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Gastrolith

 

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Gastrolith



 
 
Gastroliths ('stomach
Stomach

In most mammals, the stomach is a hollow muscular organ of the gastrointestinal tract involved in the second phase of digestion, following mastication....
 stones' or 'gizzard
Gizzard

The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including birds, reptiles, earthworms and some fish....
 stones') are rocks
Rock (geology)

In geology, rock is a naturally occurring solid aggregate of minerals and/or mineraloids.The Earth's outer solid layer, the lithosphere, is made of rock....
, which are or have been held inside the digestive tract
Gastrointestinal tract

The digestive tract is the system of Organ s within multicellular animals that takes in food, digestion it to extract energy and nutrients, and expels the remaining waste....
 of an animal. Among living vertebrates, gastroliths are common among herbivorous
Herbivore

Herbivory is a form of predation in which an organism, known as an herbivore, heterotrophs principally autotrophs such as plants, algae and photosynthesizing bacteria....
 birds, crocodiles, alligators, seals
Seals

Seals, or Seales, as a surname may refer to:*Brady Seals , American country music artist*Bruce Seals , American basketball player*Dan Seals , American musician...
 and sea lions
Sea Lion

For other uses of the term "sea lion", see Sea lion .Sea lions are any of seven species in six genera of modern pinnipeds including one extinct ....
. Domestic fowl
Fowl

Fowl is a term for birds; fowl belong to one of two order , namely the gamefowl or landfowl and the waterfowl . Studies of anatomical and molecular similarities suggest these two groups were close evolutionary relatives; together, they form the fowl clade which is scientifically known as Galloanserae ....
, for instance, require access to 'grit', for the purpose of food-grinding. Gastroliths are retained in the very muscular gizzard
Gizzard

The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including birds, reptiles, earthworms and some fish....
 and serve the masticatory function of teeth, in an animal without suitable grinding teeth.






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Gastroliths ('stomach
Stomach

In most mammals, the stomach is a hollow muscular organ of the gastrointestinal tract involved in the second phase of digestion, following mastication....
 stones' or 'gizzard
Gizzard

The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including birds, reptiles, earthworms and some fish....
 stones') are rocks
Rock (geology)

In geology, rock is a naturally occurring solid aggregate of minerals and/or mineraloids.The Earth's outer solid layer, the lithosphere, is made of rock....
, which are or have been held inside the digestive tract
Gastrointestinal tract

The digestive tract is the system of Organ s within multicellular animals that takes in food, digestion it to extract energy and nutrients, and expels the remaining waste....
 of an animal. Among living vertebrates, gastroliths are common among herbivorous
Herbivore

Herbivory is a form of predation in which an organism, known as an herbivore, heterotrophs principally autotrophs such as plants, algae and photosynthesizing bacteria....
 birds, crocodiles, alligators, seals
Seals

Seals, or Seales, as a surname may refer to:*Brady Seals , American country music artist*Bruce Seals , American basketball player*Dan Seals , American musician...
 and sea lions
Sea Lion

For other uses of the term "sea lion", see Sea lion .Sea lions are any of seven species in six genera of modern pinnipeds including one extinct ....
. Domestic fowl
Fowl

Fowl is a term for birds; fowl belong to one of two order , namely the gamefowl or landfowl and the waterfowl . Studies of anatomical and molecular similarities suggest these two groups were close evolutionary relatives; together, they form the fowl clade which is scientifically known as Galloanserae ....
, for instance, require access to 'grit', for the purpose of food-grinding. Gastroliths are retained in the very muscular gizzard
Gizzard

The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including birds, reptiles, earthworms and some fish....
 and serve the masticatory function of teeth, in an animal without suitable grinding teeth. The grain size
Grain size

*particle size provides general definitions of the particle size notion.*particle size provides specific details for particle size of soils, powders, gravel, etc....
 of the gastrolith depends upon the size of the animal and its special needs. Particles as small as sand
Sand

Sand is a naturally occurring granular material composed of finely divided rock and mineral particles.As the term is used by geologists, sand particles range in diameter from 0.0625 to 2 millimeters....
 and stones the size of cobble
Cobble

Cobble may refer to:* A particular size of rock, larger than gravel* Cobblestone, partially-rounded rocks used for road paving* Hammerstone, a prehistoric stone tool...
s or greater have been found. Some extinct animals, such as sauropod dinosaurs, appear to have used stones to grind tough plant matter. Gastroliths have only rarely been found in association with fossils of theropod dinosaurs and a trituration
Trituration

Trituration is the name of several different methods of processing materials.Trituration is also the name of the process for reducing the particle size of a substance by grinding, as by grinding of powders in a Mortar and pestle with a pestle....
 of their food with the stones is not plausible. Aquatic animals, such as plesiosaurs, may have used them as ballast, to help balance themselves or to decrease their buoyancy, as crocodiles do. More research is needed to understand the function of the stones in aquatic animals. While some fossil gastroliths are rounded and polished, many stones in living birds are not polished at all. Gastroliths associated with dinosaur
Dinosaur

Dinosaurs were the dominant vertebrate animals of Landform ecosystems for over 160 million years, from the late Triassic Period until the end of the Cretaceous Period , when most of them became extinct in the Cretaceous?Tertiary extinction event....
 fossils can be several kilograms in weight. Stones swallowed by ostriches can also reach a length of more than 10 cm.

Geologist
Geologist

For other uses, see Geologist .A geologist is a contributor to the science of geology, studying the physical structure and processes of the Earth and planets of the solar system ....
s usually require several pieces of evidence before they will accept that a rock was used by a dinosaur to aid its digestion
Digestion

Digestion is the mechanical and chemical breaking down of food into smaller components, to a form that can be Absorption, for instance, by a blood stream....
. First, the stone must be unlike the rock found in its geological vicinity. Secondly, it should be rounded and polished, because inside a dinosaur's gizzard any genuine gastrolith would have been acted upon by other stones and fibrous materials in a process similar to the action of a rock tumbler. Lastly, the stone must be found with the bone
Bone

Bones are rigid organ that form part of the endoskeleton of vertebrates. They function to move, support, and protect the various organs of the body, produce red blood cell and white blood cells and store minerals....
s of the dinosaur which ingested it. It is this last criterion that causes trouble in identification, as smooth stones found without context can (possibly erroneously in some cases) be dismissed as having been polished by water or wind. Whittle (1988,9) pioneered scanning electron microscope
Scanning electron microscope

The scanning electron microscope is a type of electron microscope that images the sample surface by scanning it with a high-energy beam of electrons in a raster scan pattern....
 analysis of wear patterns on gastroliths. Wings (2003) found that ostrich
Ostrich

The ostrich Struthio camelus is a large flightless bird native to Africa . It is the only living species of its family , Struthionidae, and its genus, Struthio....
 gastroliths would be deposited outside the skeleton if the carcass was deposited in an aquatic environment for as little as a few days following death. He concludes that this is likely to hold true for all birds (with the possible exception of moa
Moa

The moa were ten species of flightless birds endemic to New Zealand. The two largest species, Dinornis robustus and Dinornis novaezelandiae, reached about in height with neck outstretched, and weighed about ....
) due to their air-filled bones which would cause a carcass deposited in water to float for the time it needs to rot sufficiently to allow gastroliths to escape.

Gastroliths can be distinguished from stream- or beach-rounded rocks by several criteria: gastroliths are highly polished on the higher surfaces, with little or no polish in depressions or crevices, often strongly resembling the surface of worn animal teeth. Stream- or beach-worn rocks, particularly in a high-impact environment, show less polishing on higher surfaces, often with many small pits or cracks on these higher surfaces. Finally, highly polished gastroliths often show long microscopic hairline scratches, presumably caused by contact with a sharp corner of a freshly swallowed stone. Since most gastroliths were scattered when the animal died and many entered a stream or beach environment, some gastroliths show a mixture of these wear features. Others were undoubtedly swallowed by other dinosaurs and highly polished gastroliths may have been swallowed repeatedly.

The American Museum of Natural History
American Museum of Natural History

The American Museum of Natural History , located on the Upper West Side, Manhattan, New York, USA, is one of the largest and most celebrated museums in the world....
 Photograph # 311488 demonstrates an articulated skeleton of a Protiguanadon mongoliense, from the Ondai Sair Formation, Lower Cretaceous
Cretaceous

The Cretaceous , usually abbreviated K for its German translation Kreide, is a geologic period from circa to million years ago . In the geologic timescale, the Cretaceous follows on the Jurassic period and is followed by the Paleogene period....
 Period of Mongolia
Mongolia

Mongolia is a landlocked country in East Asia and Central Asia. It borders Russia to the north and People's Republic of China to the south, east and west....
, showing a collection of about 40 gastroliths inside the rib cage
Rib

In vertebrate anatomy, ribs are the long curved bones which form the ribcage. In most vertebrates, ribs surround the chest and protect the lungs, heart, and other internal Organ s of the thorax....
, about midway between shoulder and pelvis.

The Early Cretaceous
Early Cretaceous

The Early Cretaceous or the Lower Cretaceous , is the earlier of the two major divisions of the Cretaceous Period . It began about 146 million years ago....
 Cedar Mountain Formation
Cedar Mountain Formation

The Cedar Mountain Formation is the name given to distinctive sedimentary rocks in eastern Utah that occur between the underlying Morrison Formation and overlying Naturita Formation ....
 of Central Utah
Utah

The State of Utah is a western United States U.S. state of the United States. It was the List of U.S. states by date of statehood admitted to the United States on January 4, 1896....
 is full of highly polished red and black chert
Chert

Chert is a fine-grained silica-rich microcrystalline, cryptocrystalline or microfibrous sedimentary rock that may contain small fossils. It varies greatly in color , but most often manifests as gray, brown, grayish brown and light green to rusty red; its color is an expression of trace elements present in the rock, and both red and green ar...
s, which may partly represent gastroliths. Interestingly, the cherts may themselves contain fossils of ancient animals, such as corals. These stones do not appear to be associated with stream deposits and are rarely more than fist-sized, which is consistent with the idea that they are gastroliths. Gastroliths have sometimes been called 'Morrison stones' because they are often found in the Morrison Formation
Morrison Formation

The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has been the most fertile source of dinosaur fossils in North America....
 (named after the town of Morrison
Morrison, Colorado

The historic Town of Morrison is a Colorado municipalities#Home_Rule_Municipality in Jefferson County, Colorado, Colorado, United States. The population was 430 at the United States Census, 2000....
, west of Denver, Colorado
Colorado

The State of Colorado is a U.S. state located in the Mountain States of the United States of America. Colorado may also be considered to be a part of the Western United States and Southwestern United States regions of the United States....
), a late Jurassic
Jurassic

The Jurassic is a geologic period that extends from about annum to  Ma, that is, from the end of the Triassic to the beginning of the Cretaceous....
 formation roughly 150 million years old. Some gastroliths are made of petrified wood
Petrified wood

Petrified wood is a type of fossil: it consists of fossil wood where all the organic life have been replaced with minerals , while retaining the original structure of the wood....
.

Paleontologists are researching new methods of identifying gastroliths that have been found disassociated from animal remains, because of the important information they can provide. If the validity of such gastroliths can be verified, it may be possible to trace gastrolithic rocks back to their original sources. This may provide important information on how dinosaurs migrated. Because the number of suspected gastroliths is large, they could provide significant new insights into the lives and behavior of dinosaurs.

See also

  • Bezoar
    Bezoar

    A bezoar is a mass found trapped in the gastrointestinal system , though they can occur in other locations.There are several varieties of bezoar, some of which have inorganic constituents and others organic compound....
  • Calculus (medicine)
    Calculus (medicine)

    A calculus is a stone that forms in an organ or duct of the body. Formation of calculi is known as lithiasis. Stones cause a number of medical conditions....
  • Coprolite
    Coprolite

    A coprolite is fossilized animal dung. Coprolites are classified as Trace fossil as opposed to body fossils, as they give evidence for the animal's behaviour rather than morphology....
  • Enterolith
    Enterolith

    An enterolith is a mineral concretion or Calculus formed anywhere in the gastrointestinal system. Enteroliths are uncommon and usually incidental findings but, once found, they require at a minimum watchful waiting....
  • Regurgitalith
    Regurgitalith

    Regurgitaliths are the fossilised remains of stomach contents that have been Regurgitation by an animal. They are trace fossils and can be subdivided into ichnotaxon....