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Cnidocyte

Cnidocyte

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A cnidocyte, cnidoblast, or nematocyte is a type of venom
Venom
Venom is the general term referring to any variety of toxins used by certain types of animals that inject it into their victims by the means of a bite or a sting...

ous cell
Cell (biology)
The cell is the basic structural and functional unit of all known living organisms. It is the smallest unit of life that is classified as a living thing, and is often called the building block of life. The Alberts text discusses how the "cellular building blocks" move to shape developing embryos....

 unique to the phylum
Phylum
In biology, a phylum The term was coined by Georges Cuvier from Greek φῦλον phylon, "race, stock," related to φυλή phyle, "tribe, clan." is a taxonomic rank below kingdom and above class. "Phylum" is equivalent to the botanical term division....

 Cnidaria
Cnidaria
Cnidaria is a phylum containing over 9,000 species of animals found exclusively in aquatic and mostly marine environments. Their distinguishing feature is cnidocytes, specialized cells that they use mainly for capturing prey. Their bodies consist of mesoglea, a non-living jelly-like substance,...

 (coral
Coral
Corals are marine animals in class Anthozoa of phylum Cnidaria typically living in compact colonies of many identical individual "polyps". The group includes the important reef builders that inhabit tropical oceans and secrete calcium carbonate to form a hard skeleton.A coral "head" is a colony of...

s, sea anemone
Sea anemone
Sea anemones are a group of water-dwelling, predatory animals of the order Actiniaria; they are named after the anemone, a terrestrial flower. Sea anemones are classified in the phylum Cnidaria, class Anthozoa, subclass Zoantharia. Anthozoa often have large polyps that allow for digestion of larger...

s, hydra
Hydra (genus)
Hydra is a genus of simple fresh-water animal possessing radial symmetry. Hydras are predatory animals belonging to the phylum Cnidaria and the class Hydrozoa. They can be found in most unpolluted fresh-water ponds, lakes, and streams in the temperate and tropical regions and can be found by...

e, jellyfish
Jellyfish
Jellyfish are free-swimming members of the phylum Cnidaria. Medusa is another word for jellyfish, and refers to any free-swimming jellyfish stages in the phylum Cnidaria...

, etc.). The cnidocyte cell provides a means for them to catch prey and defend themselves from predators. Despite being morphologically simple, lacking a skeleton and usually being sessile
Sessility (zoology)
In zoology, sessility is a characteristic of animals which are not able to move about. They are usually permanently attached to a solid substrate of some kind, such as a part of a plant or dead tree trunk, a rock, or the hull of a ship in the case of barnacles. Corals lay down their own...

, cnidarians prey on fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

 and crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s. A cnidocyte fires a structure that contains the toxin
Toxin
A toxin is a poisonous substance produced within living cells or organisms; man-made substances created by artificial processes are thus excluded...

, from a characteristic sub-cellular organelle
Organelle
In cell biology, an organelle is a specialized subunit within a cell that has a specific function, and is usually separately enclosed within its own lipid bilayer....

 called a cnidocyst (also known as a cnida or nematocyst). This is responsible for the stings delivered by jellyfish.

Structure and action



Each cnidocyte cell contains an organelle called a cnidocyst (or nematocyst), which comprises a bulb-shape capsule containing a coiled hollow thread-like structure attached to it. The externally-oriented side of the cell also has a hair-like trigger called a cnidocil. When the trigger is activated, the shaft of the cnidocyst penetrates the target organism, and the hollow thread is everted into it. This discharge takes no more than a few microsecond
Microsecond
A microsecond is an SI unit of time equal to one millionth of a second. Its symbol is µs.A microsecond is equal to 1000 nanoseconds or 1/1000 millisecond...

s, and is able to reach acceleration
Acceleration
In physics, acceleration is the rate of change of velocity with time. In one dimension, acceleration is the rate at which something speeds up or slows down. However, since velocity is a vector, acceleration describes the rate of change of both the magnitude and the direction of velocity. ...

s of about 40,000 g
G-force
The g-force associated with an object is its acceleration relative to free-fall. This acceleration experienced by an object is due to the vector sum of non-gravitational forces acting on an object free to move. The accelerations that are not produced by gravity are termed proper accelerations, and...

. Recent research suggests the process to occur as fast as 700 nanoseconds, thus reaching an acceleration of up to 5,410,000 g. After penetration, the toxic content of the nematocyst is injected into the target organism. The rapid activity of the injected neurotoxins immediately paralyzes the mobile prey, thus allowing the sessile cnidarian to devour it.

Discharge mechanism


The nematocyst capsule stores a large concentration of calcium
Calcium
Calcium is the chemical element with the symbol Ca and atomic number 20. It has an atomic mass of 40.078 amu. Calcium is a soft gray alkaline earth metal, and is the fifth-most-abundant element by mass in the Earth's crust...

 ion
Ion
An ion is an atom or molecule in which the total number of electrons is not equal to the total number of protons, giving it a net positive or negative electrical charge. The name was given by physicist Michael Faraday for the substances that allow a current to pass between electrodes in a...

s, which are released from the capsule into the cytoplasm
Cytoplasm
The cytoplasm is a small gel-like substance residing between the cell membrane holding all the cell's internal sub-structures , except for the nucleus. All the contents of the cells of prokaryote organisms are contained within the cytoplasm...

 of the cnidocyte when the trigger is activated. This causes a large concentration gradient of calcium across the cnidocyte plasma membrane. The resulting osmotic pressure
Osmotic pressure
Osmotic pressure is the pressure which needs to be applied to a solution to prevent the inward flow of water across a semipermeable membrane....

 causes a rapid influx of water into the cell. This increase in water volume in the cytoplasm forces the coiled nematocyst to eject rapidly. The coiled nematocyst is a hollow tube that exists inside the cell in an "inside out" condition. The pressure of water flowing into the cnidocyte forces the water into the tubular nematocyst causing it to right itself as it comes rushing out of the cell with enough force to impale a prey organism.


Prey detection


A nematocyte is able under some conditions to fire independently, but this presents several problems for the cnidarian. First, it must avoid stinging itself. Second, it must replace cnidocytes after discharge, as they are "single use" cells, and this costs a lot of energy. In order to regulate discharge, cnidocytes are connected as "batteries", containing several types of nematocytes connected to supporting cells and neurons. The supporting cells contain chemosensor
Chemosensor
A chemoreceptor, also known as chemosensor, is a sensory receptor that transduces a chemical signal into an action potential. In more general terms, a chemosensor detects certain chemical stimuli in the environment.- Classes :...

s, which, together with the mechanoreceptor
Mechanoreceptor
A mechanoreceptor is a sensory receptor that responds to mechanical pressure or distortion. There are four main types in the glabrous skin of humans: Pacinian corpuscles, Meissner's corpuscles, Merkel's discs, and Ruffini corpuscles...

 on the cnidocyte (cnidocil), allow only the right combination of stimuli to cause discharge, such as prey swimming, and chemicals found in prey cuticle
Cuticle
A cuticle , or cuticula, is a term used for any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticles" are non-homologous; differing in their origin, structure, function, and chemical composition...

 or cuteous.

Types of nematocytes


Over 30 types of nematocytes are found in different cnidarians. They can be divided into the following groups:
  1. Penetrant: a harpoon-like structure used to penetrate, such as a nematocyst
  2. Glutinant: a sticky surface used to stick to prey
  3. Volvent: a lasso-like string that is fired at prey and wraps around a cellular projection on the prey, such as a spirocyst
  4. Ptychocyst: a special type of nematocyte found on burrowing (tube) anemones, which help create the tube in which the animal lives.


Depending on the species, one or several types can appear simultaneously on the organism.

Nematocyst toxicity


Nematocysts are very efficient weapons. A single nematocyst has been shown to suffice in paralyzing a small arthropod
Arthropod
An arthropod is an invertebrate animal having an exoskeleton , a segmented body, and jointed appendages. Arthropods are members of the phylum Arthropoda , and include the insects, arachnids, crustaceans, and others...

 (Drosophila
Drosophila
Drosophila is a genus of small flies, belonging to the family Drosophilidae, whose members are often called "fruit flies" or more appropriately pomace flies, vinegar flies, or wine flies, a reference to the characteristic of many species to linger around overripe or rotting fruit...

larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

). The most deadly cnidocytes (to humans, at least) are found on the body of a box jellyfish
Box jellyfish
Box jellyfish are cnidarian invertebrates distinguished by their cube-shaped medusae. Box jellyfish are known for the extremely potent venom produced by some species: Chironex fleckeri, Carukia barnesi and Malo kingi are among the most venomous creatures in the world...

. One member of this family, the sea wasp, Chironex fleckeri
Chironex fleckeri
Chironex fleckeri, commonly known as sea wasp, is a species of Box jellyfish found in coastal waters from northern Australia and New Guinea north to the Philippines and Vietnam. It has been described as "the most lethal jellyfish in the world", with at least 63 known deaths from 1884 to...

, is "claimed to be the most venomous marine animal known," according to the Australian Institute of Marine Science
Australian Institute of Marine Science
The Australian Institute of Marine Science is a state-of-the-art tropical marine research centre located primarily at Cape Ferguson, 50km by road east of Townsville in North Queensland, Australia. It was established in 1972, by the Commonwealth of Australia...

. It causes excruciating pain to humans, often followed by death, sometimes within two or three minutes. The chance of survival if stung while swimming alone is "virtually zero." Other cnidarians, such as the jellyfish Cyanea capillata
Lion's mane jellyfish
The lion's mane jellyfish is the largest known species of jellyfish. Its range is confined to cold, boreal waters of the Arctic, northern Atlantic, and northern Pacific Oceans, seldom found farther south than 42°N latitude. Similar jellyfish, which may be the same species, are known to inhabit...

(the "Lion's Mane
The Adventure of the Lion's Mane
"The Adventure of the Lion's Mane", one of the 56 Sherlock Holmes short stories written by British author Sir Arthur Conan Doyle, is one of 12 stories in the cycle collected as The Case Book of Sherlock Holmes. It is notable for being narrated by Holmes himself, instead of by Dr...

" made famous by Sherlock Holmes
Sherlock Holmes
Sherlock Holmes is a fictional detective created by Scottish author and physician Sir Arthur Conan Doyle. The fantastic London-based "consulting detective", Holmes is famous for his astute logical reasoning, his ability to take almost any disguise, and his use of forensic science skills to solve...

) or the hydrozoan Physalia physalis (Portuguese Man o' War
Portuguese Man o' War
The Portuguese Man o' War , also known as the Portuguese man-of-war, man-of-war, or bluebottle, is a jelly-like marine invertebrate of the family Physaliidae...

, "Bluebottle") can cause extremely painful and sometimes fatal stings. On the other side, aggregating
Shoaling and schooling
In biology, any group of fish that stay together for social reasons are said to be shoaling , and if, in addition, the group is swimming in the same direction in a coordinated manner, they are said to be schooling . In common usage, the terms are sometimes used rather loosely...

 sea anemones may have the lowest sting intensity, perhaps due to the inability of the nematocysts to penetrate the skin, providing only a feeling of that similar to touching sticky candies to human fingers. Besides feeding and defense, sea anemone and coral colonies use cnidocytes to sting one another in order to defend or win space.

Venom
Venom
Venom is the general term referring to any variety of toxins used by certain types of animals that inject it into their victims by the means of a bite or a sting...

 from animals such as cnidarians, scorpion
Scorpion
Scorpions are predatory arthropod animals of the order Scorpiones within the class Arachnida. They have eight legs and are easily recognized by the pair of grasping claws and the narrow, segmented tail, often carried in a characteristic forward curve over the back, ending with a venomous stinger...

s and spider
Spider
Spiders are air-breathing arthropods that have eight legs, and chelicerae with fangs that inject venom. They are the largest order of arachnids and rank seventh in total species diversity among all other groups of organisms...

s may be species-specific. A substance that is weakly toxic for humans or other mammals may be strongly toxic to the natural prey or predators of the venomous animal. Such specificity has been used to create new medicines and bioinsecticides.

Animals in the phylum Ctenophora ("sea-gooseberries" or "comb jellies") are transparent and jelly-like but have no nematocysts, and are harmless to humans.

Certain types of sea slugs, such as the nudibranch aeolids, are known to undergo kleptocnidae (in addition to kleptoplasty
Kleptoplasty
Kleptoplasty or kleptoplastidy is a symbiotic phenomenon whereby plastids from algae are sequestered by host organisms. The alga is eaten normally and partially digested, leaving the plastid intact. The plastids are maintained within the host, temporarily retaining functional photosynthesis for use...

), whereby the organisms store nematocysts of digested prey at the tips of their cerata.

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