Papuan Hornbill
Encyclopedia
The Blyth's Hornbill is also known as Papuan Hornbill. Its local name in Tok Pisin
Tok Pisin
Tok Pisin is a creole spoken throughout Papua New Guinea. It is an official language of Papua New Guinea and the most widely used language in that country...

 is kokomo. It is a large hornbill
Hornbill
Hornbills are a family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly-colored and sometimes has a casque on the upper mandible. Both the common English and the scientific name of the family...

 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 inhabiting the forest canopy in the Wallacea
Wallacea
Wallacea is a biogeographical designation for a group of Indonesian islands separated by deep water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as Lombok, Sumbawa, Flores, Sumba, Timor, Halmahera, Buru, Seram, and...

 and Melanesia
Melanesia
Melanesia is a subregion of Oceania extending from the western end of the Pacific Ocean to the Arafura Sea, and eastward to Fiji. The region comprises most of the islands immediately north and northeast of Australia...

.

Previously, this hornbill was placed in the genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 Aceros
Aceros
Aceros is a genus of large south-east Asian hornbills . This genus sometimes includes all species generally placed in the genus Rhyticeros. Alternatively, Aceros has been considered monotypic, only including the Rufous-necked Hornbill, in which case all of the remaining are moved to the Rhyticeros...

. It has in the past been often lumped
Lumpers and splitters
Lumping and splitting refers to a well-known problem in any discipline which has to place individual examples into rigorously defined categories. The lumper/splitter problem occurs when there is the need to create classifications and assign examples to them, for example schools of literature,...

 with the Plain-pouched Hornbill
Plain-pouched Hornbill
The Plain-pouched Hornbill is a species of hornbill in the Bucerotidae family. It is found in forests in southern Burma, adjacent parts of western Thailand and in northern Peninsular Malaysia....

 (R. subruficollis), and sometimes considered to include the Narcondam Hornbill
Narcondam Hornbill
The Narcondam Hornbill is a species of hornbill in the Bucerotidae family. It is endemic to the Indian island of Narcondam in the Andamans. Males and females have distinct plumages.-Description:...

 (R. narcondami) and the Wreathed Hornbill
Wreathed Hornbill
The Wreathed Hornbill , also known as the Bar-pouched Wreathed Hornbill, is a species of hornbill found in forests from far north-eastern India and Bhutan, east and south through mainland south-east Asia and the Greater Sundas, except Sulawesi. It is 75–100 cm long...

 (R. undulatus) as subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

.

Description

Up to 91 cm in length, the adult male has a mainly black plumage
Plumage
Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season. Within species there can also be a...

 with a golden or orange-buff coloured head, white throat and a white tail. Its irides
Iris (anatomy)
The iris is a thin, circular structure in the eye, responsible for controlling the diameter and size of the pupils and thus the amount of light reaching the retina. "Eye color" is the color of the iris, which can be green, blue, or brown. In some cases it can be hazel , grey, violet, or even pink...

 are reddish brown, surrounded by naked pale blue skin around the eye. The female is a smaller predominantly black bird with a white throat and tail. Both sexes have a very large, horn-coloured, bill and casque
Casque
Casque can refer to:* French word for helmet* An enlargement on the upper mandible of the bill of some species of birds, including many hornbills*Hornbill ivory: the casque of the Helmeted Hornbill, collected as a decorative material...

. Both sexes of young birds resemble the male. Adults have up to eight folds on the pale casque, depending on age, while young birds have none.

In flight the sound of its wings is loud and distinctive, a rushing noise that has been compared to the sound of steam escaping from a steam locomotive
Steam locomotive
A steam locomotive is a railway locomotive that produces its power through a steam engine. These locomotives are fueled by burning some combustible material, usually coal, wood or oil, to produce steam in a boiler, which drives the steam engine...

. As well as the noise produced by its wings, the Papuan Hornbill has a range of far-reaching, guttural grunting and laughing calls.

Subspecies

Various subspecies have been described across its range:
  • R. p. plicatus (Forster, 1781)South Moluccas
    Maluku Islands
    The Maluku Islands are an archipelago that is part of Indonesia, and part of the larger Maritime Southeast Asia region. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone...

  • R. p. ruficollis (Vieillot, 1816)North Moluccas
    North Maluku
    North Maluku is a province of northeastern Indonesia. It covers the northern part of the Maluku Islands, which are split between it and the province of Maluku. Maluku province used to cover the entire group...

     and West Papua, eastwards to the Southern Highlands and Simbu Province
    Simbu Province
    Simbu, also known as Chimbu, is a Highlands Region province in Papua New Guinea. The province has an area of 6,100 km² and a population of 259,703 . The capital of the province is Kundiawa...

    , Papua New Guinea
    Papua New Guinea
    Papua New Guinea , officially the Independent State of Papua New Guinea, is a country in Oceania, occupying the eastern half of the island of New Guinea and numerous offshore islands...

  • R. p. jungei Mayr, 1937 – Eastern New Guinea
    New Guinea
    New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

    , west as far as the Fly River
    Fly River
    The Fly at , is the second longest river, after the Sepik, in Papua New Guinea. The Fly is the largest river in Oceania, the largest in the world without a single dam in its catchment, and overall ranks as the twenty-fifth largest river in the world by volume of discharge...

     region
  • R. p. dampieri Mayr, 1934Bismarck Archipelago
    Bismarck Archipelago
    The Bismarck Archipelago is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea.-History:...

  • R. p. harterti Mayr, 1934Bougainville
    Bougainville Island
    Bougainville Island is the main island of the Autonomous Region of Bougainville of Papua New Guinea. This region is also known as Bougainville Province or the North Solomons. The population of the province is 175,160 , which includes the adjacent island of Buka and assorted outlying islands...

     and Buka Island
    Buka Island
    Buka Island is the second largest island in the Papua New Guinean province of Bougainville.- History :Buka was first occupied by humans in paleolithic times, some 30,000 years ago...

    s
  • R. p. mendanae Hartert, 1924Solomon Islands
    Solomon Islands
    Solomon Islands is a sovereign state in Oceania, east of Papua New Guinea, consisting of nearly one thousand islands. It covers a land mass of . The capital, Honiara, is located on the island of Guadalcanal...

     from Choiseul
    Choiseul Island
    Choiseul Island, native name Lauru, is the largest island of the Choiseul Province, Solomon Islands, at .-Description:This island is named after Étienne François, duc de Choiseul....

     to Guadalcanal
    Guadalcanal
    Guadalcanal is a tropical island in the South-Western Pacific. The largest island in the Solomons, it was discovered by the Spanish expedition of Alvaro de Mendaña in 1568...

     and Malaita
    Malaita
    Malaita is the largest island of the Malaita Province in the Solomon Islands. A tropical and mountainous island, Malaita's pristine river systems and tropical forests have not been exploited. Malaita is the most populous island of the Solomon Islands, with 140,000 people or more than a third of the...


Distribution and ecology

The Papuan Hornbill occurs throughout lowland forests, from sea level up to 1,200-1,500 m ASL
Above mean sea level
The term above mean sea level refers to the elevation or altitude of any object, relative to the average sea level datum. AMSL is used extensively in radio by engineers to determine the coverage area a station will be able to reach...

, in the Moluccas, New Guinea
New Guinea
New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

, the Bismarck Archipelago
Bismarck Archipelago
The Bismarck Archipelago is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea.-History:...

, and as far east as the Solomon Islands
Solomon Islands
Solomon Islands is a sovereign state in Oceania, east of Papua New Guinea, consisting of nearly one thousand islands. It covers a land mass of . The capital, Honiara, is located on the island of Guadalcanal...

. It is the only hornbill
Hornbill
Hornbills are a family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly-colored and sometimes has a casque on the upper mandible. Both the common English and the scientific name of the family...

 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 native to New Guinea, and one of the largest flying birds of the region.

Its diet consists mainly of fruits – especially figs
Ficus
Ficus is a genus of about 850 species of woody trees, shrubs, vines, epiphytes, and hemiepiphyte in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extending into the semi-warm temperate zone. The Common Fig Ficus is a genus of...

 (Ficus) –, occasionally supplemented with insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s and other small animals.

The Papuan Hornbill nests in a large rainforest
Rainforest
Rainforests are forests characterized by high rainfall, with definitions based on a minimum normal annual rainfall of 1750-2000 mm...

 tree hollow
Tree hollow
A tree hollow or tree hole is a semi-enclosed cavity which has naturally formed in the trunk or branch of a tree. These are predominantly found in old trees, whether living or not...

 from 18 m to at least 30 m above the ground. The female is restricted to the nest cavity throughout the incubation and nestling period, being largely sealed within with the entrance plastered up by a mixture of fruit pulp and rotten wood, leaving only a narrow aperture through which she is fed by the male. The clutch size is about two eggs.

Status and relationship with humans

Still widespread throughout its large habitat range, the Papuan Hornbill is evaluated as a Species of Least Concern
Least Concern
Least Concern is an IUCN category assigned to extant taxon or lower taxa which have been evaluated but do not qualify for any other category. As such they do not qualify as threatened, Near Threatened, or Conservation Dependent...

 on the IUCN Red List
IUCN Red List
The IUCN Red List of Threatened Species , founded in 1963, is the world's most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the world's main authority on the conservation status of species...

 of Threatened Species.

However, it is subject to hunting pressure by some tribal groups, with its feathers used in headdress
Headgear
Headgear, headwear or headdress is the name given to any element of clothing which is worn on one's head.Headgear serve a variety of purposes:...

es, its bill being valued as a personal adornment, and the lower mandible used as a spear
Spear
A spear is a pole weapon consisting of a shaft, usually of wood, with a pointed head.The head may be simply the sharpened end of the shaft itself, as is the case with bamboo spears, or it may be made of a more durable material fastened to the shaft, such as flint, obsidian, iron, steel or...

 point. As a consequence, it is becoming rarer in some areas of New Guinea. On the other hand, this species has withstood tens of millennia of human hunting pressure, so as long as sufficient habitat
Habitat
* Habitat , a place where a species lives and grows*Human habitat, a place where humans live, work or play** Space habitat, a space station intended as a permanent settlement...

is preserved, it is unlikely that hunting alone is a significant threat.

External links

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