Kepodactylus
Encyclopedia
Kepodactylus is an extinct genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of pterodactyloid pterosaur
Pterosaur
Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...

 from the Kimmeridgian
Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age or stage in the Late or Upper Jurassic epoch or series. It spans the time between 155.7 ± 4 Ma and 150.8 ± 4 Ma . The Kimmeridgian follows the Oxfordian and precedes the Tithonian....

-Tithonian
Tithonian
In the geologic timescale the Tithonian is the latest age of the Late Jurassic epoch or the uppermost stage of the Upper Jurassic series. It spans the time between 150.8 ± 4 Ma and 145.5 ± 4 Ma...

-age Upper Jurassic Morrison Formation
Morrison Formation
The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone and limestone and is light grey, greenish...

 of Colorado
Colorado
Colorado is a U.S. state that encompasses much of the Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains...

, USA.

In 1992 a team from the Denver Museum of Natural History dug up a specimen of the dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

 Stegosaurus
Stegosaurus
Stegosaurus is a genus of armored stegosaurid dinosaur. They lived during the Late Jurassic period , some 155 to 150 million years ago in what is now western North America. In 2006, a specimen of Stegosaurus was announced from Portugal, showing that they were present in Europe as well...

 stenops
in the Garden Park
Garden Park, Colorado
Garden Park, in southcentral Colorado, is known for its Jurassic dinosaurs and the role the specimens played in the infamous bone wars of the late 19th century. Located north of Cañon City, Colorado, the name originates from the area providing vegetables to the miners at nearby Cripple Creek in...

, Colorado
Colorado
Colorado is a U.S. state that encompasses much of the Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains...

. In the quarry also disarticulated smaller bones from other animals were found, among which those of a pterosaur, new to science.

In 1996 Jerald Harris and Kenneth Carpenter
Kenneth Carpenter
Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life...

 named the new genus. The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 is Kepodactylus insperatus. The genus name is derived from Greek, kepos, "garden", a reference to Garden Park and daktylos, "finger", referring to the typical wing finger of pterosaurs. The specific name means "unhoped-for" in Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

, alluding to the fact that the researchers hoped to find a dinosaur, and did not expect a pterosaur.

The genus is based on holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 DMNH 21684
Denver Museum of Nature and Science
The Denver Museum of Nature & Science is a municipal natural history and science museum in Denver, Colorado. It is a resource for informal science education in the Rocky Mountain region. A variety of exhibitions, programs, and activities help museum visitors learn about the natural history of...

, consisting of a cervical
Neck
The neck is the part of the body, on many terrestrial or secondarily aquatic vertebrates, that distinguishes the head from the torso or trunk. The adjective signifying "of the neck" is cervical .-Boner anatomy: The cervical spine:The cervical portion of the human spine comprises seven boney...

 vertebra, humerus
Humerus
The humerus is a long bone in the arm or forelimb that runs from the shoulder to the elbow....

, several finger bones, and a metatarsal. Kepodactylus was similar to Mesadactylus
Mesadactylus
Mesadactylus is an extinct genus of pterosaur from the Kimmeridgian-Tithonian-age Upper Jurassic Morrison Formation of Colorado, USA....

but larger (wingspan around 2.5 m [8.2 ft]), and with additional pneumatic foramina (holes to allow air from air sacks to enter the bones) in the vertebrae and humerus. The describers concluded that the species was a member of the Pterodactyloidea
Pterodactyloidea
Pterodactyloidea forms one of the two suborders of pterosaurs , and contains the most derived members of this group of flying reptiles...

 and within this group, using the phylogeny of David Unwin, a member of a clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

 that is now known as Lophocratia. It was regarded as a potentially valid genus in the most recent review of Morrison pterosaurs.
Unwin considered it a basal member of the Dsungaripteroidea
Dsungaripteroidea
Dsungaripteroidea is a group of pterosaurs within the suborder Pterodactyloidea.-Classification:Listing of families and genera after Unwin 2006 unless otherwise noted.* Superfamily Dsungaripteroidea** Herbstosaurus** Kepodactylus...

sensu Unwin.
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