Mesadactylus
Encyclopedia
Mesadactylus is an extinct genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of pterosaur
Pterosaur
Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...

 from the Kimmeridgian
Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age or stage in the Late or Upper Jurassic epoch or series. It spans the time between 155.7 ± 4 Ma and 150.8 ± 4 Ma . The Kimmeridgian follows the Oxfordian and precedes the Tithonian....

-Tithonian
Tithonian
In the geologic timescale the Tithonian is the latest age of the Late Jurassic epoch or the uppermost stage of the Upper Jurassic series. It spans the time between 150.8 ± 4 Ma and 145.5 ± 4 Ma...

-age Upper Jurassic Morrison Formation
Morrison Formation
The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone and limestone and is light grey, greenish...

 of Colorado
Colorado
Colorado is a U.S. state that encompasses much of the Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains...

, USA.

The genus was named in 1989 by James Jensen
James A. Jensen
James A. Jensen , was an American paleontologist. His extensive collecting program at BYU in the Utah-Colorado region which spanned 23 years was comparable in terms of the number of specimens collected to that of Barnum Brown during the early 20th century. He was given the name "Dinosaur Jim"...

 and Kevin Padian
Kevin Padian
Kevin Padian is a Professor of Integrative Biology at the University of California, Berkeley, Curator of Paleontology, University of California Museum of Paleontology and President of the National Center for Science Education. Padian's area of interest is in vertebrate evolution, especially the...

. The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 is Mesadactylus ornithosphyos. The holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 is BYU 2024, a synsacrum
Synsacrum
The synsacrum is a skeletal structure, mainly described in birds and dinosaurs, in which the sacrum is extended by incorporation of additional fused or partially-fused caudal or lumbar vertebrae. The ilium of the pelvis is attached to the synsacrum and, in some species, may also be fused to it...

 of seven sacral vertebrae, featuring a for pterosaurs unique complete fusion of the spinae into a supraneural blade, a character, as the specific name indicates more typical for birds, at first leading Jensen to assign the fossil to a bird, Paleopteryx.

Further referred associated remains include arms bones, pectoral girdle bones, vertebrae (including cervix
Neck
The neck is the part of the body, on many terrestrial or secondarily aquatic vertebrates, that distinguishes the head from the torso or trunk. The adjective signifying "of the neck" is cervical .-Boner anatomy: The cervical spine:The cervical portion of the human spine comprises seven boney...

 and sacral
Sacrum
In vertebrate anatomy the sacrum is a large, triangular bone at the base of the spine and at the upper and back part of the pelvic cavity, where it is inserted like a wedge between the two hip bones. Its upper part connects with the last lumbar vertebra, and bottom part with the coccyx...

), and femorae
Femur
The femur , or thigh bone, is the most proximal bone of the leg in tetrapod vertebrates capable of walking or jumping, such as most land mammals, birds, many reptiles such as lizards, and amphibians such as frogs. In vertebrates with four legs such as dogs and horses, the femur is found only in...

. Additional material was described in 2004 (including a partial braincase) and 2006; in the latter publication, the authors suggested that its larger contemporary Kepodactylus
Kepodactylus
Kepodactylus is an extinct genus of pterodactyloid pterosaur from the Kimmeridgian-Tithonian-age Upper Jurassic Morrison Formation of Colorado, USA....

could be the same animal, although there are minor differences.

Jensen and Padian classified Mesadactylus as a pterodactyloid. In 2007 S. Christopher Bennett claimed that the holotype and the referred material came from different forms and that, while the last was indeed of a pterodactyloid nature, the synsacrum belonged to a member of the Anurognathidae.
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