Angustinaripterus
Encyclopedia
Angustinaripterus was a basal pterosaur
Pterosaur
Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...

, belonging to the Breviquartossa, and discovered at Dashanpu
Dashanpu Formation
The Dashanpu Formation is a Mid to Late Jurassic rock formation in China, most notable for the wealth of dinosaurs that have been excavated from the area...

 near Zigong
Zigong
Zigong , ancient name Ziliujing and Gongjing, is a prefecture-level city and the third largest city in Sichuan Province, in southwest China.-Geography:...

 in the Szechuan province of China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

.

Angustinaripterus was named in 1983 by He Xinlu.
The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 is Angustinaripterus longicephalus. The genus name is derived from Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 angustus, "narrow" and naris, "nostril", combined with Latinized Greek pteron, "wing". The specific name is derived from Latin longus, "long", and Greek kephale, "head".

The holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

, ZDM T8001, is a single skull with lower jaws, found in 1981 by researchers from the Zigong Historical Museum of the Salt Industry, in the Xiaximiao Formation (Bathonian
Bathonian
In the geologic timescale the Bathonian is an age or stage of the Middle Jurassic. It lasted from approximately 167.7 Ma to around 164.7 Ma...

).

The skull, of which the left side is severely damaged, is very elongated and flat. The back part is missing; in its conserved state it has a length of 192 millimetres; the total length in a complete state was estimated at 201 millimetres. On its top is a low crest, two to three millimetres high. The nares are long, slit-like and positioned above and in front of the large skull openings, the fenestrae antorbitales, with which they are not confluent. Of the jaws, which are very straight, the front part is lacking. There are six pairs of teeth in the maxillae
Maxilla
The maxilla is a fusion of two bones along the palatal fissure that form the upper jaw. This is similar to the mandible , which is also a fusion of two halves at the mental symphysis. Sometimes The maxilla (plural: maxillae) is a fusion of two bones along the palatal fissure that form the upper...

 and three pairs in the praemaxillae. In the mandible
Mandible
The mandible pronunciation or inferior maxillary bone forms the lower jaw and holds the lower teeth in place...

 there are at least ten pairs of teeth, perhaps twelve. The back teeth are small, the front teeth are very long, robust and curved, pointing moderately forwards. At the front they form a large, intermeshing "prey grab", that may have been used to snatch fish from the water surface. The teeth of Angustinaripterus resemble those of Dorygnathus
Dorygnathus
Dorygnathus was a genus of pterosaur that lived in Europe during the Early Jurassic period, 180 million years ago when shallow seas flooded much of the continent. It had a short 1.5 meter wingspan, and a relatively small triangular sternum, which is where its flight muscles attached. Its skull...

.

He placed Angustinaripterus into the Rhamphorhynchidae. Because of the derived morphology and the large geographical distance with comparable European forms He also created a special subfamily Angustinaripterinae, of which Angustinaripterus itself is the only known member; because of this redundancy the concept is rarely used. He concluded that Angustinaripterus was directly related to the Scaphognathinae. David Unwin however, considers it a member of the other rhamphorhynchid subgroup: the Rhamphorhynchinae.

Peter Wellnhofer
Peter Wellnhofer
Peter Wellnhofer is a German paleontologist at the "Bayerische Staatssammlung fur Paläontologie" in Munich. He is best known for his work on the various fossil specimens of Archaeopteryx or "Urvogel", the first known bird...

in 1991, assuming the skull length was 16.5 centimetres, estimated the wingspan at 1.6 metres.

Literature

  • He, Xinlu; Yang, Daihuan; & Su, Chunkang, 1983, "A New Pterosaur from the Middle Jurassic of Dashanpu, Zigong, Sichuan", Journal of the Chengdu College of Geology supplement 1, pp. 27-33 [title in 1991 English translation by Will Downs]
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