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Dactylorhiza

 
Dactylorhiza

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Dactylorhiza



 
 
Dactylorhiza Necker ex Nevski 1937, is a genus of terrestrial (ground-dwelling)
Terrestrial plant

A terrestrial plant is one that grows on land. Other types of plants are aquatic ecosystem , epiphytic and lithophytes ....
 plants in the orchid family (Orchidaceae
Orchidaceae

Orchidaceae is the largest Family of the flowering plants . Its name is derived from the genus Orchis.The Royal Botanic Gardens, Kew list 880 genus and nearly 22,000 accepted species, but the exact number is unknown because of taxonomic disputes....
).

The name Dactylorhiza is derived from Greek words d??t???? "daktylos" (finger) and ???a "rhiza" (root), referring to the palmately two- to five-lobed tubers of this genus. Dactylorhiza were previously classified under Orchis
Orchis

Orchis is a genus in the orchid family . This genus gets its name from the Greek ????? orchis, meaning "testicle", from the appearance of the paired subterranean Glossary of botanical terms....
 which has two round tubers .

These orchids are distributed throughout the subarctic and temperate northern hemisphere : in Europe
Europe

Europe is, conventionally, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally divided from Asia to its east by the water divide of the Ural Mountains, the Ural , the Caspian Sea, and by the Caucasus Mountains to the southeast....
, from Scandinavia
Scandinavia

Scandinavia is a historical and geographical subregion in northern Europe that includes the Scandinavian Peninsula. It consists of the kingdoms of Norway, Sweden, and Denmark; some authorities also include Finland and some might even include Iceland....
 to North Africa
North Africa

North Africa or Northern Africa is the northernmost region of the African continent, separated by the Sahara from Sub-Saharan Africa.Geopolitically, the United Nations subregion of Northern Africa includes the following seven countries or territories:...
; also on Madeira
Madeira

Madeira is a Portugal archipelago in the north Atlantic Ocean that lies between and . It is one of the Autonomous regions of Portugal, with Madeira Island and Porto Santo Island being the only inhabited islands....
, Iceland
Iceland

Iceland, officially the Republic of Iceland , is an island country located in the North Atlantic Ocean between mainland Europe and Greenland....
, West Asia
Asia

Asia is the world's largest and most populous continent. It covers 8.6% of the Earth's total surface area and, with over 4 billion people, it contains more than 60% of the world's current human population....
, North Asia, the Himalayas
Himalayas

The Himalaya Range or Himalayas for short , meaning "abode of snow" ), is a mountain range in Asia, separating the Indian subcontinent from the Tibetan Plateau....
, North America
North America

North America is the northern continent of the Americas, situated in the Earth's northern hemisphere and almost totally in the western hemisphere....
 and even in Alaska
Alaska

Alaska is the largest U.S. state of the United States by area; it is situated in the northwest extremity of the North American continent, with Canada to the east, the Arctic Ocean to the north, and the Pacific Ocean to the west and south, with Russia further west across the Bering Strait....
.

These terrestrial orchids grow in basic soils in wet meadows, bogs, heathland and in areas sparsely populated by trees.






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Encyclopedia


Dactylorhiza Necker ex Nevski 1937, is a genus of terrestrial (ground-dwelling)
Terrestrial plant

A terrestrial plant is one that grows on land. Other types of plants are aquatic ecosystem , epiphytic and lithophytes ....
 plants in the orchid family (Orchidaceae
Orchidaceae

Orchidaceae is the largest Family of the flowering plants . Its name is derived from the genus Orchis.The Royal Botanic Gardens, Kew list 880 genus and nearly 22,000 accepted species, but the exact number is unknown because of taxonomic disputes....
).

The name Dactylorhiza is derived from Greek words d??t???? "daktylos" (finger) and ???a "rhiza" (root), referring to the palmately two- to five-lobed tubers of this genus. Dactylorhiza were previously classified under Orchis
Orchis

Orchis is a genus in the orchid family . This genus gets its name from the Greek ????? orchis, meaning "testicle", from the appearance of the paired subterranean Glossary of botanical terms....
 which has two round tubers .

These orchids are distributed throughout the subarctic and temperate northern hemisphere : in Europe
Europe

Europe is, conventionally, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally divided from Asia to its east by the water divide of the Ural Mountains, the Ural , the Caspian Sea, and by the Caucasus Mountains to the southeast....
, from Scandinavia
Scandinavia

Scandinavia is a historical and geographical subregion in northern Europe that includes the Scandinavian Peninsula. It consists of the kingdoms of Norway, Sweden, and Denmark; some authorities also include Finland and some might even include Iceland....
 to North Africa
North Africa

North Africa or Northern Africa is the northernmost region of the African continent, separated by the Sahara from Sub-Saharan Africa.Geopolitically, the United Nations subregion of Northern Africa includes the following seven countries or territories:...
; also on Madeira
Madeira

Madeira is a Portugal archipelago in the north Atlantic Ocean that lies between and . It is one of the Autonomous regions of Portugal, with Madeira Island and Porto Santo Island being the only inhabited islands....
, Iceland
Iceland

Iceland, officially the Republic of Iceland , is an island country located in the North Atlantic Ocean between mainland Europe and Greenland....
, West Asia
Asia

Asia is the world's largest and most populous continent. It covers 8.6% of the Earth's total surface area and, with over 4 billion people, it contains more than 60% of the world's current human population....
, North Asia, the Himalayas
Himalayas

The Himalaya Range or Himalayas for short , meaning "abode of snow" ), is a mountain range in Asia, separating the Indian subcontinent from the Tibetan Plateau....
, North America
North America

North America is the northern continent of the Americas, situated in the Earth's northern hemisphere and almost totally in the western hemisphere....
 and even in Alaska
Alaska

Alaska is the largest U.S. state of the United States by area; it is situated in the northwest extremity of the North American continent, with Canada to the east, the Arctic Ocean to the north, and the Pacific Ocean to the west and south, with Russia further west across the Bering Strait....
.

These terrestrial orchids grow in basic soils in wet meadows, bogs, heathland and in areas sparsely populated by trees. They are tuber
Tuber

Tubers are various types of modified plant structures that are enlarged to store nutrients. They are used by plants to overwinter and regrow the next year and as a means of asexual reproduction....
ous geophytes. In a thickened underground stem, they can store a large amount of water to survive arid conditions.

The long leaves
Leaf

In botany, a leaf is an above-ground plant Organ specialized for photosynthesis. For this purpose, a leaf is typically flat and thin, to expose the cells containing chloroplast to light over a broad area, and to allow light to penetrate fully into the tissues....
 are lanceolate and, in most species, also speckled. They grow along a rather long stem
Plant stem

A stem is one of two main structural axes of a vascular plant. The stem is normally divided into nodes and internodes, the nodes hold buds which grow into one or more leaf, inflorescence , conifer cones or other stems etc....
 which reaches a height of 70-90 cm. Leaves higher on the stem are shorter than leaves lower on the stem.

The inflorescence
Inflorescence

An inflorescence is a group or cluster of flowers arranged on a Plant stem that is composed of a main branch or a complicated arrangement of branches....
, compared to the length of the plant, is rather short. It consists of a compact raceme
Raceme

A raceme is a type of inflorescence that is unbranched and indeterminate growth and bears pedicellate flowers — flowers having short floral stalks called Pedicel s — along the axis....
 with 25-50 flowers. These develop from axillary buds. The dominant colors are all shades of pink to red, sprinkled with darker speckles.

Many 'species' in this genus hybridise so readily that species boundaries themselves are vague, with regular name changes and no clear answers. A few species colonise very well onto fresh industrial wastes such as pulverised fuel ash
Pulverised Fuel Ash

Pulverised fuel ash , is a by product of pulverised fuel firing fired power stations. The fuel is pulverised into a fine powder, mixed with heated air and burned....
, where vast hybrid swarms can appear for a decade or more, before ecological succession replaces them.

Species

There are many species that cross-fertilize, giving an immense variation and complicating classification.

  • Dactylorhiza alpestris : Alpine Dactylorhiza (Pyrenees, Alps, Carpathians).
  • Dactylorhiza angustata (France).
  • Dactylorhiza aristata : Keyflower (E. China to Alaska).
    • Dactylorhiza aristata var. aristata : Keyflower (E. China to Alaska).
    • Dactylorhiza aristata var. kodiakensis : Kodiak Keyflower (Aleutian Is. to SW. Alaska).
  • Dactylorhiza armeniaca (Turkey) - has become synonym of Dactylorhiza euxina subsp. armeniaca (Hedrén) Kreutz
  • Dactylorhiza atlantica Kreutz & Vlaciha (Morocco)
  • Dactylorhiza baldshuanica (C. Asia).
  • Dactylorhiza baltica (Eastern Europe) (synonym of Dactylorhiza longifolia (Neuman) Aver.)
  • Dactylorhiza baumanniana (N. Greece).
    • Dactylorhiza baumanniana subsp. smolikana (B. Willing & E. Willing) H. Baumann & R. Lorenz (Greece)
  • Dactylorhiza bohemica (EC. Europe).
  • Dactylorhiza cordigera (Fr.) Soó (SE. Europe to Ukraine).
    • Dactylorhiza cordigera subsp. bosniaca (N. Balkan Pen).
    • Dactylorhiza cordigera subsp. cordigera (SE. Europe to Ukraine).
    • Dactylorhiza cordigera var. graeca (H.Baumann) Presser)
    • Dactylorhiza cordigera subsp. pindica (B. Willing & E. Willing) H. Baumann & R. Lorenz (NW. Greece).
    • Dactylorhiza cordigera var. rhodopeia Presser (Greece, Southeastern Europe, Europe)
    • Dactylorhiza cordigera subsp. siculorum (Romania to W. Ukraine).
  • Dactylorhiza ebudensis (Wief. ex R.M. Bateman & Denholm) P. Delforge : Hebridean
    Hebrides

    The Hebrides comprise a widespread and diverse archipelago off the west coast of Scotland. There are two main groups, the Inner and Outer Hebrides....
     Marsh Orchid
  • Dactylorhiza elata (Poir.) Soó : Stately Dactylorhiza (W. Europe to NW. Africa).
    • Dactylorhiza elata subsp. ambigua (Martrin-Donos) Kreutz
    • Dactylorhiza elata subsp. brennensis (W. Europe).
    • Dactylorhiza elata subsp. elata (NW. Africa.
    • Dactylorhiza elata subsp. mauritanica B.Baumann & H. Baumann (Morocco, Algeria)
    • Dactylorhiza elata subsp. sesquipedalis (SW. Europe to Sicilia).
  • Dactylorhiza euxina (Nevski) Czerep.
    • Dactylorhiza euxina subsp. armeniaca (Hedrén) Kreutz
  • Dactylorhiza flavescens (Turkey to C. Asia).
  • Dactylorhiza foliosa : Richly-leaved Dactylorhiza (Madeira).
  • Dactylorhiza fuchsii (Druce) Soó : Common Spotted Orchid, Fuch's Dactylorhiza (Europe to Siberia).
    • Dactylorhiza fuchsii subsp. carpatica (Batousek & Kreutz) Kreutz
    • Dactylorhiza fuchsii subsp. fuchsii (Europe to Siberia).
    • Dactylorhiza fuchsii subsp. hebridensis (W. Europe).
    • Dactylorhiza fuchsii subsp. meyeri (Rchb.f.) Kulikov & E.G.Philippov
    • Dactylorhiza fuchsii subsp. okellyi (Ireland, W. Great Britain).
    • Dactylorhiza fuchsii subsp. psychrophila (Europe to Siberia).
    • Dactylorhiza fuchsii var. sooana (Borsos) Kreutz (Hungary)
    • Dactylorhiza fuchsii var. sudetica (Poech ex Rchb.f.) H.Baumann
  • Dactylorhiza gervasiana (Sicilia to S. Italy).
  • Dactylorhiza graeca (N. Greece) - has become synonym of Dactylorhiza cordigera var. graeca (H.Baumann) Presser)
  • Dactylorhiza graggeriana (W. Himalaya).
  • Dactylorhiza hatagirea (Pakistan to SE. Tibet).
  • Dactylorhiza iberica (Greece to Iran).
  • Dactylorhiza ilgazica (N. Turkey) - now synonym of Dactylorhiza urvilleana subsp. ilgazica (Kreutz) Kreutz


Dactylorhiza Incarnata
Dactylorhiza Majalis Web
Dactylorhiza Majalis
  • Dactylorhiza incarnata
    Dactylorhiza incarnata

    Dactylorhiza incarnata is an orchid....
     (L.) Soó : Early Marsh Orchid
    • Dactylorhiza incarnata var. baumgartneriana (B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter) P.Delforge
    • Dactylorhiza incarnata subsp. coccinea
    • Dactylorhiza incarnata subsp. cruenta (Europe to Turkey).
    • Dactylorhiza incarnata subsp. gemmana (W. Europe).
    • Dactylorhiza incarnata subsp. incarnata (Europe to Mongolia).
    • Dactylorhiza incarnata nothosubsp. krylovii (W. Europe to Siberia).
    • Dactylorhiza incarnata subsp. lobelii (Norway to The Netherlands).
    • Dactylorhiza incarnata subsp. ochroleuca (Europe).
    • Dactylorhiza incarnata subsp. pulchella (Europe).
    • Dactylorhiza incarnata nothosubsp. versicolor (Europe)
  • Dactylorhiza insularis : Island Dactylorhiza (W. Medit. to WC. Italy).
  • Dactylorhiza kafiriana (NE. Afghanistan to W. Himalaya).
  • Dactylorhiza kalopissii E.Nelson (N. Greece).
    • Dactylorhiza kalopissii subsp. macedonica (J.Hölzinger & Künkele) Kreutz
    • Dactylorhiza kalopissii subsp. pythagorae (Gölz & H.R.Reinhard) Kreutz
  • Dactylorhiza kulikalonica (C. Asia).
  • Dactylorhiza lapponica
    Dactylorhiza lapponica

    Dactylorhiza lapponica is an orchid....
     (Laest.ex Hartm.) Soó (N. Europe).
    • Dactylorhiza lapponica subsp. angustata (Arv.-Touv.) Kreutz
    • Dactylorhiza lapponica subsp. rhaetica H. Baumann & R. Lorenz (Alps of Austria, Germany, Switzerland, Italy and France)
  • Dactylorhiza lapponica subsp. russowii (Klinge) H. Baumann & R. Lorenz (C. Europe to Siberia).
  • Dactylorhiza libanotica (Lebanon)
  • Dactylorhiza longifolia (Europe to C. Asia).
  • Dactylorhiza macedonica (N. Greece) - now a synonym of Dactylorhiza kalopissii subsp. macedonica (J.Hölzinger & Künkele) Kreutz
  • Dactylorhiza maculata
    Dactylorhiza maculata

    Heath Spotted-orchid is an orchid....
     (L.) Soó : Heath Spotted Orchid, Moorland Spotted Orchid (NW. Africa, Europe to Siberia).
    • Dactylorhiza maculata subsp. battandieri (N. Algeria).
    • Dactylorhiza maculata subsp. caramulensis (W. Europe).
    • Dactylorhiza maculata subsp. elodes (Europe).
    • Dactylorhiza maculata subsp. ericetorum : Heath Spotted Orchid (W. Europe).
    • Dactylorhiza maculata subsp. islandica (Iceland).
    • Dactylorhiza maculata subsp. kolaensis (Montell) Kreutz
    • Dactylorhiza maculata subsp. maculata (Europe to Siberia).
    • Dactylorhiza maculata subsp. maurusia (Morocco)
    • Dactylorhiza maculata subsp. podesta (Netherlands).
    • Dactylorhiza maculata subsp. rhoumensis (Great Britain)
    • Dactylorhiza maculata subsp. savogiensis (D.Tyteca & Gathoye) Kreutz
    • Dactylorhiza maculata subsp. schurii (Carpathians).
    • Dactylorhiza maculata subsp. sennia (Vollmar) Kreutz
    • Dactylorhiza maculata subsp. transsilvanica (SC. & SE. Europe).
  • Dactylorhiza magna (C. Asia).
  • Dactylorhiza majalis (Rchb.) P.F.Hunt & Summerh. : Broad-leaved Marsh Orchid, Western Marsh Orchid, Fan Orchid, Common Marsh Orchid (Europe).
    • Dactylorhiza majalis var. brevifolia (Rchb.f.) Kreutz
    • Dactylorhiza majalis subsp. calcifugiens (Denmark)
    • Dactylorhiza majalis subsp. cambrensis (Coastal Great Britain and Denmark) (now synonym of Dactylorhiza purpurella (T.Stephenson & T.A.Stephenson) Soó var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm 2005)
    • Dactylorhiza majalis subsp. majalis (Europe).
    • Dactylorhiza majalis subsp. occidentalis (W. & SW. Ireland, N. Great Britain).
    • Dactylorhiza majalis subsp. parvimajalis (D.Tyteca & Gathoye) Kreutz
    • Dactylorhiza majalis subsp. purpurella : Early Marsh Orchid, Northern Marsh Orchid (NW. Europe).
    • Dactylorhiza majalis subsp. sphagnicola
    • Dactylorhiza majalis subsp. turfosa (Alps to W. Carpathians) - has become a synonym of Dactylorhiza traunsteineri subsp. turfosa (F.Proch.) Kreutz
  • Dactylorhiza markusii : Markus' Dactylorhiza (N. Portugal to W. Spain and Italy).
  • Dactylorhiza nieschalkiorum (N. Turkey).
  • Dactylorhiza occidentalis (Pugsley) P. Delforge : Irish Marsh Orchid
  • Dactylorhiza osmanica (Turkey to Syria).
    • Dactylorhiza osmanica var. anatolica (Turkey).
    • Dactylorhiza osmanica var. osmanica (Turkey to Syria).
  • Dactylorhiza pindica (NW. Greece).
  • Dactylorhiza praetermissa : Leopard Marsh Orchid, Southern Marsh Orchid (W. & NW. Europe)
  • Dactylorhiza purpurella
    Dactylorhiza purpurella

    Dactylorhiza purpurella is an orchid....
     (T.Stephenson & T.A.Stephenson) Soó : Northern Marsh Orchid (Great Britain, Ireland).
    • Dactylorhiza purpurella var. maculosa (T. Stephenson)
    • Dactylorhiza purpurella var. purpurella
    • Dactylorhiza purpurella var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm 2005
  • Dactylorhiza pythagorae (E. Aegean Is.) - now a synonym of Dactylorhiza kalopissii subsp. pythagorae (Gölz & H.R.Reinhard) Kreutz
  • Dactylorhiza romana : Roman Dactylorhiza (Mediterranean)
  • Dactylorhiza saccifera (Brongn.) Soó : Sack-carrying Dactylorhiza (Mediterranean).
    • Dactylorhiza saccifera subsp. bithynica (H.Baumann) Kreutz
    • Dactylorhiza saccifera subsp. gervasiana (Tod.) Kreutz
  • Dactylorhiza salina (Caucasus to Amur)
  • Dactylorhiza sambucina : Elder-flowered Orchid (Europe). Photos
  • Dactylorhiza sudetica (Europe to Siberia)
  • Dactylorhiza traunsteineri
    Dactylorhiza traunsteineri

    Dactylorhiza traunsteineri is an orchid....
     (Saut. ex Rchb.) Soó : Narrow-leaved Marsh Orchid, Traunstein's Dactylorhiza (Europe to W. Siberia).
    • Dactylorhiza traunsteineri subsp. carpatica (Slovakia) - has become synonym of Dactylorhiza fuchsii subsp. carpatica (Batousek & Kreutz) Kreutz
    • Dactylorhiza traunsteineri subsp. turfosa (F.Proch.) Kreutz
    • Dactylorhiza traunsteineri subsp. curvifolia (N. & NE. Europe).
    • Dactylorhiza traunsteineri subsp. traunsteineri
      Dactylorhiza traunsteineri subsp. traunsteineri

      Dactylorhiza traunsteineri subsp. traunsteineri or Pugsley's Marsh Orchid occurs from Europe to West Siberia.The "traunsteinerioides" form is endemism to Great Britain and Ireland in scattered localities....
       (Europe to W. Siberia).
    • Dactylorhiza traunsteineri subsp. wirtgenii (Höppner) Kreutz
    • Dactylorhiza traunsteinerioides (Pugsley)Landwehr (synonym of Dactylorhiza traunsteineri subsp. traunsteineri)
  • Dactylorhiza umbrosa (W. & C. Asia to Siberia)
  • Dactylorhiza urvilleana (Steud.) H.Baumann & Künkele (N. & NE. Turkey to Iran)
    • Dactylorhiza urvilleana subsp. bithynica (H.Baumann) H. Baumann & R. Lorenz
    • Dactylorhiza urvilleana subsp. phoenissa B. Baumann & H. Baumann (Lebanon)
    • Dactylorhiza urvilleana subsp. ilgazica (Kreutz) Kreutz
  • Dactylorhiza viridis : Frog orchid (Subarctic and subalpine Northern Hemisphere).
    • Dactylorhiza viridis var. virescens (Temp. Asia, N. America)
    • Dactylorhiza viridis var. viridis (Subarctic and subalpine Northern Hemisphere)


Hybrids

Dactylorhiza X Aschersoniana 190505
Dactylorhiza X Braunii 030606
Note : nothosubspecies = a hybrid subspecies; nothovarietas = subvariety.
  • Dactylorhiza × abantiana (D. iberica × D. nieschalkiorum) (Turkey).
  • Dactylorhiza × aldenii (D. iberica × D. kalopissii) (Greece).
  • Dactylorhiza × altobracensis (D. maculata × D. sambucina) (France, Austria).
  • Dactylorhiza × aschersoniana (D. incarnata × D. majalis) (W. & C. Europe).
    • Dactylorhiza × aschersoniana nothosubsp. aschersoniana (W. & C. Europe).
    • Dactylorhiza × aschersoniana nothosubsp. templinensis (D. incarnata subsp. ochroleuca × D. majalis) (C. Europe).
    • Dactylorhiza × aschersoniana nothovar. uliginosa (D. incarnata subsp.pulchella × D. majalis) (C. Europe).
  • Dactylorhiza × baicalica (D. incarnata subsp. cruenta × D. salina) (Siberia).
  • Dactylorhiza × balabaniana (D. iberica × D. urvilleana) (Turkey).
  • Dactylorhiza × bayburtiana (D. euxina × D. umbrosa) (Turkey).
  • Dactylorhiza × beckeriana (C. Europe).
  • Dactylorhiza × boluiana (D. nieschalkiorum × D. saccifera) (Turkey).
  • Dactylorhiza × bourdonii (D. brennensis × D. incarnata) (France).
  • Dactylorhiza × braunii (D. fuchsii × D. majalis) (Europe).
    • Dactylorhiza × braunii nothosubsp. braunii (Europe).
    • Dactylorhiza × braunii nothosubsp. lilacina (D fuchsii × D. majalis subsp. turfosa) (EC. Europe).
    • Dactylorhiza × braunii nothosubsp. monticola (D. fuchsii subsp. psychrophila × D. majalis) (Europe).
    • Dactylorhiza × braunii nothosubsp. smitakii (D. fuchsii subsp. sooana × D. majalis) (EC. Europe). te
  • Dactylorhiza × breviceras (D. osmanica × D. urvilleana) (Turkey).
  • Dactylorhiza × carnea (D. incarnata × D. maculata subsp. ericetorum) (W. Europe).
    • Dactylorhiza × carnea nothosubsp. ampolai (D. incarnata subsp. cruenta × D. maculata) (Europe).
    • Dactylorhiza × carnea nothosubsp. carnea (W. Europe).
    • Dactylorhiza × carnea nothosubsp. maculatiformis. (D. incarnata × D. maculata) (W. Europe).
  • Dactylorhiza × claudiopolitana (D. incarnata × D. schurii) (Europe.
  • Dactylorhiza × conigerum (D. maculata × D. viridis) (W. Europe).
  • Dactylorhiza × csatoi (D. cordigera × D. maculata) (SE. Europe).
  • Dactylorhiza czerniakowskae (C. Asia).
  • Dactylorhiza × daunia (D. romana × D. saccifera) (S. Europe).
  • Dactylorhiza × delamainii (D. elata subsp. sesquipedalis × D. maculata) (SW. Europe).
  • Dactylorhiza × dinglensis (D. maculata subsp. ericetorum × D. majalis subsp. occidentalis) (W. Europe).
    • Dactylorhiza × dinglensis nothosubsp. dinglensis (W. Europe).
    • Dactylorhiza × dinglensis nothosubsp. robertsii (D. maculata subsp. ericetorum × D. majalis subsp. cambrensi) (Great Britain).
    • Dactylorhiza × dinglensis nothosubsp. senayi (D. maculata subsp. elodes × D. majalis) (Europe).
    • Dactylorhiza × dinglensis nothosubsp. townsendiana (D. maculata subsp. ericetorum × D. majalis) (Europe).
    • Dactylorhiza × dinglensis nothosubsp. vermeuleniana (D. maculata × D. majalis) (W. Europe).
  • Dactylorhiza × drucei (D. majalis × D. viridis) (W. Europe)
  • Dactylorhiza × dubreuilhii (D. elata subsp. sesquipedalis × D. incarnata) (W. Europe).
  • Dactylorhiza × dufftiana (D. majalis × D. traunsteineri) (Europe).
  • Dactylorhiza × dufftii (D. incarnata × D. traunsteineri) (Europe).
    • Dactylorhiza × dufftii nothosubsp. dufftii (Europe).
    • Dactylorhiza × dufftii nothosubsp. gotlandica (D incarnata subsp. ochroleuca × D. traunsteineri) (Europe). Tuber geophyte
    • Dactylorhiza × dufftii nothosubsp. stenkyrkae (D. incarnata subsp. cruenta × D. traunsteineri) (Europe).
  • Dactylorhiza × erdingeri (D. sambucina × D. viridis) (W. Europe).
  • Dactylorhiza euxina (NE. Turkey to Caucasus).
    • Dactylorhiza euxinavar. euxina (NE. Turkey to Caucasus).
    • Dactylorhiza euxinavar. markowitschii (NE. Turkey to Caucasus).
  • Dactylorhiza × flixensis (D. incarnata subsp. pulchella × D. traunsteineri.) (Switzerland).
  • Dactylorhiza × formosa (D. maculata subsp. ericetorum × D. purpurella) (W. Europe).
  • Dactylorhiza × fourkensis (D. baumanniana × D. sambucina) (Greece).
  • Dactylorhiza × gabretana (D. incarnata × D. maculata × D. sambucina) (Europe).
  • Dactylorhiza × genevensis (D. incarnata × D. latifolia × D. maculata) (Europe).
  • Dactylorhiza × godferyana (D. majalis × D. praetermissa) (W. Europe).
  • Dactylorhiza × grandis (D. fuchsii × D. praetermissa) (W. Europe).
  • Dactylorhiza × guilhotii (D. incarnata × D. viridis) (W. Europe).
  • Dactylorhiza × guillaumeae (D. incarnata × D. sambucina) (W. Europe).
  • Dactylorhiza × gustavssonii (D. iberica × D. saccifera) (Greece to Turkey).
  • Dactylorhiza × hallii (D maculata subsp. ericetorum × D. praetermissa) (W. Europe).
    • Dactylorhiza × hallii nothosubsp. hallii (W. Europe).
    • Dactylorhiza × hallii nothosubsp. nummiana (D. maculata subsp. elodes × D. praetermissa) (W. Europe).
  • Dactylorhiza × hochreutinerana (D. alpestris × D. incarnata) (W. Europe).
  • Dactylorhiza × insignis (D. praetermissa × D. purpurella) (W. Europe).
  • Dactylorhiza × ishorica (''D. incarnata'' × ''D. longifolia'') (European Russia).
  • ''Dactylorhiza'' × ''jenensis'' (''D. maculata'' subsp. ''ericetorum'' × ''D. traunsteineri'') (W. & NC. Europe)
  • ''Dactylorhiza'' × ''jestrebiensis'' (''D. bohemica'' × ''D. majalis'') (EC. Europe).
  • ''Dactylorhiza'' × ''juennensis'' (''D. fuchsii'' × ''D. lapponica'') (C. Europe).
  • ''Dactylorhiza'' × ''katarana'' (''D kalopissii'' × ''D. saccifera'') (Greece).
  • ''Dactylorhiza'' × ''kelleriana'' (''D. fuchsii'' × ''D. traunsteineri'') (Europe).
  • ''Dactylorhiza'' × ''kerasovinensis'' (''D. pindica'' × ''D. saccifera'') (Greece).
  • ''Dactylorhiza'' × ''kerneriorum'' (''D. fuchsii'' × ''D. incarnata'') (Europe).
    • ''Dactylorhiza'' × ''kerneriorum'' nothosubsp. ''kerneriorum'' (Europe).
    • ''Dactylorhiza'' × ''kerneriorum'' nothosubsp. ''lillsundica'' (''D. fuchsii'' × ''D. incarnata'' subsp. ''ochroleuca'') (N. & W. Europe).
    • ''Dactylorhiza'' × ''kerneriorum'' nothosubsp. ''variablis'' (''D. fuchsii'' subsp. ''hebridensis'' × ''D. incarnata'') (W. Europe). *''Dactylorhiza'' × ''komiensis'' (''D. hebridensis'' × ''D. maculata'') (E. Europe).
  • ''Dactylorhiza'' × ''kopdagiana'' (''D. iberica'' × ''D. umbrosa'') (Turkey).
  • ''Dactylorhiza'' × ''koutsourana'' (''D. baumanniana'' × ''D. smolikana'') (Greece).
  • ''Dactylorhiza'' × ''kuuskiae'' (''D.longifolia'' × ''D. traunsteineri'') (E. Europe).
  • ''Dactylorhiza'' × ''latirella'' (''D. incarnata'' × ''D. purpurella'') (W. Europe).
  • ''Dactylorhiza'' × ''lehmannii'' (''D. incarnata'' × ''D. russowii'') (Europe).
  • ''Dactylorhiza'' × ''megapolitana'' (''D. fuchsii'' × ''D. russowii'') (C. Europe).
  • ''Dactylorhiza'' × ''metsowonensis'' (''D. kalopissii'' × ''D. sambucina'') (Greece).
  • ''Dactylorhiza'' × ''mixtum'' (''D. fuchsii'' × ''D. viridis'') (W. Europe).
  • ''Dactylorhiza'' × ''mulignensis'' (''D. incarnata'' subsp. ''pulchella'' × ''D. majalis'') (C. Europe).
  • ''Dactylorhiza'' × ''nevskii'' (''D. osmanica'' × ''D. umbrosa'') (Turkey).
  • ''Dactylorhiza'' × ''ornonensis'' (''D. elata'' subsp. ''sesquipedalis'' × ''D. incarnata'' × ''D. maculata'') (W. Europe).
  • ''Dactylorhiza'' × ''paridaeniana'' (''D. elata'' subsp. ''sesquipedalis'' × ''D. praetermissa'') (W. Europe).
  • ''Dactylorhiza'' × ''pontica'' (''D. urvilleana'' × ''D. viridis'') (Turkey)
  • ''Dactylorhiza'' × ''prochazkana'' (''D. bohemica'' × ''D. maculata'') (EC. Europe).
  • ''Dactylorhiza'' × ''renzii'' (''D. incarnata'' × ''D. nieschalkiorum'') (Turkey).
  • ''Dactylorhiza'' × ''rizeana'' (''D. euxina'' × ''D. urvilleana'') (Turkey).
  • ''Dactylorhiza'' × ''rombucina'' (''D. romana'' × ''D. sambucina'') (C. Europe).
  • ''Dactylorhiza'' × ''ruppertii'' (''D. majalis'' × ''D. sambucina'') (Europe).
  • ''Dactylorhiza'' × ''salictina'' (''D. pindica'' × ''D. smolikana'') (Greece).
  • ''Dactylorhiza'' × ''serbica'' (''D. incarnata'' × ''D. saccifera'') (Europe).
  • ''Dactylorhiza'' × ''serreana'' (''D. graeca'' × ''D. lagotis'') (Greece).
  • ''Dactylorhiza'' × ''sivasiana'' (''D. umbrosa'' × ''D. urvilleana'') (Turkey).
  • ''Dactylorhiza'' × ''sooi'' (''D. alpestris'' × ''D. fuchsii.'') (Europe).
  • ''Dactylorhiza'' × ''souflikensis'' (''D. baumanniana'' × ''D. pindica'') (Greece).
  • ''Dactylorhiza'' × ''stagni-novi'' (''D. brennensis'' × ''D. fuchsii'') (Europe).
  • ''Dactylorhiza'' × ''szaboiana'' (''D. cordigera'' × ''D. sudetica'') (SE. Europe).
  • ''Dactylorhiza'' × ''transiens'' (''D. fuchsii'' × ''D. maculata'' subsp. ''ericetorum'') (Europe)
    • ''Dactylorhiza'' × ''transiens'' nothosubsp. ''corylensis'' (''D. fuchsii'' subsp. ''hebridensis'' × ''D. maculata'')
    • ''Dactylorhiza'' × ''transiens''subsp. ''ericetorum'' (Europe). Tuber geophyte
    • ''Dactylorhiza'' × ''transiens'' nothosubsp. ''transiens'' (Europe). Tuber geophyte
  • ''Dactylorhiza'' × ''turcestanicum'' (''D. umbrosa'' × ''D. viridis'') (C. Asia).
  • ''Dactylorhiza'' × ''vallis-peenae'' (''D. majalis'' × ''D. russowii'') (C. Europe).
  • ''Dactylorhiza'' × ''venusta'' (''D. fuchsii'' × ''D. purpurella'') (Europe).
    • ''Dactylorhiza'' × ''venusta'' nothosubsp. ''hebridella'' (''D. fuchsii'' subsp. ''hebridensis'' × ''D. purpurella'') (Great Britain).
    • ''Dactylorhiza'' × ''venusta'' nothosubsp. ''venusta'' (Europe)
  • ''Dactylorhiza'' × ''viridella'' (''D. purpurella'' × ''D. viridis'') (W. Europe).
  • ''Dactylorhiza'' × ''vitosana'' (''D. saccifera'' × ''D. sambucina'') (SE. Europe).
  • ''Dactylorhiza'' × ''vogtiana'' (''D. iberica'' × ''D. incarnata'') (Turkey).
  • ''Dactylorhiza'' × ''vorasica'' (''D. cordigera'' × ''D. sambucina'') (Greece).
  • ''Dactylorhiza'' × ''weissenbachiana'' (''D. incarnata'' × ''D. lapponica'') (C. Europe).
  • ''Dactylorhiza'' × ''wiefelspuetziana'' (''D. maculata'' × ''D. sphagnicola'') (W. Europe).


This list follows the ''World checklist of monocotyledons'', periodically amended from the "Orchid Research Newsletter", both of Kew Botanical Gardens.

External links