Theriosuchus
Encyclopedia
Theriosuchus is an extinct genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of atoposaurid
Atoposauridae
Atoposauridae is a family of crocodile-like archosaurs. Members of the family have been found from France, Spain, Portugal, Romania, Germany, Kazakhstan, the United Kingdom, the United States, Cameroon, and Thailand.-Phylogeny:...

 mesoeucrocodylia
Mesoeucrocodylia
Mesoeucrocodylia is the name of the clade that includes Eusuchia and the paraphyletic group Mesosuchia. The group appeared during the Early Jurassic, and continues to the present day....

n from the Late Jurassic
Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.2 ± 4.0 to 145.5 ± 4.0 million years ago , which is preserved in Upper Jurassic strata. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age...

 of Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, the Early Cretaceous
Early Cretaceous
The Early Cretaceous or the Lower Cretaceous , is the earlier or lower of the two major divisions of the Cretaceous...

 of Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 and the Late Cretaceous
Late Cretaceous
The Late Cretaceous is the younger of two epochs into which the Cretaceous period is divided in the geologic timescale. Rock strata from this epoch form the Upper Cretaceous series...

 of Romania
Romania
Romania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...

. The five species are T. pusillus from England
England
England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...

 (Owen, 1879), T. ibericus from Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

 (Brinkmann, 1992), T. guimarotae from Portugal
Portugal
Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...

 (Schwarz & Salisbury, 2005), T. sympiestodon from Romania (Jeremy E. Martin, Márton Rabi & Zoltán Csiki, 2010) and T. grandinaris from Berriasian
Berriasian
In the geological timescale, the Berriasian is an age or stage of the Early or Lower Creteceous. It is the oldest or lowest subdivision in the entire Cretaceous. It spanned between 145.5 ± 4.0 Ma and 140.2 ± 3.0 Ma...

-Barremian
Barremian
The Barremian is an age in the geologic timescale between 130.0 ± 1.5 Ma and 125.0 ± 1.0 Ma). It is a subdivision of the Early Cretaceous epoch...

 of Thailand
Thailand
Thailand , officially the Kingdom of Thailand , formerly known as Siam , is a country located at the centre of the Indochina peninsula and Southeast Asia. It is bordered to the north by Burma and Laos, to the east by Laos and Cambodia, to the south by the Gulf of Thailand and Malaysia, and to the...

 (Lauprasert et al., 2011).

Theriosuchus was a common terrestrial mesoeucrocodylian in Europe during the Cretaceous. Fossils belonging to the genus are abundant at several European sites. It was one of the few basal mesoeucrocodylians to exist in Laurasia during the Late Cretaceous, a time when many basal forms such as notosuchia
Notosuchia
Notosuchia is a suborder of primarily Gondwanan mesoeucrocodylian crocodylomorphs that lived during the Cretaceous. Fossils have been found from South America, Africa, and Asia...

ns and araripesuchids
Araripesuchus
Araripesuchus is a genus of extinct crocodyliform whose species existed during the Cretaceous period of the late Mesozoic era some 125 to 65 million years ago. They are generally considered to be notosuchians , characterized by their varied teeth types and distinct skull elements...

 were present in Gondwana
Gondwana
In paleogeography, Gondwana , originally Gondwanaland, was the southernmost of two supercontinents that later became parts of the Pangaea supercontinent. It existed from approximately 510 to 180 million years ago . Gondwana is believed to have sutured between ca. 570 and 510 Mya,...

. Some other basal mesoeucrocodylians present in Europe at this time were Doratodon
Doratodon
Doratodon is an extinct genus of crocodylomorph that was once believed to be a dinosaur.- Species :The type species is D. carcharidens from the Late Cretaceous of Austria, whilst the second known species is D. ibericus from the Late Cretaceous of Spain.-References:*Company, J., Suberbiola, X.P.,...

and Ischyrochampsa
Ischyrochampsa
Ischyrochampsa is an extinct genus of Late Cretaceous mesoeucrocodylian. Fossils of the type species I. meridionalis are late Campanian in age and were found in the commune of Saint-Estève-Janson in Bouches-du-Rhône, France. Material is also known from Spain. At an estimated length of over ,...

.

There is a 58-million-year gap in the fossil record of the genus that spans between the last occurrence of Early Cretaceous
Early Cretaceous
The Early Cretaceous or the Lower Cretaceous , is the earlier or lower of the two major divisions of the Cretaceous...

 species and the appearance of the Late Cretaceous T. sympiestodon. Described in 2010, T. sympiestodon was found in the Haţeg Basin
Hateg
Hațeg is a town in Hunedoara County, Romania with a population of 12,507. Three villages are administered by the town: Nălațvad, Silvașu de Jos and Silvașu de Sus.Țara Hațegului is the region around Hațeg town...

 of Romania
Romania
Romania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...

. During the Late Cretaceous, this area was part of Haţeg Island
Hateg Island
Hațeg Island was a large offshore island in the Tethys Sea during the Cretaceous period. It was situated in an area corresponding to the region around modern-day Hațeg, Romania...

, which was part of an archipelago
Archipelago
An archipelago , sometimes called an island group, is a chain or cluster of islands. The word archipelago is derived from the Greek ἄρχι- – arkhi- and πέλαγος – pélagos through the Italian arcipelago...

 that spanned the Tethys Ocean
Tethys Ocean
The Tethys Ocean was an ocean that existed between the continents of Gondwana and Laurasia during the Mesozoic era before the opening of the Indian Ocean.-Modern theory:...

.
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