George's taxonomic arrangement of Dryandra
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Alex George
Alex George
Alexander Segger George is a Western Australian botanist. He is the authority on the plant genera Banksia and Dryandra...

's taxonomic arrangement of Dryandra
Dryandra
Banksia ser. Dryandra is a series of 94 species of shrub to small tree in the plant genus Banksia. It was considered a separate genus named Dryandra until early 2007, when it was merged into Banksia on the basis of extensive molecular and morphological evidence that Banksia was paraphyletic with...

 was the first modern-day arrangement of that taxon. First published in
Nuytsia
Nuytsia (journal)
Nuytsia is a peer-reviewed journal published by the Western Australian Herbarium. It publishes papers on systematic botany, giving preference to papers related to the flora of Western Australia. Nearly twenty percent of Western Australia's plant taxa have been published in Nuytsia. First published...

in 1996, it superseded the arrangement of
Bentham's taxonomic arrangement of Dryandra
George Bentham's taxonomic arrangement of Dryandra was published in 1870, in Volume 5 of Bentham's Flora Australiensis...

 George Bentham
George Bentham
George Bentham CMG FRS was an English botanist, characterized by Duane Isely as "the premier systematic botanist of the nineteenth century".- Formative years :...

, which had stood for over a hundred years; it would later form the basis for George's 1999 treatment of
Dryandra for the Flora of Australia
Flora of Australia (series)
The Flora of Australia is a 59 volume series describing the vascular plants, bryophytes and lichens present in Australia and its external territories...

. In accordance with contemporary thinking, George treated Dryandra as a genus, dividing it into three subgenera, the largest of which was divided into 24 series. The arrangement stood until 2007, when Dryandra was transferred into Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

as B. ser. Dryandra. No alternative has yet been proposed.

Background

The dryandras are a group of 94 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 in the plant family Proteaceae
Proteaceae
Proteaceae is a family of flowering plants distributed in the Southern Hemisphere. The family comprises about 80 genera with about 1600 species. Together with the Platanaceae and Nelumbonaceae they make up the order Proteales. Well known genera include Protea, Banksia, Embothrium, Grevillea,...

. Endemic to south-west Western Australia
Western Australia
Western Australia is a state of Australia, occupying the entire western third of the Australian continent. It is bounded by the Indian Ocean to the north and west, the Great Australian Bight and Indian Ocean to the south, the Northern Territory to the north-east and South Australia to the south-east...

, they grow in forms varying from prostrate shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

s to small trees.
Dryandra was originally published at genus rank in 1810, and was still considered a genus in the 1990s when George published his infrageneric classification.

George's arrangement

George's arrangement of
Dryandra was first published in his 1996 article "New taxa and a new infragenetic classification in Dryandra R.Br. (Proteaceae: Grevilleoideae)", which was published in Nuytsia
Nuytsia (journal)
Nuytsia is a peer-reviewed journal published by the Western Australian Herbarium. It publishes papers on systematic botany, giving preference to papers related to the flora of Western Australia. Nearly twenty percent of Western Australia's plant taxa have been published in Nuytsia. First published...

10(3). This formed the basis for his 1999 treatment of Dryandra for the Flora of Australia
Flora of Australia (series)
The Flora of Australia is a 59 volume series describing the vascular plants, bryophytes and lichens present in Australia and its external territories...

 series, in which he published one new species. In 2005, he published a further six new taxa.

George's arrangement remained current until Austin Mast
Austin Mast
Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently an associate professor within the Department of Biological Science at Florida State University , and has been director of FSU's since August 2003.One of his...

 and Kevin Thiele
Kevin Thiele
Kevin R. Thiele is curator of the Western Australian Herbarium. His research interests include the systematics of the plant families Proteaceae, Rhamnaceae and Violaceae, and the conservation ecology of grassy woodland ecosystems...

 transferred
Dryandra into Banksia in 2007. Mast and Thiele were not then ready to tender a new arrangement, so as an interim measure they transferred Dryandra into Banksia at series
Series (botany)
Series is a low-level taxonomic rank below that of section but above that of species.In botany, a series is a subdivision of a genus...

, thus minimising the nomenclatural disruption of the transfer. Thus George's taxonomic arrangement of
Dryandra was set aside. In effecting the transfer, Mast and Thiele overlooked the six taxa published by George in 2005; these taxa thus retain their original, and now invalid, names.

George's 1999 taxonomic arrangement of
Dryandra, with 2005 amendments, is as follows:
Genus Dryandra (now Banksia ser. Dryandra)
Subgenus Dryandra
Dryandra subg. Dryandra
Dryandra subg. Dryandra is an obsolete series within the former genus Dryandra . It was first published at sectional rank as Dryandra verae in 1830, before being renamed Eudryandra in 1847, the replaced by the autonym at subgenus rank in 1996...

Series Floribundae
Dryandra ser. Floribundae
Dryandra ser. Floribundae is an obsolete series within the former genus Dryandra . It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into...

D. sessilis (now B. sessilis)
D. sessilis var. sessilis (now B. sessilis var. sessilis
Banksia sessilis var. sessilis
Banksia sessilis var. sessilis is the most populous and most widespread variety of Banksia sessilis .-Description:The leaves of this variety are larger than those of B. sessilis var. cygnorum but smaller than those of B. sessilis var. cordata. It differs from B. sessilis var...

)
D. sessilis var. flabellifolia (now B. sessilis var. flabellifolia
Banksia sessilis var. flabellifolia
Banksia sessilis var. flabellifolia is a variety of Banksia sessilis .-Description:It grows as a shrub up to five metres high. Unlike other varieties of B. sessilis, it has hairless stems. The leaves are fan shaped, with a long, toothless lower margin, and a toothed end...

)
D. sessilis var. cordata (now B. sessilis var. cordata
Banksia sessilis var. cordata
Banksia sessilis var. cordata is a variety of Banksia sessilis , with unusually large leaves and flower heads. It is a rare variety that is restricted to the extreme south-west corner of Western Australia.-Description:...

)
D. sessilis var. cygnorum (now B. sessilis var. cygnorum
Banksia sessilis var. cygnorum
Banksia sessilis var. cygnorum is a variety of Banksia sessilis .-Description:It grows as a shrub up to five metres high, with a down of fine hairs on the stems. This variety is has the smallest leaves of any B. sessilis variety, being no more than three centimetres long, and less than two...

)
Series Armatae
Dryandra ser. Armatae
Dryandra ser. Armatae is an obsolete series within the former genus Dryandra . It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.-According...

D. cuneata (now B. obovata
Banksia obovata
Banksia obovata, commonly known as Wedge-leaved Dryandra, is a shrub endemic to Western Australia.It was known as Dryandra cuneata until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. fuscobractea (now B. fuscobractea
Banksia fuscobractea
Banksia fuscobractea is a shrub endemic to Western Australia....

)
D. armata (now B. armata
Banksia armata
Banksia armata, commonly known as Prickly Dryandra, is a shrub endemic to Western Australia.-Description:B. armata grows as a spreading or upright shrub, up to three metres in height. It has deeply serrated leaves. Its inflorescences are usually bright yellow, but may be pink.-Distribution and...

)
D. armata var. armata (now B. armata var. armata
Banksia armata var. armata
Banksia armata var. armata is a variety of shrub endemic to Western Australia.-Description:B. armata var. armata grows as a spreading shrub, up to three metres in height, with deeply serrated leaves and a bright yellow inflorescence.-Distribution and habitat:It occurs throughout much...

)
D. armata var. ignicida (now B. armata var. ignicida
Banksia armata var. ignicida
Banksia armata var. ignicida is a variety of shrub endemic to Western Australia.-Description:B. armata var. ignicida grows as an upright shrub, up to three metres in height, with deeply serrated leaves and yellow or sometimes pink inflorescences...

)
D. prionotes (now B. prionophylla
Banksia prionophylla
Banksia prionophylla is a shrub endemic to Western Australia. Known only from a single population of around 70 plants in a remote part of Western Australia, it is considered rare but not endangered...

D. arborea (now B. arborea
Banksia arborea
Banksia arborea, commonly known as Yilgarn Dryandra, is a plant endemic to Western Australia, notable as it is the only dryandra to grow to tree-like proportions...

)
D. hirsuta (now B. hirta
Banksia hirta
Banksia hirta is a shrub endemic to Western Australia.It was known as Dryandra hirsuta until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. pallida (now B. pallida
Banksia pallida
Banksia pallida is a shrub endemic to Western Australia....

)
D. purdieana (now B. purdieana
Banksia purdieana
Banksia purdieana is a shrub endemic to Western Australia....

)
D. xylothemelia (now B. xylothemelia
Banksia xylothemelia
Banksia xylothemelia is a sprawling woody shrub of the Proteaceae family endemic to southern Western Australia, one of the many species commonly known as dryandras and until recently called Dryandra xylothemelia...

)
D. cirsioides (now B. cirsioides
Banksia cirsioides
Banksia cirsioides is a shrub endemic to Western Australia. It was known as Dryandra cirsioides until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-Description:...

)
D. acanthopoda (now B. acanthopoda
Banksia acanthopoda
Banksia acanthopoda is a species of shrub in the Proteaceae family. It grows as a small spreading shrub to 2 m high and has prickly leaves and yellow composite flower heads, called inflorescences, composed of 50 to 60 individual yellow flowers...

)
D. squarrosa (now B. squarrosa
Banksia squarrosa
Banksia squarrosa, commonly known as Pingle, is a shrub endemic to Western Australia.-Taxonomy:Specimens of B. squarrosa were first collected from near King George Sound in 1829 by William Baxter, and published by Robert Brown as Dryandra squarrosa the following year...

)
D. squarrosa subsp. squarrosa (now B. squarrosa subsp. squarrosa
Banksia squarrosa subsp. squarrosa
Banksia squarrosa subsp. squarrosa is a subspecies of Banksia squarrosa. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra squarrosa subsp. squarrosa until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As with other...

)
D. squarrosa subsp. argillacea (now B. squarrosa subsp. argillacea)
D. hewardiana (now B. hewardiana
Banksia hewardiana
Banksia hewardiana is a shrub endemic to Western Australia. It was first published as Dryandra hewardiana by Carl Meissner in 1856. In 1870, George Bentham published what he held to be a closely related species under the name Dryandra patens, but in 1999 Alex George declared this a synonym of...

)
D. wonganensis (now B. wonganensis
Banksia wonganensis
Banksia wonganensis is a large shrub endemic to Western Australia that, until 2007, was previously known as Dryandra wonganensis. It occurs within a small area in the vicinity of Wongan Hills. It grows on lateritic soils in open woodland or amongst dense shrub...

)
D. trifontinalis (now B. trifontinalis
Banksia trifontinalis
Banksia trifontinalis, commonly known as Three Springs Dryandra, is a shrub endemic to Western Australia.-Description:It grows as an openly branched shrub, and can reach up to two metres high. It has long, narrow, serrated leaves, and a yellow inflorescence.-Taxonomy:The type specimen was collected...

)
D. stricta (now B. strictifolia
Banksia strictifolia
Banksia strictifolia is a shrub endemic to Western Australia.It was known as Dryandra stricta until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele. As the name Banksia stricta had already been published in reference to the plant now known as Pimelea...

)
D. echinata (now B. echinata
Banksia echinata
Banksia echinata is a shrub endemic to Western Australia....

)
D. polycephala (now B. polycephala
Banksia polycephala
Banksia polycephala, commonly known as Many-headed Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra polycephala until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

)
D. subpinnatifida (now B. subpinnatifida
Banksia subpinnatifida
Banksia subpinnatifida is a shrub endemic to Western Australia....

)
D. subpinnatifida var. subpinnatifida (now B. subpinnatifida var. subpinnatifida
Banksia subpinnatifida var. subpinnatifida
Banksia subpinnatifida var. subpinnatifida is a variety of Banksia subpinnatifida. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra subpinnatifida var. subpinnatifida until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia....

)
D. subpinnatifida var. imberbis (now B. subpinnatifida var. imberbis)
D. longifolia (now B. prolata
Banksia prolata
Banksia prolata is a shrub endemic to Western Australia.It was known as Dryandra longifolia until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. longifolia subsp. longifolia (now B. prolata subsp. prolata
Banksia prolata subsp. prolata
Banksia prolata subsp. prolata is a subspecies of Banksia prolata. It was known as Dryandra longifolia subsp. longifolia until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia longifolia had already been used, Mast and Thiele had to choose a new...

)
D. longifolia subsp. calcicola (now B. prolata subsp. calcicola
Banksia prolata subsp. calcicola
Banksia prolata subsp. calcicola is a subspecies of Banksia prolata. It was known as Dryandra longifolia subsp. calcicola until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia longifolia had already been used, Mast and Thiele had to choose a new...

)
D. longifolia subsp. archeos (now B. prolata subsp. archeos
Banksia prolata subsp. archeos
Banksia prolata subsp. archeos is a subspecies of Banksia prolata. It was known as Dryandra longifolia subsp. archeos until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia longifolia had already been used, Mast and Thiele had to choose a new specific...

)
D. borealis (now B. borealis
Banksia borealis
Banksia borealis is a shrub endemic to Western Australia....

)
D. borealis subsp. borealis (now B. borealis subsp. borealis
Banksia borealis subsp. borealis
Banksia borealis subsp. borealis is a subspecies of Banksia borealis. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra borealis subsp. borealis until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As with other members...

)
D. borealis subsp. elatior (now B. borealis subsp. elatior)
Series Marginatae
D. pulchella (now B. bella
Banksia bella
Banksia bella, commonly known as the Silver Wongan Dryandra or Wongan Dryandra, is a shrub endemic to Western Australia. Unusual amongst dryandras, its foliage is glaucous-grey in colour....

)
Series Folliculosae
D. fraseri (now B. fraseri
Banksia fraseri
Banksia fraseri is a shrub endemic to Western Australia. It was known as Dryandra fraseri until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-Description:...

)
D. fraseri var. fraseri (now B. fraseri var. fraseri
Banksia fraseri var. fraseri
Banksia fraseri var. fraseri is a variety of Banksia fraseri. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra fraseri var. fraseri until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As with other members of Banksia ser...

)
D. fraseri var. crebra (now B. fraseri var. crebra)
D. fraseri var. effusa (now B. fraseri var. effusa)
D. fraseri var. ashbyi (now B. fraseri var. ashbyi)
D. fraseri var. oxycedra (now B. fraseri var. oxycedra)
Series Acrodontae
D. sclerophylla (now B. sclerophylla
Banksia sclerophylla
Banksia sclerophylla is a shrub endemic to Western Australia. It was known as Dryandra sclerophylla until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

)
D. kippistiana (now B. kippistiana
Banksia kippistiana
Banksia kippistiana is a shrub endemic to Western Australia. It was known as Dryandra kippistiana until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

)
D. kippistiana var. kippistiana (now B. kippistiana var. kippistiana)
D. kippistiana var. paenepeccata (now B. kippistiana var. paenepeccata)
D. carlinoides (now B. carlinoides
Banksia carlinoides
Banksia carlinoides, commonly known as Pink Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra carlinoides until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

)
D. tridentata (now B. tridentata
Banksia tridentata
Banksia tridentata, commonly known as Yellow Honeypot, is a shrub endemic to Western Australia. It was published as Dryandra tridentata in 1856, and remained in that genus for 150 years, although in 1893 Benjamin Daydon Jackson accidentally listed it in Index Kewensis under Banksia, thus...

)
Series Capitellatae
Dryandra ser. Capitellatae
Dryandra ser. Capitellatae is an obsolete series within the former genus Dryandra . It was published by Alex George in 1996, but discarded in 2007 when Austin Mast and Kevin Thiele sank Dryandra into Banksia.-Publication:George published the series in his 1996 "New taxa and a new infrageneric...

D. serratuloides (now B. serratuloides
Banksia serratuloides
Banksia serratuloides is a shrub endemic to Western Australia. It was known as Dryandra serratuloides until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

)
D. serratuloides subsp. serratuloides (now B. serratuloides subsp. serratuloides
Banksia serratuloides subsp. serratuloides
Banksia serratuloides subsp. serratuloides is a subspecies of Banksia serratuloides. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra serratuloides subsp. serratuloides until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into...

)
D. serratuloides subsp. perissa (now B. serratuloides subsp. perissa)
D. meganotia (now B. meganotia
Banksia meganotia
Banksia meganotia is a shrub endemic to Western Australia.An assessment of the potential impact of climate change on this species found that it was likely to be driven to extinction by loss of habitat by 2080, even under mild climate change scenarios....

)
Series Ilicinae
Dryandra ser. Ilicinae
Dryandra ser. Ilicinae is an obsolete series within the former genus Dryandra . It was first published by Carl Meissner in 1856, but was discarded by George Bentham in 1870...

D. praemorsa (now B. undata
Banksia undata
Banksia undata, commonly known as Urchin Dryandra, is a shrub endemic to Western Australia.It was known as Dryandra praemorsa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. praemorsa var. praemorsa (now B. undata var. undata
Banksia undata var. undata
Banksia undata var. undata is a shrub endemic to Western Australia.It was known as Dryandra praemorsa var. praemorsa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. praemorsa var. splendens (now B. undata var. splendens
Banksia undata var. splendens
Banksia undata var. splendens is a shrub endemic to Western Australia.It was first published by Alex George in 1996, based on a specimen collected by him two years earlier. The name given was Dryandra praemorsa var. splendens, the varietal epithet referring to the "splendid" flower heads...

)
D. quercifolia (now B. heliantha
Banksia heliantha
Banksia heliantha, commonly known as Oak-leaved Dryandra, is a shrub endemic to Western Australia.It was known as Dryandra quercifolia until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. anatona (now B. anatona
Banksia anatona
Banksia anatona is a shrub endemic to Western Australia....

)
Series Dryandra
D. formosa (now B. formosa
Banksia formosa
Banksia formosa, commonly known as Showy Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra formosa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

)
D. nobilis (now B. nobilis
Banksia nobilis
Banksia nobilis, commonly known as Golden Dryandra, Great Dryandra or Kerosene Bush, is a shrub endemic to Western Australia. It occurs on lateritic rises from Eneabba to Katanning in the state's Southwest Botanic Province. With large pinnatifid leaves with triangular lober, and a golden or reddish...

)
D. nobilis subsp. nobilis (now B. nobilis subsp. nobilis
Banksia nobilis subsp. nobilis
Banksia nobilis subsp. nobilis is a subspecies of Banksia nobilis. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra nobilis subsp. nobilis until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As with other members of...

)
D. nobilis subsp. fragrans (now B. nobilis subsp. fragrans)
D. stuposa (now B. stuposa
Banksia stuposa
Banksia stuposa is a shrub endemic to Western Australia. It was known as Dryandra stuposa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-Description:...

)
Series Foliosae
D. mucronulata (now B. mucronulata
Banksia mucronulata
Banksia mucronulata, commonly known as Swordfish Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra mucronulata until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

)
D. mucronulata subsp. mucronulata (now B. mucronulata subsp. mucronulata
Banksia mucronulata subsp. mucronulata
Banksia mucronulata subsp. mucronulata is a subspecies of Banksia mucronulata. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra mucronulata subsp. mucronulata until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As with...

)
D. mucronulata subsp. retrorsa (now B. mucronulata subsp. retrorsa)
D. baxteri (now B. biterax
Banksia biterax
Banksia biterax is a shrub endemic to Western Australia.It was known as Dryandra baxteri until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. foliosissima (now B. foliosissima
Banksia foliosissima
Banksia foliosissima is a shrub endemic to Western Australia....

)
Series Decurrentes
D. comosa (now B. comosa
Banksia comosa
Banksia comosa, commonly known as Wongan Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra comosa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

)
Series Tenuifoliae
D. tenuifolia (now B. tenuis
Banksia tenuis
Banksia tenuis is a shrub endemic to Western Australia. It was known as Dryandra tenuifolia until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

)
D. tenuifolia var. tenuifolia (now B. tenuis var. tenuis
Banksia tenuis var. tenuis
Banksia tenuis var. tenuis is a variety of Banksia tenuis. It was known as Dryandra tenuifolia var. tenuifolia until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia tenuifolia had already been used, Mast and Thiele had to choose a new specific epithet...

)
D. tenuifolia var. reptans (now B. tenuis var. reptans
Banksia tenuis var. reptans
Banksia tenuis var. reptans is a variety of Banksia tenuis. It was known as Dryandra tenuifolia var. reptans until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia tenuifolia had already been used, Mast and Thiele had to choose a new specific epithet...

)
D. obtusa (now B. obtusa
Banksia obtusa
Banksia obtusa, commonly known as Shining Honeypot, is a shrub endemic to Western Australia. It was known as Dryandra obtusa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

)
Series Runcinatae
D. ferruginea (now B. rufa
Banksia rufa
Banksia rufa is a shrub endemic to Western Australia. It was first published by Carl Meisner in 1855, where it was given the name Dryandra ferruginea by Richard Kippist. The following year, Meisner published what was purportedly a distinct species, Dryandra runcinata...

)
D. ferruginea subsp. magma (now B. rufa subsp. magma)
D. ferruginea subsp. tutanningensis (now B. rufa subsp. tutanningensis
Banksia rufa subsp. tutanningensis
Banksia rufa subsp. tutanningensis is a subspecies of Banksia rufa. It was known as Dryandra ferruginea subsp. tutanningensis until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia ferruginea had already been used, Mast and Thiele had to choose a new...

)
D. ferruginea subsp. ferruginea (now B. rufa subsp. rufa
Banksia rufa subsp. rufa
Banksia rufa subsp. rufa is a subspecies of Banksia rufa. It was known as Dryandra ferruginea subsp. ferruginea until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia ferruginea had already been used, Mast and Thiele had to choose a new specific epithet...

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D. ferruginea subsp. pumila (now B. rufa subsp. pumila
Banksia rufa subsp. pumila
Banksia rufa subsp. pumila is a subspecies of Banksia rufa. It was known as Dryandra ferruginea subsp. pumila until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia ferruginea had already been used, Mast and Thiele had to choose a new specific epithet...

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D. ferruginea subsp. obliquiloba (now B. rufa subsp. obliquiloba
Banksia rufa subsp. obliquiloba
Banksia rufa subsp. obliquiloba is a subspecies of Banksia rufa. It was known as Dryandra ferruginea subsp. obliquiloba until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia ferruginea had already been used, Mast and Thiele had to choose a new specific...

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D. ferruginea subsp. chelomacarpa (now B. rufa subsp. chelomacarpa
Banksia rufa subsp. chelomacarpa
Banksia rufa subsp. chelomacarpa is a subspecies of Banksia rufa. It was known as Dryandra ferruginea subsp. chelomacarpa until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia ferruginea had already been used, Mast and Thiele had to choose a new...

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D. ferruginea subsp. flavescens (now B. rufa subsp. flavescens
Banksia rufa subsp. flavescens
Banksia rufa subsp. flavescens is a subspecies of Banksia rufa. It was known as Dryandra ferruginea subsp. flavescens until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia ferruginea had already been used, Mast and Thiele had to choose a new specific...

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D. corvijuga (now B. corvijuga
Banksia corvijuga
Banksia corvijuga is a shrub endemic to Western Australia....

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D. epimicta (now B. epimicta
Banksia epimicta
Banksia epimicta is a shrub endemic to Western Australia....

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D. proteoides (now B. proteoides
Banksia proteoides
Banksia proteoides, commonly known as King Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra proteoides until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

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Series Triangulares
D. drummondii (now B. drummondii
Banksia drummondii
Banksia drummondii, commonly known as Drummond's Dryandra, is a shrub endemic to Western Australia. Depending on the subspecies, flowering occurs either mid-year, or at year's end.-Description:...

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D. drummondii subsp. drummondii (now B. drummondii subsp. drummondii)
D. drummondii subsp. hiemalis (now B. drummondii subsp. hiemalis)
D. drummondii subsp. macrorufa (now B. drummondii subsp. macrorufa)
D. octotriginta (now B. octotriginta
Banksia octotriginta
Banksia octotriginta is a shrub endemic to Western Australia.An assessment of the potential impact of climate change on this species found that it was likely to be driven to extinction by loss of habitat by 2080, even under mild climate change scenarios....

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D. catoglypta (now B. catoglypta
Banksia catoglypta
Banksia catoglypta is a shrub endemic to Western Australia....

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Series Aphragma
Dryandra ser. Aphragma
Dryandra ser. Aphragma is an obsolete series within the former genus Dryandra . It was first published at sectional rank by Robert Brown in 1830, and was retained at that rank until 1999, when Alex George demoted it to a series...

D. pteridifolia (now B. pteridifolia
Banksia pteridifolia
Banksia pteridifolia, commonly known as Tangled Honeypot, is a shrub endemic to Western Australia. It was known as Dryandra pteridifolia until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

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D. pteridifolia subsp. inretita (now B. pteridifolia subsp. inretita
Banksia pteridifolia subsp. pteridifolia
Banksia pteridifolia subsp. pteridifolia is a subspecies of Banksia pteridifolia. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra pteridifolia subsp. pteridifolia until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As...

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D. pteridifolia subsp. pteridifolia (now B. pteridifolia subsp. pteridifolia
Banksia pteridifolia subsp. pteridifolia
Banksia pteridifolia subsp. pteridifolia is a subspecies of Banksia pteridifolia. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra pteridifolia subsp. pteridifolia until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As...

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D. pteridifolia subsp. vernalis (now B. pteridifolia subsp. vernalis)
D. fililoba (now B. fililoba
Banksia fililoba
Banksia fililoba is a shrub endemic to Western Australia....

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D. shanklandiorum (now B. shanklandiorum
Banksia shanklandiorum
Banksia shanklandiorum is a shrub endemic to Western Australia....

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D. nervosa (now B. alliacea
Banksia alliacea
Banksia alliacea is a shrub endemic to Western Australia.It was known as Dryandra nervosa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

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D. blechnifolia (now B. pellaeifolia
Banksia pellaeifolia
Banksia pellaeifolia is a shrub endemic to Western Australia.It was known as Dryandra blechnifolia until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

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D. porrecta (now B. porrecta
Banksia porrecta
Banksia porrecta is a shrub endemic to Western Australia.An assessment of the potential impact of climate change on this species found that its range is likely to contract by between 50% and 80% by 2080, depending on the severity of the change....

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D. aurantia (now B. aurantia
Banksia aurantia
Banksia aurantia is a shrub endemic to Western Australia....

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D. calophylla (now B. calophylla
Banksia calophylla
Banksia calophylla is a shrub endemic to Western Australia. It was known as Dryandra calophylla until 2007, when all Dryandra species were transferred to Banksia by research botanists Austin Mast and Kevin Thiele.-External links:...

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D. lepidorhiza (now B. lepidorhiza
Banksia lepidorhiza
Banksia lepidorhiza is a shrub endemic to Western Australia....

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Series Ionthocarpae
D. ionthocarpa (now B. ionthocarpa
Banksia ionthocarpa
Banksia ionthocarpa is a shrub endemic to Western Australia. It was known as Dryandra ionthocarpa until 2007.-Taxonomy:B. ionthocarpa was first discovered in 1987 by professional seed supplier Peter Luscombe. The following year specimens were collected by both Margaret Pieroni and Alex George...

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D. ionthocarpa subsp. ionthocarpa (now B. ionthocarpa subsp. ionthocarpa
Banksia ionthocarpa subsp. ionthocarpa
Banksia ionthocarpa subsp. ionthocarpa is a subspecies of Banksia ionthocarpa. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra ionthocarpa subsp. ionthocarpa until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As with...

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D. ionthocarpa subsp. chrysophoenix (now B. ionthocarpa subsp. chrysophoenix)
Series Inusitatae
D. idiogenes (now B. idiogenes
Banksia idiogenes
Banksia idiogenes is a shrub endemic to Western Australia. It was known as Dryandra idiogenes until 2007.-Taxonomy:B. idiogenes was first discovered by Alex George in 1986, and published by him in 1996 as Dryandra idiogenes, the specific epithet coming from the Greek idiogenes , in reference...

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Series Subulatae
D. subulata (now B. subulata
Banksia subulata
Banksia subulata, commonly known as Awled Honeypot, is a shrub endemic to Western Australia.-Taxonomy:It was first published as Dryandra subulata by Charles Gardner in 1964. In Alex George's 1996 arrangement, this species was recognised as quite distinctive, being unusual in its long, unserrated...

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Series Gymnocephalae
D. cynaroides (now B. cynaroides
Banksia cynaroides
Banksia cynaroides is a shrub endemic to Western Australia.An assessment of the potential impact of climate change on this species found that it was likely to be driven to extinction by loss of habitat by 2080, even under mild climate change scenarios....

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D. erythrocephala (now B. erythrocephala
Banksia erythrocephala
Banksia erythrocephala is a shrub endemic to Western Australia.An assessment of the potential impact of climate change on this species found that it was likely to be driven to extinction by loss of habitat by 2080, even under mild climate change scenarios....

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D. erythrocephala var. erythrocephala (now B. erythrocephala var. erythrocephala
Banksia erythrocephala var. erythrocephala
Banksia erythrocephala var. erythrocephala is a variety of Banksia erythrocephala. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra erythrocephala var. erythrocephala until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia....

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D. erythrocephala var. inopinata (now B. erythrocephala var. inopinata)
D. horrida (now B. horrida
Banksia horrida
Banksia horrida, commonly known as Prickly Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra horrida until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

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D. vestita (now B. vestita
Banksia vestita
Banksia vestita, commonly known as Summer Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra vestita until 2007.-Description:...

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D. viscida (now B. viscida
Banksia viscida
Banksia viscida, commonly known as Sticky Dryandra, is a small shrub known only from four laterite hills in semi-arid inland Western Australia. Known until recently as Dryandra viscida, it is thought to be rare but not threatened....

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D. mimica (now B. mimica
Banksia mimica
Banksia mimica, commonly known as Summer Honeypot, is a shrub endemic to Western Australia. Until 2007 it was known as Dryandra mimica; in 2007 all Dryandra species were transferred into Banksia....

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D. speciosa (now B. splendida
Banksia splendida
Banksia splendida, commonly known as Shaggy Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra speciosa until 2007-Taxonomy:...

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D. speciosa subsp. speciosa (now B. speciosa subsp. splendida
Banksia splendida subsp. splendida
Banksia splendida subsp. splendida is a subspecies of Banksia splendida. It was known as Dryandra speciosa subsp. speciosa until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia speciosa had already been used, Mast and Thiele had to choose a new...

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D. speciosa subsp. macrocarpa (now B. speciosa subsp. macrocarpa
Banksia splendida subsp. macrocarpa
Banksia splendida subsp. macrocarpa is a subspecies of Banksia splendida. It was known as Dryandra speciosa subsp. macrocarpa until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a species named Banksia speciosa, Mast and Thiele had to choose a new...

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D. shuttleworthiana (now B. shuttleworthiana
Banksia shuttleworthiana
Banksia shuttleworthiana, commonly known as Bearded Dryandra, is a shrub endemic to Western Australia, and is found between the towns of Geraldton and Gingin. It was known as Dryandra shuttleworthiana until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin...

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Series Plumosae
Dryandra ser. Plumosae
Dryandra ser. Plumosae is an obsolete series within the former genus Dryandra . It was published by Alex George in 1996, but discarded in 2007 when Austin Mast and Kevin Thiele sank Dryandra into Banksia....

D. plumosa (now B. plumosa
Banksia plumosa
Banksia plumosa is a shrub endemic to Western Australia. It was known as Dryandra plumosa until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

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D. plumosa subsp. plumosa (now B. plumosa subsp. plumosa
Banksia plumosa subsp. plumosa
Banksia plumosa subsp. plumosa is a subspecies of Banksia plumosa. As an autonym, it is defined as encompassing the type material of the species. It was known as Dryandra plumosa subsp. plumosa until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. As with other members of...

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D. plumosa subsp. denticulata (now B. plumosa subsp. denticulata)
D. pseudoplumosa (now B. pseudoplumosa
Banksia pseudoplumosa
Banksia pseudoplumosa is a shrub endemic to Western Australia....

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D. montana (now B. montana
Banksia montana
Banksia montana is a shrub endemic to Western Australia....

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Series Concinnae
D. concinna (now B. concinna
Banksia concinna
Banksia concinna is a shrub endemic to Western Australia. It was known as Dryandra concinna until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

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D. serra (now B. serra
Banksia serra
Banksia serra, commonly known as Serrate-leaved Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra polycephala until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.An assessment of the potential impact of climate change on this...

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D. foliolata (now B. foliolata
Banksia foliolata
Banksia foliolata is a shrub endemic to Western Australia. It was known as Dryandra foliolata until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

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Series Obvallatae
D. fasciculata (now B. fasciculata
Banksia fasciculata
Banksia fasciculata is a shrub endemic to Western Australia....

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D. conferta (now B. densa
Banksia densa
Banksia densa is a shrub endemic to Western Australia. It was known as Dryandra conferta until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

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D. conferta var. conferta (now B. densa var. densa
Banksia densa var. densa
Banksia densa var. densa is a variety of Banksia densa. It was known as Dryandra conferta var. conferta until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia conferta had already been used, Mast and Thiele had to choose a new specific epithet for D....

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D. conferta var. parva (now B. densa var. parva
Banksia densa var. parva
Banksia densa var. parva is a variety of Banksia densa. It was known as Dryandra conferta var. parva until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia conferta, Mast and Thiele had to choose a new specific epithet for D....

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D. columnaris (now B. columnaris
Banksia columnaris
Banksia columnaris is a shrub endemic to Western Australia....

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D. platycarpa (now B. platycarpa
Banksia platycarpa
Banksia platycarpa is a shrub endemic to Western Australia....

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D. seneciifolia (now B. seneciifolia
Banksia seneciifolia
Banksia seneciifolia is a shrub endemic to Western Australia. It was known as Dryandra polycephala until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele.-External links:...

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D. rufistylis (now B. rufistylis
Banksia rufistylis
Banksia rufistylis is a shrub endemic to Western Australia....

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D. insulanemorecincta (now B. insulanemorecincta
Banksia insulanemorecincta
Banksia insulanemorecincta is a shrub endemic to Western Australia....

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Series Pectinatae
D. nana (now B. nana
Banksia nana
Banksia nana, commonly known as Dwarf Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra nana until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

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Series Acuminatae
D. preissii (now B. acuminata
Banksia acuminata
Banksia acuminata is a rare prostrate shrub endemic to south-west Western Australia. It was published in 1848 as Dryandra preissii, but transferred into Banksia as B. acuminata in 2007.-Description:...

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Series Niveae
Dryandra ser. Niveae
Dryandra ser. Niveae is an obsolete series within the former genus Dryandra . It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.-According to...

D. arctotidis (now B. arctotidis
Banksia arctotidis
Banksia arctotidis is a shrub endemic to Western Australia. It was known as Dryandra arctotidis until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

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D. tortifolia (now B. tortifolia
Banksia tortifolia
Banksia tortifolia is a shrub endemic to Western Australia. It was first published as Dryandra tortifolia in 1855. In 1870, George Bentham demoted it to a variety of Dryandra arctotidis , but this was overturned by Alex George in 1999...

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D. stenoprion (now B. stenoprion
Banksia stenoprion
Banksia stenoprion is a shrub endemic to Western Australia....

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D. cypholoba (now B. cypholoba
Banksia cypholoba
Banksia cypholoba is a shrub endemic to Western Australia....

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D. lindleyana (now B. dallanneyi
Banksia dallanneyi
Banksia dallanneyi, commonly known as Couch Honeypot, is a prostrate shrub endemic to Western Australia. It grows to a height and width of up to 0.5 m, although it is very common for many plants to be growing side by side; thus the clump may be several metres across.It was known as Dryandra...

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D. lindleyana subsp. lindleyana (now B. dallanneyi subsp. dallanneyi
Banksia dallanneyi subsp. dallanneyi
Banksia dallanneyi subsp. dallanneyi is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. lindleyana until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia lindleyana had already been used, Mast and Thiele had to choose a...

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D. lindleyana subsp. pollosta (now B. dallanneyi subsp. pollosta
Banksia dallanneyi subsp. pollosta
Banksia dallanneyi subsp. pollosta is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. pollosta until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia lindleyana, Mast and Thiele had to choose a new...

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D. lindleyana subsp. media (now B. dallanneyi subsp. media
Banksia dallanneyi subsp. media
Banksia dallanneyi subsp. media is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. media until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia lindleyana, Mast and Thiele had to choose a new...

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D. lindleyana subsp. agricola (now B. dallanneyi subsp. agricola
Banksia dallanneyi subsp. agricola
Banksia dallanneyi subsp. agricola is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. agricola until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia lindleyana, Mast and Thiele had to choose a new...

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D. lindleyana subsp. sylvestris (now B. dallanneyi subsp. sylvestris
Banksia dallanneyi subsp. sylvestris
Banksia dallanneyi subsp. sylvestris is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. sylvestris until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia lindleyana, Mast and Thiele had to choose a...

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D. brownii (now B. brunnea
Banksia brunnea
Banksia brunnea is a shrub endemic to Western Australia.It was known as Dryandra brownii until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele...

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D. nivea (now B. nivea
Banksia nivea
Banksia nivea, commonly known as Honeypot Dryandra, is a shrub endemic to Western Australia. First described as Banksia nivea, it was transferred to Dryandra as Dryandra nivea by Robert Brown in 1810, and remained in that genus until 2007, when all Dryandra species were transferred to Banksia by...

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D. nivea subsp. nivea (now B. nivea subsp. nivea)
D. nivea subsp. uliginosa (now B. nivea subsp. uliginosa)
Subgenus Hemiclidia
Dryandra subg. Hemiclidia
Dryandra subg. Hemiclidia is an obsolete plant taxon that encompassed material that is now included in Banksia. Published at genus rank as Hemiclidia by Robert Brown in 1830, it was set aside by George Bentham in 1870, but reinstated at subgenus rank by Alex George in 1996...

D. falcata (now B. falcata
Banksia falcata
Banksia falcata, commonly known as Prickly Dryandra, is a shrub endemic to Western Australia. It was known as Dryandra falcata until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele....

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D. glauca (now B. glaucifolia
Banksia glaucifolia
Banksia glaucifolia is a shrub endemic to Western Australia.It was known as Dryandra glauca until 2007, when all Dryandra species were transferred to Banksia by Austin Mast and Kevin Thiele. The name "Banksia glauca" had already been published in reference to Banksia integrifolia subsp...

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Subgenus Diplophragma
Dryandra subg. Diplophragma
Dryandra subg. Diplophragma is an obsolete subgenus within the former genus Dryandra . It was first published by Robert Brown in 1830, but was discarded by George Bentham in 1870...

D. bipinnatifida (now B. bipinnatifida
Banksia bipinnatifida
Banksia bipinnatifida is a shrub endemic to Western Australia.-Description:It grows as a prostrate shrub with an underground stem and a lignotuber. It has few leaves; these are deeply lobes, and the lobes are themselves deeply lobed, giving the impression of a bipinnate leaf structure...

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See also

  • Brown's taxonomic arrangement of Dryandra
    Brown's taxonomic arrangement of Dryandra
    Robert Brown's taxonomic arrangement of Dryandra was the first arrangement of what is now Banksia ser. Dryandra. His initial arrangement was published in 1810, and a further arrangement, including an infrageneric classification, followed in 1830...

  • Meissner's taxonomic arrangement of Dryandra
    Meissner's taxonomic arrangement of Dryandra
    Carl Meissner's taxonomic arrangement of Dryandra, now Banksia ser. Dryandra, was published in 1856 as part of his chapter on the Proteaceae in A. P. de Candolle's Prodromus systematis naturalis regni vegetabilis...

  • Bentham's taxonomic arrangement of Dryandra
    Bentham's taxonomic arrangement of Dryandra
    George Bentham's taxonomic arrangement of Dryandra was published in 1870, in Volume 5 of Bentham's Flora Australiensis...

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