Lettered olive
Encyclopedia
The lettered olive, Oliva sayana, is a species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of large predatory sea snail
Snail
Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

, a marine
Marine (ocean)
Marine is an umbrella term. As an adjective it is usually applicable to things relating to the sea or ocean, such as marine biology, marine ecology and marine geology...

 gastropod mollusc in the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Olividae, the olive shell
Olive Shell
Olive shells, olive snails, or olives, scientific name Olividae, are a family of medium to large predatory sea snails with smooth, shiny, elongated oval-shaped shells. The shells often show various muted but attractive colors, and may be patterned also...

s, olive snails, or olives.

Subspecies

Oliva sayana contains the following subspecies:
  • Oliva sayana sarasotensis Petuch & Sargent, 1986
  • Oliva sayana sayana Ravenel, 1834
  • Oliva sayana texana Petuch & Sargent, 1986

Distribution

The species' range is from North Carolina
North Carolina
North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

 to Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

 and the Gulf states of North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

, ingluding Louisiana
Louisiana
Louisiana is a state located in the southern region of the United States of America. Its capital is Baton Rouge and largest city is New Orleans. Louisiana is the only state in the U.S. with political subdivisions termed parishes, which are local governments equivalent to counties...

 and Texas
Texas
Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

; It also occurs in Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

, including Campeche State, Yucatán State
Yucatán
Yucatán officially Estado Libre y Soberano de Yucatán is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 106 municipalities and its capital city is Mérida....

 and Quintana Roo
Quintana Roo
Quintana Roo officially Estado Libre y Soberano de Quintana Roo is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 10 municipalities and its capital city is Chetumal....

, and may also occur in Brazil
Brazil
Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

.

Habitat

The lettered olive typically lives in near-shore waters, on shallow sand flats near inlets. The empty shell is occasionally, or sometimes commonly, washed up onto ocean beaches.

Shell description

The shell
Gastropod shell
The gastropod shell is a shell which is part of the body of a gastropod or snail, one kind of mollusc. The gastropod shell is an external skeleton or exoskeleton, which serves not only for muscle attachment, but also for protection from predators and from mechanical damage...

 of this species can be about 6 cm (2½ in) long (maximum reported size reaches 9.1 cm). It is a smooth, shiny, cylindrical-shaped shell with a short spire
Spire (mollusc)
A spire is a descriptive term for part of the coiled shell of mollusks. The word is a convenient aid in describing shells, but it does not refer to a very precise part of shell anatomy: the spire consists of all of the whorls except for the body whorl...

. The aperture
Aperture (mollusc)
The aperture is an opening in certain kinds of mollusc shells: it is the main opening of the shell, where part of the body of the animal emerges for locomotion, feeding, etc....

 is narrow and extending almost the length of shell, continuing around the bottom and ending in a notch on the other side. The suture is V-cut and deep. The lower part of the whorl
Whorl (mollusc)
A whorl is a single, complete 360° revolution or turn in the spiral growth of a mollusc shell. A spiral configuration of the shell is found in of numerous gastropods, but it is also found in shelled cephalopods including Nautilus, Spirula and the large extinct subclass of cephalopods known as the...

 is just above where the suture extends outward and then at a sharp shoulder drops into the suture.

The shell coloration can vary from cream to a greyish exterior with reddish-brown zigzag markings. The common name
Common name
A common name of a taxon or organism is a name in general use within a community; it is often contrasted with the scientific name for the same organism...

 of this species is derived from the darker surface markings that sometimes resemble letters.

Life habits

This olive, like all olives, is a carnivore
Carnivore
A carnivore meaning 'meat eater' is an organism that derives its energy and nutrient requirements from a diet consisting mainly or exclusively of animal tissue, whether through predation or scavenging...

: it captures bivalves and small crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s with its foot and takes them below the sand surface to digest.

Its presence is sometimes detected at very low tides by the trails it leaves when it crawls below the surface on semi-exposed sand flats.

Females lay floating, round egg capsules that are often found in beach drift. Young are free swimming.

Human use

Colonists and early Native Americans made jewelry from these shells.

The lettered olive is the state shell of South Carolina
South Carolina
South Carolina is a state in the Deep South of the United States that borders Georgia to the south, North Carolina to the north, and the Atlantic Ocean to the east. Originally part of the Province of Carolina, the Province of South Carolina was one of the 13 colonies that declared independence...

.
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