Pagurus bernhardus
Encyclopedia
Pagurus bernhardus is the common marine hermit crab
Hermit crab
Hermit crabs are decapod crustaceans of the superfamily Paguroidea. Most of the 1100 species possess an asymmetrical abdomen which is concealed in an empty gastropod shell that is carried around by the hermit crab.-Description:...

 of Europe's
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 coasts. It is sometimes referred to as the common hermit crab or soldier crab. It is about 3.5 centimetres (1.4 in) long, and is found in both rocky and sandy areas, from the Arctic
Arctic
The Arctic is a region located at the northern-most part of the Earth. The Arctic consists of the Arctic Ocean and parts of Canada, Russia, Greenland, the United States, Norway, Sweden, Finland, and Iceland. The Arctic region consists of a vast, ice-covered ocean, surrounded by treeless permafrost...

 waters of Iceland
Iceland
Iceland , described as the Republic of Iceland, is a Nordic and European island country in the North Atlantic Ocean, on the Mid-Atlantic Ridge. Iceland also refers to the main island of the country, which contains almost all the population and almost all the land area. The country has a population...

, Svalbard
Svalbard
Svalbard is an archipelago in the Arctic, constituting the northernmost part of Norway. It is located north of mainland Europe, midway between mainland Norway and the North Pole. The group of islands range from 74° to 81° north latitude , and from 10° to 35° east longitude. Spitsbergen is the...

 and Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

 as far south as southern Portugal
Portugal
Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...

, but its range does not extend as far as the Mediterranean Sea
Mediterranean Sea
The Mediterranean Sea is a sea connected to the Atlantic Ocean surrounded by the Mediterranean region and almost completely enclosed by land: on the north by Anatolia and Europe, on the south by North Africa, and on the east by the Levant...

. It can be found in pools on the upper shore and at the mean tide level down to a depth of approximately 140 metres (459.3 ft), with smaller specimens generally found in rock pools around the middle shore and lower shore regions, with larger individuals at depth. P. bernhardus is an omnivorous
Omnivore
Omnivores are species that eat both plants and animals as their primary food source...

 detritivore
Detritivore
Detritivores, also known as detritophages or detritus feeders or detritus eaters or saprophages, are heterotrophs that obtain nutrients by consuming detritus . By doing so, they contribute to decomposition and the nutrient cycles...

 that opportunistically scavenges
Scavenger
Scavenging is both a carnivorous and herbivorous feeding behavior in which individual scavengers search out dead animal and dead plant biomass on which to feed. The eating of carrion from the same species is referred to as cannibalism. Scavengers play an important role in the ecosystem by...

 for carrion
Carrion
Carrion refers to the carcass of a dead animal. Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters include vultures, hawks, eagles, hyenas, Virginia Opossum, Tasmanian Devils, coyotes, Komodo dragons, and burying beetles...

, and which can also filter feed when necessary.

Pagurus bernhardus uses shells
Seashell
A seashell or sea shell, also known simply as a shell, is a hard, protective outer layer created by an animal that lives in the sea. The shell is part of the body of the animal. Empty seashells are often found washed up on beaches by beachcombers...

 of a number of gastropod species for protection, including Littorina littorea
Common Periwinkle
The common periwinkle or winkle, scientific name Littorina littorea, is a species of small edible sea snail, a marine gastropod mollusk which has gills and an operculum, and is classified within the family Littorinidae, the periwinkles....

, Littorina obtusata
Littorina obtusata
Littorina obtusata, common name the flat periwinkle, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles.-Distribution:This marine species occurs wherever brown seaweeds grow...

, Nucella lapillus
Dog Whelk
The dog whelk, dogwhelk, or Atlantic dogwinkle, scientific name Nucella lapillus, is a species of predatory sea snail, a carnivorous marine gastropod mollusc in the family Muricidae, the rock snails....

and Buccinum
Buccinum
Buccinum is a genus of medium-sized sea snails with an operculum, marine gastropod mollusks in the family Buccinidae, the true whelks.Snails in this genus are commonly called whelks, a name that is however shared with several related and unrelated species...

. In the warmer parts of its range, the sea anemone
Sea anemone
Sea anemones are a group of water-dwelling, predatory animals of the order Actiniaria; they are named after the anemone, a terrestrial flower. Sea anemones are classified in the phylum Cnidaria, class Anthozoa, subclass Zoantharia. Anthozoa often have large polyps that allow for digestion of larger...

 Calliactis parasitica
Calliactis parasitica
Calliactis parasitica is a species of sea anemone associated with hermit crabs. It lives in the eastern Atlantic Ocean and Mediterranean Sea at depths between the intertidal zone and . It is up to in size, with up to 700 tentacles, and is very variable in colour. The relationship between C...

is often found growing on the shell occupied by Pagurus bernhardus. In colder waters, this rôle is filled by Hormathia digitata.
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