Steganinae
Encyclopedia
The Steganinae Hendel, 1917, is the smaller of two subfamilies in the fruit fly family Drosophilidae
Drosophilidae
Drosophilidae is a diverse, cosmopolitan family of flies, which includes fruit flies. Another family of flies called Tephritidae also includes fruit flies. The best known species of Drosophilidae is Drosophila melanogaster, within the genus Drosophila, and this species Is used extensively for...

. The other subfamily is the Drosophilinae
Drosophilinae
The Drosophilinae are the largest subfamily in the Drosophilidae. The other subfamily is the Steganinae.-Phylogeny:Many molecular studies have addressed small parts of the phylogenetic tree. Most of these studies are limited to species of the genus Drosophila...

.

Diagnosis

The subfamily is monophyletic (Grimali 1990; Sidorenko 2002), but can not be characterised by a single morphological character that distinguishes this subfamily from the Drosophilinae
Drosophilinae
The Drosophilinae are the largest subfamily in the Drosophilidae. The other subfamily is the Steganinae.-Phylogeny:Many molecular studies have addressed small parts of the phylogenetic tree. Most of these studies are limited to species of the genus Drosophila...

. See for a discussion on this subfamily "Drosophila
Drosophila
Drosophila is a genus of small flies, belonging to the family Drosophilidae, whose members are often called "fruit flies" or more appropriately pomace flies, vinegar flies, or wine flies, a reference to the characteristic of many species to linger around overripe or rotting fruit...

: A Laboratory Handbook" by M. Ashburner
Michael Ashburner
Michael Ashburner FRS is a biologist and emeritus Professor in the Department of Genetics at University of Cambridge. He is also the former joint-head of the European Bioinformatics Institute of the European Molecular Biology Laboratory .Born in Sussex, England, Ashburner attended High Wycombe...

, S. Hawley, K. Golic (not reproduced here due to copyrights).
  • D. A. Grimaldi 1990. A phylogenetic, revised classification of genera in the Drosophilidae
    Drosophilidae
    Drosophilidae is a diverse, cosmopolitan family of flies, which includes fruit flies. Another family of flies called Tephritidae also includes fruit flies. The best known species of Drosophilidae is Drosophila melanogaster, within the genus Drosophila, and this species Is used extensively for...

     (Diptera
    Diptera
    Diptera , or true flies, is the order of insects possessing only a single pair of wings on the mesothorax; the metathorax bears a pair of drumstick like structures called the halteres, the remnants of the hind wings. It is a large order, containing an estimated 240,000 species, although under half...

    ). Bulletin of the American Museum of Natural History 197: 1-128.
  • V. S. Sidorenko 2002. Phylogeny of the tribe Steganini Hendel and some related taxa (Diptera
    Diptera
    Diptera , or true flies, is the order of insects possessing only a single pair of wings on the mesothorax; the metathorax bears a pair of drumstick like structures called the halteres, the remnants of the hind wings. It is a large order, containing an estimated 240,000 species, although under half...

    , Drosophilidae
    Drosophilidae
    Drosophilidae is a diverse, cosmopolitan family of flies, which includes fruit flies. Another family of flies called Tephritidae also includes fruit flies. The best known species of Drosophilidae is Drosophila melanogaster, within the genus Drosophila, and this species Is used extensively for...

    ). Far Eastern Entomologist 111: 1-20.

Phylogeny

The phylogenetic relationships within the subfamily have not yet been confirmed by molecular studies, and are partially based on traditional cladistic
Cladistics
Cladistics is a method of classifying species of organisms into groups called clades, which consist of an ancestor organism and all its descendants . For example, birds, dinosaurs, crocodiles, and all descendants of their most recent common ancestor form a clade...

 methods using primarily morphological characteristics and partly based on expert opinions. The classification below might be not approved by all scholars in the field, but gives a reasonable overview of our current knowledge:

Tribe: Steganini
Subtribe: Steganina
Genus: Eostegana
Genus: Stegana

Subtribe: Leucophengina
Genus: Allopygaea
Genus: Cacoxenus
Genus: Leucophenga


Tribe: Gitonini
Subtribe: Gitonina
Genus: Amiota
Genus: Apenthecia
Genus: Apsiphortica
Genus: Cacoxenus
Genus: Crincosia
Genus: Erima
Genus: Gitona
Genus: Paraleucophenga
Genus: Paraphortica
Genus: Phortica

Subtribe: Acletoxenina
Genus: Acletoxenus
Genus: Hyalistata
Genus: Mayagueza
Genus: Pseudiastata
Genus: Rhinoleucophenga
Genus: Trachyleucophenga

Subtribe unknown
Genus: Electrophortica
Genus: Pararhinoleucophenga
Genus: Parastegana
Genus: Pseudostegana


Tribe unknown
Genus: Soederbomia
Genus: Pyrgometopa


For species within the various genera, see fix Taxodros
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