Nottingham Catchfly
Encyclopedia
Silene nutans is a flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

 in the genus Silene
Silene
Silene is a genus of flowering plants in the family Caryophyllaceae. Common names include campion and catchfly....

, most commonly known as Nottingham Catchfly.

Description

Silene nutans is a diploid, mainly outcrossing
Outcrossing
Outcrossing is the practice of introducing unrelated genetic material into a breeding line. It increases genetic diversity, thus reducing the probability of all individuals being subject to disease or reducing genetic abnormalities...

, herb
Herb
Except in botanical usage, an herb is "any plant with leaves, seeds, or flowers used for flavoring, food, medicine, or perfume" or "a part of such a plant as used in cooking"...

aceous, perennial plant
Perennial plant
A perennial plant or simply perennial is a plant that lives for more than two years. The term is often used to differentiate a plant from shorter lived annuals and biennials. The term is sometimes misused by commercial gardeners or horticulturalists to describe only herbaceous perennials...

.
It grows up to 25–80 cm (9.8–31.5 ) tall, from a branching, wood
Wood
Wood is a hard, fibrous tissue found in many trees. It has been used for hundreds of thousands of years for both fuel and as a construction material. It is an organic material, a natural composite of cellulose fibers embedded in a matrix of lignin which resists compression...

y stock with a thick taproot
Taproot
A taproot is an enlarged, somewhat straight to tapering plant root that grows vertically downward. It forms a center from which other roots sprout laterally.Plants with taproots are difficult to transplant...

. The lower leaves are up to 75 millimetres (3 in) long, spathulate and have a long stalk, while leaves higher on the plant are lanceolate, subsessile and acute; all the leaves are covered in soft hairs. The flowers are 18 millimetre (0.708661417322835 in) wide, 12 millimetre (0.47244094488189 in) long, and drooping, on short, viscid stalks. The petals are white or pinkish and divided into two narrow lobes. Each flower remains open for three nights as a means of preventing self-fertilisation; the flower reveals one whorl
Whorl (botany)
In botany, a whorl is an arrangement of sepals, petals, leaves, or branches in which all the parts are attached at the same point and surround or wrap around the stem.There are four whorls in a general flower...

 of stamen
Stamen
The stamen is the pollen producing reproductive organ of a flower...

s on the first night, the second whorl of stamens on the second night, and the three styles on the third night. The seeds are 10–22 mm (0.393700787401575–0.866141732283465 ) wide and kidney
Kidney
The kidneys, organs with several functions, serve essential regulatory roles in most animals, including vertebrates and some invertebrates. They are essential in the urinary system and also serve homeostatic functions such as the regulation of electrolytes, maintenance of acid–base balance, and...

-shaped.

Distribution

Silene nutans is widespread across Europe, from southern Spain and Italy north to the British Isles and Scandinavia, and is also found across large parts of Asia. It has been introduced to North America, where it is known as the Eurasian catchfly. It is found in the U.S. states of Michigan
Michigan
Michigan is a U.S. state located in the Great Lakes Region of the United States of America. The name Michigan is the French form of the Ojibwa word mishigamaa, meaning "large water" or "large lake"....

, Ohio
Ohio
Ohio is a Midwestern state in the United States. The 34th largest state by area in the U.S.,it is the 7th‑most populous with over 11.5 million residents, containing several major American cities and seven metropolitan areas with populations of 500,000 or more.The state's capital is Columbus...

, New York
New York
New York is a state in the Northeastern region of the United States. It is the nation's third most populous state. New York is bordered by New Jersey and Pennsylvania to the south, and by Connecticut, Massachusetts and Vermont to the east...

, Vermont
Vermont
Vermont is a state in the New England region of the northeastern United States of America. The state ranks 43rd in land area, , and 45th in total area. Its population according to the 2010 census, 630,337, is the second smallest in the country, larger only than Wyoming. It is the only New England...

 and Maine
Maine
Maine is a state in the New England region of the northeastern United States, bordered by the Atlantic Ocean to the east and south, New Hampshire to the west, and the Canadian provinces of Quebec to the northwest and New Brunswick to the northeast. Maine is both the northernmost and easternmost...

.

Silene nutans can sometimes be found in the very widespread MG1 (Arrhenatherum elatius grassland)
British NVC community MG1
British NVC community MG1 is one of the mesotrophic grassland communities in the British National Vegetation Classification system.It is a very widespread community throughout the British lowlands...

 community of the British National Vegetation Classification
British National Vegetation Classification
The British National Vegetation Classification or NVC is a system of classifying natural habitat types in Great Britain according to the vegetation they contain....

, and thus can be found where Arrhenatherum elatius
Arrhenatherum elatius
-Introduction:Arrhenatherum elatius, with the common names false oat-grass, tall oat-grass, tall meadow oat, onion couch and tuber oat-grass, is a perennial species of grass, common in the temperate regions of Europe....

(false oat grass) or Dactylis glomerata
Dactylis glomerata
Dactylis glomerata, Cock's-foot and orchard grass, is a common species of grass in the genus Dactylis, native throughout most of Europe, temperate Asia, and northern Africa.-Distribution:...

(cocksfoot) occur.

Ecology

Silene nutans is a steppe
Steppe
In physical geography, steppe is an ecoregion, in the montane grasslands and shrublands and temperate grasslands, savannas, and shrublands biomes, characterized by grassland plains without trees apart from those near rivers and lakes...

 species across most of its range. At the periphery of its distribution
Species distribution
Species distribution is the manner in which a biological taxon is spatially arranged. Species distribution is not to be confused with dispersal, which is the movement of individuals away from their area of origin or from centers of high population density. A similar concept is the species range. A...

, it has a patchy distribution in xeric habitats, such as open grassland
Grassland
Grasslands are areas where the vegetation is dominated by grasses and other herbaceous plants . However, sedge and rush families can also be found. Grasslands occur naturally on all continents except Antarctica...

s and on rock outcrop
Outcrop
An outcrop is a visible exposure of bedrock or ancient superficial deposits on the surface of the Earth. -Features:Outcrops do not cover the majority of the Earth's land surface because in most places the bedrock or superficial deposits are covered by a mantle of soil and vegetation and cannot be...

s at forest margins, on both acidic and alkaline substrates (pH
PH
In chemistry, pH is a measure of the acidity or basicity of an aqueous solution. Pure water is said to be neutral, with a pH close to 7.0 at . Solutions with a pH less than 7 are said to be acidic and solutions with a pH greater than 7 are basic or alkaline...

 3.8–8.0). In the far north of its range, S. nutans is characteristic of maritime cliff
Cliff
In geography and geology, a cliff is a significant vertical, or near vertical, rock exposure. Cliffs are formed as erosion landforms due to the processes of erosion and weathering that produce them. Cliffs are common on coasts, in mountainous areas, escarpments and along rivers. Cliffs are usually...

s.

S. nutans flowers during the night, and produces a strong floral scent to attract its pollinator
Pollinator
A pollinator is the biotic agent that moves pollen from the male anthers of a flower to the female stigma of a flower to accomplish fertilization or syngamy of the female gamete in the ovule of the flower by the male gamete from the pollen grain...

s, which are mostly night-flying moths. Chemical compound
Chemical compound
A chemical compound is a pure chemical substance consisting of two or more different chemical elements that can be separated into simpler substances by chemical reactions. Chemical compounds have a unique and defined chemical structure; they consist of a fixed ratio of atoms that are held together...

s in the scent include benzyl acetate
Benzyl acetate
Benzyl acetate is an organic compound with the molecular formula C9H10O2. It is the ester formed by condensation of benzyl alcohol and acetic acid....

 and benzaldehyde
Benzaldehyde
Benzaldehyde is an organic compound consisting of a benzene ring with a formyl substituent. It is the simplest aromatic aldehyde and one of the most industrially useful. This colorless liquid has a characteristic pleasant almond-like odor...

.

Silene nutans is the host plant for the leaf mining
Leaf miner
Leaf miner is a term used to describe the larvae of many different species of insect which live in and eat the leaf tissue of plants. The vast majority of leaf-mining insects are moths , sawflies and flies , though some beetles and wasps also exhibit this behavior.Like Woodboring beetles, leaf...

 moth, Coleophora galbulipennella
Coleophora galbulipennella
Coleophora galbulipennella is a moth of the family Coleophoridae or leaf miner. It is known from Europe, where it is found from Sweden to Spain, Italy and Greece and from Great Britain to southern Russia.The wingspan is 15–17 mm...

.

Nottingham

The common name
Common name
A common name of a taxon or organism is a name in general use within a community; it is often contrasted with the scientific name for the same organism...

 Nottingham Catchfly commemorates the former occurrence of S. nutans on the walls of Nottingham Castle
Nottingham Castle
Nottingham Castle is a castle in Nottingham, England. It is located in a commanding position on a natural promontory known as "'Castle Rock'", with cliffs high to the south and west. In the Middle Ages it was a major royal fortress and occasional royal residence...

, and the species was chosen to represent the unitary authority
Unitary authority
A unitary authority is a type of local authority that has a single tier and is responsible for all local government functions within its area or performs additional functions which elsewhere in the relevant country are usually performed by national government or a higher level of sub-national...

 of Nottingham
Nottingham
Nottingham is a city and unitary authority in the East Midlands of England. It is located in the ceremonial county of Nottinghamshire and represents one of eight members of the English Core Cities Group...

 as its County flower
County flowers of the United Kingdom
The following are the flowers selected for the historic counties of the United Kingdom in Plantlife's 2002 "County Flowers" campaign...

. Despite this association, Nottingham Catchflies no longer occur in either the city of Nottingham or the wider county of Nottinghamshire
Nottinghamshire
Nottinghamshire is a county in the East Midlands of England, bordering South Yorkshire to the north-west, Lincolnshire to the east, Leicestershire to the south, and Derbyshire to the west...

.
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