox euclensis Blackburn, 1892 (Australia)
- Trox eversmanni Krynicky, 1832 (Central Europe to Siberia)
- Trox fabricii Reiche, 1853 (Spain, Sicily to Northern Africa)
- Trox fascicularis Wiedemann, 1821 (Southern Africa)
- Trox fascifer LeConte, 1854 (California)
- Trox floridanus Howden & Vaurie, 1957 (Florida)
- Trox formosanus Nomura, 1973 (Taiwan)
- Trox foveicollisTrox foveicollisTrox foveicollis is a beetle of the Family Trogidae....
Harold, 1857 (USA) - Trox fronteraTrox fronteraTrox frontera is a beetle of the Family Trogidae....
Vaurie, 1955 (Texas) - Trox gansuensis Ren, 2003 (China)
- Trox gemmulatusTrox gemmulatusTrox gemmulatus is a beetle of the Family Trogidae....
HornGeorge Henry HornGeorge Henry Horn was a U.S. entomologist who specialized in the study of beetles.Born in Philadelphia, Horn attended the University of Pennsylvania, from which he graduated with a degree in medicine in 1861...
, 1874 (California) - Trox gigas Harold, 1872 (Australia)
- Trox gonoderus Fairmaire, 1901 (Madagascar)
- Trox granuliceps Haaf, 1954 (Australia)
- Trox granulipennis Fairmaire, 1852 (Northern Africa to Spain and Middle East)
- Trox gunki Scholtz, 1980 (South Africa)
- Trox hamatusTrox hamatusTrox hamatus is a beetle of the Family Trogidae....
Robinson, 1940 (USA) - Trox hispidus (PontoppidanErik PontoppidanErik Pontoppidan was a Danish author, bishop, historian and antiquary, born in Aarhus August 24, 1698; died in Copenhagen December 20, 1764. He was educated in Fredericia , after which he was a private tutor in Norway, and then studied in Holland, and in London and Oxford, England...
, 1763) (Europe) - Trox horridus Fabricius, 1775 (South Africa)
- Trox howdenorum Scholtz, 1986 (Western Australia)
- Trox howelli Howden & Vaurie, 1957 (Florida, Texas)
- Trox ineptus Balthasar, 1931 (TransbaikalTransbaikalTransbaikal, Trans-Baikal, Transbaikalia , or Dauria is a mountainous region to the east of or "beyond" Lake Baikal in Russia. The alternative name, Dauria, is derived from the ethnonym of the Daur people. It stretches for almost 1000 km from north to south from the Patomskoye Plateau and North...
) - Trox insularis ChevrolatLouis Alexandre Auguste ChevrolatLouis Alexandre Auguste Chevrolat was a French entomologist, born 29 March 1799 at Paris and died16 December 1884 in the same city.In government service in Paris, this amateur entomologist studied, mainly, Coleoptera and birds...
, 1864 (Southern USA, Cuba) - Trox kerleyi Masumoto, 1996 (Thailand)
- Trox kiuchii Masumoto, 1996 (Thailand)
- Trox klapperichi Pittino, 1983 (Turkey to Saudi Arabia, Middle East)
- Trox kyotensis Ochi & Kawahara, 2000 (Japan)
- Trox lama Pittino, 1985 (Tibet)
- Trox laticollis LeConte, 1854 (New York)
- Trox leonardii Pittino, 1983 (Spain to North Africa, Israel)
- Trox levis Haaf, 1953 (South Africa)
- Trox litoralis Pittino, 1991 (South Europe: Italy to Greece)
- Trox luridus Fabricius, 1781 (Southern Africa)
- Trox lutosus MarshamThomas MarshamThomas Marsham was an English entomologist, specializing on beetles.- Biography :He married a Miss Symes of Ufford, Northants, and had two daughters...
, 1802 (Great Britain) - Trox mandli Balthasar, 1931 (Transbaikal)
- Trox mariae Scholtz, 1986 (Western Australia)
- Trox mariettae Scholtz, 1986 (North Australia)
- Trox marshalli Haaf, 1957 (Australia)
- Trox martini (ReitterEdmund ReitterEdmund Reitter was a Austrian entomologist , writer and a collector.Edmund Reitter was above all well-known as an expert on the beetles of the Palaearctic....
, 1892) (North Africa) - Trox matsudai Ochi & Hori, 1999 (Japan)
- Trox maurus Herbst, 1790 (unknown distribution)
- Trox montanus Kolbe, 1891 (Africa)
- Trox monteithi Scholtz, 1986 (Australia)
- Trox morticinii PallasPeter Simon PallasPeter Simon Pallas was a German zoologist and botanist who worked in Russia.- Life and work :Pallas was born in Berlin, the son of Professor of Surgery Simon Pallas. He studied with private tutors and took an interest in natural history, later attending the University of Halle and the University...
, 1781 (Central Asia) - Trox mutsuensis Nomura, 1937 (Japan)
- Trox nama Kolbe, 1908 (Southern Africa)
- Trox nanniscus Peringuey, 1901 (South Africa)
- Trox nasutus Harold, 1872 (South Africa)
- Trox natalensis Haaf, 1954 (South Africa)
- Trox necopinus Scholtz, 1986 (Zambia)
- Trox niger Rossi, 1792
- Trox nigrociliatus Kolbe, 1904 (Ethiopia)
- Trox nigroscobinus Scholtz, 1986 (Western Australia)
- Trox niponensis Lewis, 1895 (Japan)
- Trox nodulosus Harold, 1872 (Sardinia, Corsica)
- Trox nohirai Nakane, 1954 (Japan)
- Trox novaecaledoniae Balthasar, 1966 (New CaledoniaNew CaledoniaNew Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...
) - Trox opacotuberculatus Motschulsky, 1860 (Japan, Taiwan)
- Trox oustaleti ScudderSamuel Hubbard ScudderSamuel Hubbard Scudder was an American entomologist and palaeontologist.Born in Boston, Massachusetts, Scudder may be most widely known for his essay on the importance of first-hand, careful observation in the natural sciences...
, 1879 (fossil: eocene; Nine-mile Creek, British ColumbiaBritish ColumbiaBritish Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...
) - Trox ovalis Haaf, 1957 (North Australia)
- Trox pampeanus Burmeister, 1876 (Argentina)
- Trox parvicollis Scholtz, 1986 (North Australia)
- Trox pastillarius Blanchard, 1846 (South America)
- Trox pellosomus Scholtz, 1986 (Australia)
- Trox penicillatus Fahraeus, 1857 (South Africa)
- Trox perhispidus Blackburn, 1904 (Australia)
- Trox perlatus GeoffroyÉtienne Louis GeoffroyÉtienne Louis Geoffroy was a French entomologist and pharmacist. He was born in Paris and died in Soissons. He followed the binomial nomenclature of Carl von Linné and devoted himself mainly to beetles....
, 1762 (Great Britain to Spain and Italy) - Trox perrieri Fairmaire, 1899 (Madagascar)
- Trox perrisii Fairmaire, 1868 (Europe, North Africa)
- Trox placosalinus Ren, 2003 (China)
- Trox planicollis Haaf, 1953 (Southern Africa)
- Trox plicatusTrox plicatusTrox plicatus is a beetle of the Family Trogidae....
Robinson, 1940 (Southern USA) - Trox poringensis Ochi, Kon & Kawahara, 2005 (Borneo, Java)
- Trox puncticollis Haaf, 1953 (Saudi Arabia)
- Trox pusillus Peringuey, 1908 (Africa)
- Trox quadridens Blackburn, 1892 (Australia)
- Trox quadrimaculatus Ballion, 1870 (TurkestanTurkestanTurkestan, spelled also as Turkistan, literally means "Land of the Turks".The term Turkestan is of Persian origin and has never been in use to denote a single nation. It was first used by Persian geographers to describe the place of Turkish peoples...
) - Trox quadrinodosus Haaf, 1954 (Australia)
- Trox regalis Haaf, 1954 (Australia)
- Trox rhyparoidesTrox rhyparoidesTrox rhyparoides is a beetle of the Family Trogidae. It is endemic to Africa....
(Harold, 1872) (Africa) - Trox rimulosus Haaf, 1957 (India)
- Trox robinsoniTrox robinsoniTrox robinsoni is a beetle of the Family Trogidae....
Vaurie, 1955 (Canada to Texas) - Trox rotundulus Haaf, 1957 (Australia)
- Trox rudebeckiTrox rudebeckiTrox rudebecki is a beetle of the Family Trogidae....
Haaf, 1958 (South Africa) - Trox sabulosus (LinnaeusCarolus LinnaeusCarl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...
, 1758) (Great Britain to Siberia) - Trox salebrosus Macleay, 1872 (Australia)
- Trox scaberTrox scaberTrox scaber is a beetle of the Family Trogidae....
(Linnaeus, 1767) (Holarctic, North Africa, South America, Australia) - Trox semicostatus Macleay, 1872 (Australia)
- Trox setifer WaterhouseFrederick George WaterhouseFrederick George Waterhouse was an English naturalist, zoologist and entomologist who made significant contributions to the study of the natural history of Australia....
, 1875 (Japan) - Trox setosipennis Blackburn, 1904 (Australia)
- Trox sonoraeTrox sonoraeTrox sonorae is a beetle of the Family Trogidae....
LeConte, 1854 (Canada to New Mexico) - Trox sordidatus Balthasar, 1936 (Southeastern Europe)
- Trox sordidusTrox sordidusTrox sordidus is a beetle of the Family Trogidae....
LeConte, 1854 (Canada to Texas) - Trox spinulosusTrox spinulosusTrox spinulosus is a beetle of the Family Trogidae....
Robinson, 1940 (USA) - Trox squamiger Roth, 1851 (Africa, Arabia)
- Trox squamosus Macleay, 1872 (Australia, New Guinea)
- Trox stellatus Harold, 1872 (Western Australia)
- Trox strandi Balthasar, 1936 (Algeria)
- Trox striatusTrox striatusTrox striatus is a beetle of the Family Trogidae....
MelsheimerFrederick Ernst MelsheimerFrederick Ernst Melsheimer, M.D. was an American entomologist noted for his work on Coleoptera. He was President of the American Entomological Society in 1853. Frederick Ernest Melsheimer's most important work was Catalogue of the described Coleoptera of the United States...
, 1846 (USA) - Trox strigosus Haaf, 1953 (South Africa)
- Trox strzeleckensis Blackburn, 1895 (Australia)
- Trox sugayai Masumoto & Kiuchi, 1995 (Japan)
- Trox sulcatusTrox sulcatusTrox sulcatus is a beetle of the Family Trogidae....
ThunbergCarl Peter ThunbergCarl Peter Thunberg aka Carl Pehr Thunberg aka Carl Per Thunberg was a Swedish naturalist and an apostle of Carl Linnaeus. He has been called "the father of South African botany" and the "Japanese Linnaeus"....
, 1787 (Southern Africa) - Trox taiwanus Masumoto, Ochi & Li, 2005 (Taiwan)
- Trox talpa Fahraeus, 1857 (South Africa)
- Trox tasmanicus Blackburn, 1904 (Tasmania)
- Trox tatei Blackburn, 1892 (Australia)
- Trox terrestrisTrox terrestrisTrox terrestris is a beetle of the Family Trogidae....
Say, 1825 (USA) - Trox tibialis Masumoto, Ochi & Li, 2005 (Taiwan)
- Trox torpidus Harold, 1872 (Central America)
- Trox transversus Reiche, 1856 (Greece, Syria, Turkey)
- Trox trilobus Haaf, 1954 (Australia, New Guinea)
- Trox tuberculatusTrox tuberculatusTrox tuberculatus is a beetle of the Family Trogidae....
(De GeerCharles De GeerBaron Charles de Geer was a Swedish industrialist and entomologist.- Life :...
, 1774) (USA) - Trox uenoi Nomura, 1961 (Japan)
- Trox unistriatusTrox unistriatusTrox unistriatus is a beetle of the Family Trogidae....
Palisot de Beauvois, 1818 (Canada to Texas) - Trox variolatusTrox variolatusTrox variolatus is a beetle of the Family Trogidae....
Melsheimer, 1846 (Canada to Mexico) - Trox villosus Haaf, 1954 (Australia)
- Trox yamayai Nakane, 1983 (Japan)
- Trox yangi Masumoto, Ochi & Li, 2005 (Taiwan)
- Trox zoufali Balthasar, 1931 (Taiwan)
External links
- World Trogidae Directory Naked list and picture by coleopterist Shinya Kawai
- Guide to New World Scarab Beetles - Trogidae from University of Nebraska State MuseumUniversity of Nebraska State MuseumThe University of Nebraska State Museum, also known as Elephant Hall, is a natural history museum featuring Nebraska biodiversity, paleontology, and cultural diversity. It was founded in 1871...
- Division of EntomologyEntomologyEntomology is the scientific study of insects, a branch of arthropodology... - http://bugguide.net/node/view/5824
- http://www.ento.csiro.au/education/insects/coleoptera_families/trogidae.html
- Key to the British species of family Trogidae