Hutchemys
Encyclopedia
Hutchemys is an extinct genus
of softshell turtles
from the late Cretaceous
(Maastrichtian
stage) to the late Paleocene
of New Mexico
, Montana
, Wyoming
and Utah
, USA. It was first named by Walter G. Joyce, Ariel Revan, Tyler R. Lyson and Igor G. Danilov in 2009
, and the type species
is Hutchemys rememdium. H. rememdium is known from the holotype
YPM PU 16795, which consists of a nearly complete postcrania
l skeleton
, and from the referred specimen YPM PU 16781, found in the Ekalaka Member of the Fort Union Formation
, Montana. Another referred specimen, YPM PU 14985, was found in the Cedar Point Quarry, Wyoming
. The second species, H. arctochelys, is known from the holotype
YPM PU 16319, a nearly complete carapace
, and from the paratypes YPM PU 16320, YPM PU 16321, YPM PU 16322, YPM PU 16238. All specimens of H. arctochelys were recovered from the same quarry of the Tongue River Member, Fort Union Formation
, near Burns Mine of Montana. A possible third species is represented by the unnamed specimen UCMP 130000 from the Paleocene Tullock Formation of Montana. Aspideretes? nassau (YPM PU 11566) from the Fort Union Formation
, Duffy’s Ranch of Sweet Grass County, Montana was also assigned to Hutchemys sp.
The generic name honors John Howard Hutchison, a turtle paleontologist. Hutchison himself named later in 2009 two new genera of extinct softshell turtles from New Mexico
, Montana
, Wyoming
and Utah
: Plastomenoides (two new species: P. lamberti (type) and P. tetanetron) and Derrisemys (new species: D. sterea (type) and new combination D. acupictus [originally Plastomenus]). In 2011, Joyce and Lyson noted that P. lamberti and Hutchemys rememdium are based on the same type specimen (YPM PU 16795) and therefore P. lamberti and Plastomenoides are objective junior synonyms of H. rememdium and Hutchemys. P. tetanetron Hutchison, 2009 is also cogeneric with H. rememdium. They also found that D. sterea is the closest relative of P. tetanetron and therefore both D. sterea and D. acupictus were reassigned to Hutchemys.
after Joyce, Revan, Lyson and Danilov, 2009:
Cladogram after Joyce and Lyson, 2011:
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of softshell turtles
Trionychidae
Trionychidae is a taxonomic family which comprises a number of turtle genera commonly known as softshells. They are also sometimes called pancake turtles. Softshells consist of some of the world's largest fresh water turtles, though many can adapt to living in highly brackish areas. Members of this...
from the late Cretaceous
Cretaceous
The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...
(Maastrichtian
Maastrichtian
The Maastrichtian is, in the ICS' geologic timescale, the latest age or upper stage of the Late Cretaceous epoch or Upper Cretaceous series, the Cretaceous period or system, and of the Mesozoic era or erathem. It spanned from 70.6 ± 0.6 Ma to 65.5 ± 0.3 Ma...
stage) to the late Paleocene
Paleocene
The Paleocene or Palaeocene, the "early recent", is a geologic epoch that lasted from about . It is the first epoch of the Palaeogene Period in the modern Cenozoic Era...
of New Mexico
New Mexico
New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...
, Montana
Montana
Montana is a state in the Western United States. The western third of Montana contains numerous mountain ranges. Smaller, "island ranges" are found in the central third of the state, for a total of 77 named ranges of the Rocky Mountains. This geographical fact is reflected in the state's name,...
, Wyoming
Wyoming
Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...
and Utah
Utah
Utah is a state in the Western United States. It was the 45th state to join the Union, on January 4, 1896. Approximately 80% of Utah's 2,763,885 people live along the Wasatch Front, centering on Salt Lake City. This leaves vast expanses of the state nearly uninhabited, making the population the...
, USA. It was first named by Walter G. Joyce, Ariel Revan, Tyler R. Lyson and Igor G. Danilov in 2009
2009 in paleontology
-Anomalocaridids:-Arachnids:-Insects:-Cephalopods:Three new species of extinct Octopoda discovered in 2009. The species - Keuppia hyperbolaris, Keuppia levante, and Styletoctopus annae - lived about 95 million years ago, and bear a strong resemblance to modern octopuses, suggesting that the...
, and the type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...
is Hutchemys rememdium. H. rememdium is known from the holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
YPM PU 16795, which consists of a nearly complete postcrania
Postcrania
Postcrania[p] in zoology and vertebrate paleontology refers to all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g...
l skeleton
Skeleton
The skeleton is the body part that forms the supporting structure of an organism. There are two different skeletal types: the exoskeleton, which is the stable outer shell of an organism, and the endoskeleton, which forms the support structure inside the body.In a figurative sense, skeleton can...
, and from the referred specimen YPM PU 16781, found in the Ekalaka Member of the Fort Union Formation
Fort Union Formation
The Fort Union Formation is a geologic unit containing sandstones, shales, and coal beds in Wyoming, Montana, and parts of adjacent states. In the Powder River Basin, it contains important economic deposits of coal, uranium, and coalbed methane....
, Montana. Another referred specimen, YPM PU 14985, was found in the Cedar Point Quarry, Wyoming
Wyoming
Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...
. The second species, H. arctochelys, is known from the holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
YPM PU 16319, a nearly complete carapace
Carapace
A carapace is a dorsal section of the exoskeleton or shell in a number of animal groups, including arthropods such as crustaceans and arachnids, as well as vertebrates such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron.-Crustaceans:In crustaceans, the...
, and from the paratypes YPM PU 16320, YPM PU 16321, YPM PU 16322, YPM PU 16238. All specimens of H. arctochelys were recovered from the same quarry of the Tongue River Member, Fort Union Formation
Fort Union Formation
The Fort Union Formation is a geologic unit containing sandstones, shales, and coal beds in Wyoming, Montana, and parts of adjacent states. In the Powder River Basin, it contains important economic deposits of coal, uranium, and coalbed methane....
, near Burns Mine of Montana. A possible third species is represented by the unnamed specimen UCMP 130000 from the Paleocene Tullock Formation of Montana. Aspideretes? nassau (YPM PU 11566) from the Fort Union Formation
Fort Union Formation
The Fort Union Formation is a geologic unit containing sandstones, shales, and coal beds in Wyoming, Montana, and parts of adjacent states. In the Powder River Basin, it contains important economic deposits of coal, uranium, and coalbed methane....
, Duffy’s Ranch of Sweet Grass County, Montana was also assigned to Hutchemys sp.
The generic name honors John Howard Hutchison, a turtle paleontologist. Hutchison himself named later in 2009 two new genera of extinct softshell turtles from New Mexico
New Mexico
New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...
, Montana
Montana
Montana is a state in the Western United States. The western third of Montana contains numerous mountain ranges. Smaller, "island ranges" are found in the central third of the state, for a total of 77 named ranges of the Rocky Mountains. This geographical fact is reflected in the state's name,...
, Wyoming
Wyoming
Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...
and Utah
Utah
Utah is a state in the Western United States. It was the 45th state to join the Union, on January 4, 1896. Approximately 80% of Utah's 2,763,885 people live along the Wasatch Front, centering on Salt Lake City. This leaves vast expanses of the state nearly uninhabited, making the population the...
: Plastomenoides (two new species: P. lamberti (type) and P. tetanetron) and Derrisemys (new species: D. sterea (type) and new combination D. acupictus [originally Plastomenus]). In 2011, Joyce and Lyson noted that P. lamberti and Hutchemys rememdium are based on the same type specimen (YPM PU 16795) and therefore P. lamberti and Plastomenoides are objective junior synonyms of H. rememdium and Hutchemys. P. tetanetron Hutchison, 2009 is also cogeneric with H. rememdium. They also found that D. sterea is the closest relative of P. tetanetron and therefore both D. sterea and D. acupictus were reassigned to Hutchemys.
Phylogeny
CladogramCladogram
A cladogram is a diagram used in cladistics which shows ancestral relations between organisms, to represent the evolutionary tree of life. Although traditionally such cladograms were generated largely on the basis of morphological characters, DNA and RNA sequencing data and computational...
after Joyce, Revan, Lyson and Danilov, 2009:
Cladogram after Joyce and Lyson, 2011: