Xerobdellidae
Encyclopedia
Xerobdellidae are a small family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of "jawed leeches". The latter might be (but probably are not) a natural, monophyletic group of hirudiniform
Hirudiniformes
The Hirudiniformes are one of the currently-accepted suborders of the proboscisless leeches . Their most well-known member is the European Medical Leech, Hirudo medicinalis, and indeed most of the blood-sucking "worms" as which leeches are generally perceived belong to this group...

 proboscisless leeches
Arhynchobdellida
The proboscisless leeches, Arhynchobdellida, are classified as an order of the Hirudinea. But leech taxonomy and systematics will eventually be revised in due time, not because there are many uncertainties about their phylogeny, but because the major clades of clitellate annelids - and whether the...

. Xerobdellidae have three jaws and five pairs of eyes, the fourth and fifth being separated by one or two eyeless segments. One each of the mere three genera
Genera
Genera is a commercial operating system and development environment for Lisp machines developed by Symbolics. It is essentially a fork of an earlier operating system originating on the MIT AI Lab's Lisp machines which Symbolics had used in common with LMI and Texas Instruments...

 placed herein occurs in Chile
Chile
Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

 (Mesobdella) and Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 (Xerobdella). Diestecostoma is more widespread and found in Central
Central America
Central America is the central geographic region of the Americas. It is the southernmost, isthmian portion of the North American continent, which connects with South America on the southeast. When considered part of the unified continental model, it is considered a subcontinent...

 and northern South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

. This peculiar distribution strongly suggests they are a relict
Relict
A relict is a surviving remnant of a natural phenomenon.* In biology a relict is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas....

 Pangaea
Pangaea
Pangaea, Pangæa, or Pangea is hypothesized as a supercontinent that existed during the Paleozoic and Mesozoic eras about 250 million years ago, before the component continents were separated into their current configuration....

n group, which had already been present by the start of the Jurassic
Jurassic
The Jurassic is a geologic period and system that extends from about Mya to  Mya, that is, from the end of the Triassic to the beginning of the Cretaceous. The Jurassic constitutes the middle period of the Mesozoic era, also known as the age of reptiles. The start of the period is marked by...

 250 million years ago.

These leeches resemble the Haemadipsidae and were included there by many authors, but this has always been controversial. Their status as a distinct family is supported by sequence
DNA sequence
The sequence or primary structure of a nucleic acid is the composition of atoms that make up the nucleic acid and the chemical bonds that bond those atoms. Because nucleic acids, such as DNA and RNA, are unbranched polymers, this specification is equivalent to specifying the sequence of...

 analysis of the nuclear 18S and 28S rDNA
RDNA
rDNA may stand for:*ribosomal DNA*recombinant DNARDNA is also an acronym for*Reformed Druids of North America...

 and mitochondrial COI
COI
COI may refer to:Places* City of Industry, CaliforniaPolitics and society* Center of Information, agency that followed the Office of Strategic Services and preceded the Central Intelligence Agency...

 gene
Gene
A gene is a molecular unit of heredity of a living organism. It is a name given to some stretches of DNA and RNA that code for a type of protein or for an RNA chain that has a function in the organism. Living beings depend on genes, as they specify all proteins and functional RNA chains...

s as well as the anatomy
Anatomy
Anatomy is a branch of biology and medicine that is the consideration of the structure of living things. It is a general term that includes human anatomy, animal anatomy , and plant anatomy...

 of their sexual organs and nephridia; the latter are located at the belly rather than along the body sides as in the Haemadipsidae proper.

They feed on blood
Blood
Blood is a specialized bodily fluid in animals that delivers necessary substances such as nutrients and oxygen to the cells and transports metabolic waste products away from those same cells....

 – typically of amphibian
Amphibian
Amphibians , are a class of vertebrate animals including animals such as toads, frogs, caecilians, and salamanders. They are characterized as non-amniote ectothermic tetrapods...

s – and small invertebrate
Invertebrate
An invertebrate is an animal without a backbone. The group includes 97% of all animal species – all animals except those in the chordate subphylum Vertebrata .Invertebrates form a paraphyletic group...

except Mesobdella, which is exclusively a blood-feeder and has been recorded to attack humans on occasion.
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