Podaxis pistillaris
Encyclopedia
Podaxis pistillaris is a very distinctive relative of the puffballs. It grows to 15 cm high and has a hard, woody stem
Plant stem
A stem is one of two main structural axes of a vascular plant. The stem is normally divided into nodes and internodes, the nodes hold buds which grow into one or more leaves, inflorescence , conifer cones, roots, other stems etc. The internodes distance one node from another...

. The large cap
Cap
A cap is a form of headgear. Caps have crowns that fit very close to the head and have no brim or only a visor. They are typically designed for warmth and, when including a visor, blocking sunlight from the eyes...

, which protects the blackish spore
Spore
In biology, a spore is a reproductive structure that is adapted for dispersal and surviving for extended periods of time in unfavorable conditions. Spores form part of the life cycles of many bacteria, plants, algae, fungi and some protozoa. According to scientist Dr...

-bearing tissue, splits, and usually falls away at maturity, allowing the spores to be dispersed by wind. Large numbers may appear after soaking rains. It thrives in desert
Desert
A desert is a landscape or region that receives an extremely low amount of precipitation, less than enough to support growth of most plants. Most deserts have an average annual precipitation of less than...

s and semi-deserts of Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

 and other countries. In the Hawaiian Island
Hawaii
Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...

s, it is frequently encountered along roadsides and in disturbed areas on the dry sides of the islands, especially in the Kona
Kona District, Hawaii
Kona is the name of a moku or district on the Big Island of Hawaii in the State of Hawaii. In the current system of administration of Hawaii County, the moku of Kona is divided into North Kona District and South Kona District . The term "Kona" is sometimes used to refer to its largest town,...

 area of Hawaii and the Kihei area of Maui
Maui
The island of Maui is the second-largest of the Hawaiian Islands at and is the 17th largest island in the United States. Maui is part of the state of Hawaii and is the largest of Maui County's four islands, bigger than Lānai, Kahoolawe, and Molokai. In 2010, Maui had a population of 144,444,...

.

Older synonyms for this species include Lycoperdon pistillare L. (1771) and Scleroderma pistillare (L.) Pers. (1801).

Spores

The spores are usually 10–14 (–16) by (8–) 9–12 µm broadly oval
Oval
An oval is any curve resembling an egg or an ellipse, such as a Cassini oval. The term does not have a precise mathematical definition except in one area oval , but it may also refer to:* A sporting arena of oval shape** a cricket field...

 to sub-globose, smooth yellow
Yellow
Yellow is the color evoked by light that stimulates both the L and M cone cells of the retina about equally, with no significant stimulation of the S cone cells. Light with a wavelength of 570–590 nm is yellow, as is light with a suitable mixture of red and green...

 to deep reddish-brown with a double wall, truncate base, and apical pore. Older spore measurements have varied considerably. Species from Australian collections appear to be more subglobose than those seen from the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

, raising the possibility that the latter are not the same species.

Organisation

It is an agaric
Agaric
An agaric is a type of fungal fruiting body characterized by the presence of a pileus that is clearly differentiated from the stipe , with lamellae on the underside of the pileus. "Agaric" can also refer to a basidiomycete species characterized by an agaric-type fruiting body...

, though it has lost hymenophoral organization and the ability to forcibly discharge its basidiospores and become "secotioid". Although considered by many to be a "stalked puffball", Podaxis pistillaris is more closely allied with the shaggy mane (Coprinus comatus
Coprinus comatus
Coprinus comatus, the shaggy ink cap, lawyer's wig, or shaggy mane, is a common fungus often seen growing on lawns, along gravel roads and waste areas. The young fruit bodies first appear as white cylinders emerging from the ground, then the bell-shaped caps open out. The caps are white, and...

) than with puffballs .
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