Woodsia ilvensis
Encyclopedia
Woodsia ilvensis, commonly known as Oblong Woodsia, is a fern
Fern
A fern is any one of a group of about 12,000 species of plants belonging to the botanical group known as Pteridophyta. Unlike mosses, they have xylem and phloem . They have stems, leaves, and roots like other vascular plants...

 found in North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 and northern Eurasia
Eurasia
Eurasia is a continent or supercontinent comprising the traditional continents of Europe and Asia ; covering about 52,990,000 km2 or about 10.6% of the Earth's surface located primarily in the eastern and northern hemispheres...

. Also known as Rusty Woodsia or Rusty Cliff Fern, it is typically found on sunny, exposed cliffs and rocky slopes and on thin, dry, acidic soils.

Distribution

Its distribution is circumpolar and is most abundant in Scandinavia
Scandinavia
Scandinavia is a cultural, historical and ethno-linguistic region in northern Europe that includes the three kingdoms of Denmark, Norway and Sweden, characterized by their common ethno-cultural heritage and language. Modern Norway and Sweden proper are situated on the Scandinavian Peninsula,...

, the Ural
Ural Mountains
The Ural Mountains , or simply the Urals, are a mountain range that runs approximately from north to south through western Russia, from the coast of the Arctic Ocean to the Ural River and northwestern Kazakhstan. Their eastern side is usually considered the natural boundary between Europe and Asia...

 and Altai mountains and the eastern United States. It is also found in Japan, Alaska
Alaska
Alaska is the largest state in the United States by area. It is situated in the northwest extremity of the North American continent, with Canada to the east, the Arctic Ocean to the north, and the Pacific Ocean to the west and south, with Russia further west across the Bering Strait...

, Canada, coastal Greenland
Greenland
Greenland is an autonomous country within the Kingdom of Denmark, located between the Arctic and Atlantic Oceans, east of the Canadian Arctic Archipelago. Though physiographically a part of the continent of North America, Greenland has been politically and culturally associated with Europe for...

 and various European locations including the Alps
Alps
The Alps is one of the great mountain range systems of Europe, stretching from Austria and Slovenia in the east through Italy, Switzerland, Liechtenstein and Germany to France in the west....

.

It is considered "Threatened" or "Endangered" in the states of Illinois
Illinois
Illinois is the fifth-most populous state of the United States of America, and is often noted for being a microcosm of the entire country. With Chicago in the northeast, small industrial cities and great agricultural productivity in central and northern Illinois, and natural resources like coal,...

, Iowa
Iowa
Iowa is a state located in the Midwestern United States, an area often referred to as the "American Heartland". It derives its name from the Ioway people, one of the many American Indian tribes that occupied the state at the time of European exploration. Iowa was a part of the French colony of New...

, and Maryland
Maryland
Maryland is a U.S. state located in the Mid Atlantic region of the United States, bordering Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware to its east...

 and "Presumed Extirpated" in Ohio
Ohio
Ohio is a Midwestern state in the United States. The 34th largest state by area in the U.S.,it is the 7th‑most populous with over 11.5 million residents, containing several major American cities and seven metropolitan areas with populations of 500,000 or more.The state's capital is Columbus...

. Also found in West Virginia
West Virginia
West Virginia is a state in the Appalachian and Southeastern regions of the United States, bordered by Virginia to the southeast, Kentucky to the southwest, Ohio to the northwest, Pennsylvania to the northeast and Maryland to the east...

 and North Carolina
North Carolina
North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

, it is the most common Woodsia
Woodsia
Woodsia is a genus of ferns in the family Woodsiaceae. It includes the following species :*Woodsia alpina S. F. Gray*Woodsia appalachiana T. M. C. Taylor*Woodsia cochisensis Windham...

species in the US.

Its UK distribution is confined to Angus
Angus
Angus is one of the 32 local government council areas of Scotland, a registration county and a lieutenancy area. The council area borders Aberdeenshire, Perth and Kinross and Dundee City...

 and the Moffat Hills
Moffat Hills
The Moffat hills are a range of hills in the Southern Uplands of Scotland. They form a distinctly triangular shape with a west facing side, a north facing side, and a south-east facing side. It is 17 kilometres from east to west across this triangle and some 16 kilometres north to south...

 in Scotland, north Wales
Wales
Wales is a country that is part of the United Kingdom and the island of Great Britain, bordered by England to its east and the Atlantic Ocean and Irish Sea to its west. It has a population of three million, and a total area of 20,779 km²...

 and Teesdale
Teesdale
Teesdale is a dale, or valley, of the east side of the Pennines in England. Large parts of Teesdale fall within the North Pennines Area of Outstanding Natural Beauty - the second largest AONB in England and Wales. The River Tees rises below Cross Fell, the highest hill in the Pennines, and its...

 and the Lake District
Lake District
The Lake District, also commonly known as The Lakes or Lakeland, is a mountainous region in North West England. A popular holiday destination, it is famous not only for its lakes and its mountains but also for its associations with the early 19th century poetry and writings of William Wordsworth...

 in England. There are fewer than 90 wild clumps in the whole of the UK, where it is on the edge of its natural range and is considered to be "Endangered".

Discovery and identification

The plant was first identified as a separate species from specimens collected in Scotland in Bolton's 1785 publication . Bolton distinguished between Acrostichum ilvense and Acrostichum alpina, now Woodsia ilvensis and Woodsia alpina
Woodsia alpina
Woodsia alpina, commonly known as Alpine Woodsia, is a fern found in northern latitudes in North America and Eurasia. Also known as Northern Woodsia or Alpine Cliff Fern, it is typically found in crevices, scree slopes and cliffs containing slate and calcareous rocks, especially...

respectively, which had previously been conflated. The genus Woodsia was established in 1810 by Robert Brown
Robert Brown (botanist)
Robert Brown was a Scottish botanist and palaeobotanist who made important contributions to botany largely through his pioneering use of the microscope...

, who named it named after the English botanist Joseph Woods
Joseph Woods
Joseph Woods FLS FGS 24 August 1776-1864 was an English Quaker architect, botanist and geologist born in the village of Stoke Newington, a few miles north of the City of London...

. "" is the genitive form of the Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 name for the island of Elba
Elba
Elba is a Mediterranean island in Tuscany, Italy, from the coastal town of Piombino. The largest island of the Tuscan Archipelago, Elba is also part of the National Park of the Tuscan Archipelago and the third largest island in Italy after Sicily and Sardinia...

.

The leaves are typically 6 inches long and 1 inch wide, with stiff, erected pointed tips and cut into 12 nearly opposite stemless leaflets. The underside of the leaves are covered in white woolly fibres, which later turn rusty brown.

Victorian collectors and modern conservation

Oblong Woodsia came under severe threat from Victorian
Victorian era
The Victorian era of British history was the period of Queen Victoria's reign from 20 June 1837 until her death on 22 January 1901. It was a long period of peace, prosperity, refined sensibilities and national self-confidence...

 fern collectors in the mid 19th century in Scotland, especially in the Moffat Hills
Moffat Hills
The Moffat hills are a range of hills in the Southern Uplands of Scotland. They form a distinctly triangular shape with a west facing side, a north facing side, and a south-east facing side. It is 17 kilometres from east to west across this triangle and some 16 kilometres north to south...

. These hills once had the most extensive UK populations of the species but there now remain only a few small colonies whose future is under threat. This period of collecting became known as Pteridomania
Pteridomania
Pteridomania or Fern-Fever was a craze for ferns. Victorian decorative arts presented the fern motif in pottery, glass, metal, textiles, wood, printed paper, and sculpture, with ferns "appearing on everything from christening presents to gravestones and memorials."-Description:Pteridomania, meaning...

 (or "fern-fever"). The Royal Botanic Garden Edinburgh
Royal Botanic Garden Edinburgh
The Royal Botanic Garden Edinburgh is a scientific centre for the study of plants, their diversity and conservation, as well as a popular tourist attraction. Originally founded in 1670 as a physic garden to grow medicinal plants, today it occupies four sites across Scotland — Edinburgh,...

plan to use cultivated specimens and a spore bank to restore depleted wild populations.
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