Loncosaurus
Encyclopedia
Loncosaurus was a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of ornithopod
Ornithopod
Ornithopods or members of the clade Ornithopoda are a group of ornithischian dinosaurs that started out as small, bipedal running grazers, and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world, and dominated the North American...

 dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

 from the Upper Cretaceous of Provincia de Santa Cruz, Argentina
Argentina
Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

. The type
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 (and only known) species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 is Loncosaurus argentinus, described by the famous Argentinian
Argentina
Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

 paleontologist Florentino Ameghino
Florentino Ameghino
Florentino Ameghino was an Argentine naturalist, paleontologist, anthropologist and zoologist.Born in Luján, son of Italian immigrants, Ameghino was a self-taught naturalist, and focused his study on the lands of the southern Pampas...

, but is considered a dubious
Nomen dubium
In zoological nomenclature, a nomen dubium is a scientific name that is of unknown or doubtful application...

 name. Details on this animal are often contradictory, befitting a genus that was long confused for a theropod.

History

Ameghino named this dinosaur in either 1898 or 1899, from a proximal femur
Femur
The femur , or thigh bone, is the most proximal bone of the leg in tetrapod vertebrates capable of walking or jumping, such as most land mammals, birds, many reptiles such as lizards, and amphibians such as frogs. In vertebrates with four legs such as dogs and horses, the femur is found only in...

 (MACN-1629) and tooth
Tooth
Teeth are small, calcified, whitish structures found in the jaws of many vertebrates that are used to break down food. Some animals, particularly carnivores, also use teeth for hunting or for defensive purposes. The roots of teeth are embedded in the Mandible bone or the Maxillary bone and are...

 found near Rio Sehuen, Santa Cruz, in either the Cardiel Formation
Cardiel Formation
The Cardiel Formation is a Mesozoic geologic formation. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation.-Paleofauna:* Clasmodosaurus spatula - "Teeth." -...

 (most sources) or the Matasiete Formation (both being Upper Cretaceous).

Either way, he thought the remains belonged to a "megalosaurid
Megalosaurid
Megalosauridae was a family of relatively primitive tetanuran theropod dinosaurs, order Saurischia. They were small-to-large carnivores with sharp teeth and three claws on each hand. Some members of this group were Megalosaurus, Eustreptospondylus, Streptospondylus and Torvosaurus...

" dinosaur, a carnivore
Carnivore
A carnivore meaning 'meat eater' is an organism that derives its energy and nutrient requirements from a diet consisting mainly or exclusively of animal tissue, whether through predation or scavenging...

, which Friedrich von Huene
Friedrich von Huene
Friedrich von Huene was a German paleontologist who named more dinosaurs in the early 20th century than anyone else in Europe.-Biography:...

 agreed with. Upon further review, von Zittel assigned it to the Coeluridae
Coeluridae
Coeluridae is a historically polyphyletic family of generally small, carnivorous dinosaurs from the late Jurassic Period...

, recognized today as a "wastebasket taxon" for small carnivorous
Carnivore
A carnivore meaning 'meat eater' is an organism that derives its energy and nutrient requirements from a diet consisting mainly or exclusively of animal tissue, whether through predation or scavenging...

 dinosaurs. The carnivore
Carnivore
A carnivore meaning 'meat eater' is an organism that derives its energy and nutrient requirements from a diet consisting mainly or exclusively of animal tissue, whether through predation or scavenging...

 tooth helped this misidentification take hold.

It was ignored for decades until Ralph Molnar
Ralph Molnar
Ralph E. Molnar is a paleontologist who had been Curator of Mammals at the Queensland Museum and more recently associated with the Museum of Northern Arizona. He is also a research associate at the Texas natural Science Centre. He co-authored descriptions of the dinosaurs Muttaburrasaurus, Kakuru,...

 reassessed it. He found that the tooth did not belong to the same animal as the femur and removed it from the type, and suggested that the femur belonged to a hypsilophodont
Hypsilophodont
Hypsilophodonts were small ornithopod dinosaurs, regarded as fast, herbivorous bipeds on the order of 1–2 meters long . They are known from Asia, Australia, Europe, New Zealand, North America, and South America, from rocks of Middle Jurassic to late Cretaceous age...

 or turtle
Turtle
Turtles are reptiles of the order Testudines , characterised by a special bony or cartilaginous shell developed from their ribs that acts as a shield...

. Professional opinion has not changed much since then, although based on size, it appears more likely to be an iguanodont
Iguanodont
Iguanodonts were herbivorous dinosaurs that lived from the mid-Jurassic to Late Cretaceous. Some members include Camptosaurus, Callovosaurus, Iguanodon, Ouranosaurus, and the hadrosaurids or "duck-billed dinosaurs". Iguanodonts were one of the first groups of dinosaurs to be found...

 than a hypsilophodont. Reviews either put it at Ornithopoda incertae sedis
Incertae sedis
, is a term used to define a taxonomic group where its broader relationships are unknown or undefined. Uncertainty at specific taxonomic levels is attributed by , , and similar terms.-Examples:*The fossil plant Paradinandra suecica could not be assigned to any...

or Iguanodontia. Oddly, a semipopular reference reassigned it to Genyodectes
Genyodectes
Genyodectes is a genus of ceratosaurian theropod dinosaur from the Lower Cretaceous of South America. The holotype material was collected from the Cañadón Grande, Departamento Paso de Indios in the Chubut Province of Argentina and consists of an incomplete snout, including the premaxillae,...

without comment, a view which has not been followed since.

Paleobiology

Coria
Rodolfo Coria
Dr. Rodolfo Coria is an Argentine paleontologist.He is best known for having directed the field study and co-naming of the Argentinosaurus in 1993, and the Giganotosaurus , in 1996 among other landmark South American dinosaurs...

 estimates the size of the Loncosaurus type individual at about 5 m (16.4 feet) long. As a small to medium-sized ornithopod, it would have been an agile bipedal herbivore
Herbivore
Herbivores are organisms that are anatomically and physiologically adapted to eat plant-based foods. Herbivory is a form of consumption in which an organism principally eats autotrophs such as plants, algae and photosynthesizing bacteria. More generally, organisms that feed on autotrophs in...

.
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