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Embryo

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Embryo



 
 
An embryo (irregularly from Greek
Greek language

Greek is an Indo-European languages native to the southern Balkan peninsula, the language of the Greek people. It forms an independent branch within Indo-European....
: , plural , lit. "that which grows," from en- "in" + bryein "to swell, be full"; the proper Latinate form would be embryum) is a multicellular
Multicellular organism

Multicellular organisms are organisms consisting of more than one cell , and having differentiated cells that perform specialized functions in the cell....
 diploid
Ploidy

Ploidy is the number of complete sets of non-homologous chromosomes in a biological cell. In humans, the somatic cells that comprise the body are diploid , but sex cells are haploid....
 eukaryote
Eukaryote

Animals, plants, fungus, and protists are eukaryotes , organisms whose Cell are organized into complex structures enclosed within Cell membrane....
 in its earliest stage of development
Development

Development may refer to:...
, from the time of first cell division
Cell division

Cell division is a process by which a cell , called the parent cell, divides into two or more cells, called daughter cells. Cell division is usually a small segment of a larger cell cycle....
 until birth
Birth

Birth is the act or process of bearing or bringing forth offspring . The offspring is brought forth from the mother. Different forms of birth are oviparity, vivipary or Ovoviviparity....
, hatching
Egg (biology)

In most birds and reptiles, an egg is the zygote, resulting from fertilization of the ovum. To enable incubation the egg is usually kept within a favourable temperature range as it nourishes and protects the growing embryo....
, or germination
Germination

Germination is the process whereby growth emerges from a period of dormancy. The most common example of germination is the sprouting of a seedling from a seed of an flowering plant or gymnosperm....
.






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Wrinkledfrog Embryos
An embryo (irregularly from Greek
Greek language

Greek is an Indo-European languages native to the southern Balkan peninsula, the language of the Greek people. It forms an independent branch within Indo-European....
: , plural , lit. "that which grows," from en- "in" + bryein "to swell, be full"; the proper Latinate form would be embryum) is a multicellular
Multicellular organism

Multicellular organisms are organisms consisting of more than one cell , and having differentiated cells that perform specialized functions in the cell....
 diploid
Ploidy

Ploidy is the number of complete sets of non-homologous chromosomes in a biological cell. In humans, the somatic cells that comprise the body are diploid , but sex cells are haploid....
 eukaryote
Eukaryote

Animals, plants, fungus, and protists are eukaryotes , organisms whose Cell are organized into complex structures enclosed within Cell membrane....
 in its earliest stage of development
Development

Development may refer to:...
, from the time of first cell division
Cell division

Cell division is a process by which a cell , called the parent cell, divides into two or more cells, called daughter cells. Cell division is usually a small segment of a larger cell cycle....
 until birth
Birth

Birth is the act or process of bearing or bringing forth offspring . The offspring is brought forth from the mother. Different forms of birth are oviparity, vivipary or Ovoviviparity....
, hatching
Egg (biology)

In most birds and reptiles, an egg is the zygote, resulting from fertilization of the ovum. To enable incubation the egg is usually kept within a favourable temperature range as it nourishes and protects the growing embryo....
, or germination
Germination

Germination is the process whereby growth emerges from a period of dormancy. The most common example of germination is the sprouting of a seedling from a seed of an flowering plant or gymnosperm....
. In humans, it is called an embryo from the moment of implantation until the end of the 8th week, whereafter it is instead called a fetus
Fetus

A fetus is a developing mammal or other viviparous vertebrate, after the embryonic stage and before childbirth. The plural is fetuses, or sometimes feti....
.

Development


The development of the embryo is called embryogenesis. In organisms that reproduce sexually
Sex

In biology, sex is a process of combining and mixing genetics traits, often resulting in the specialization of organisms into male and female types ....
, once a sperm
Spermatid

The term spermatid refers to the haploid male gametid that results from division of secondary spermatocytes. As a result of meiosis, each spermatid contains only half of the genetic material present in the original primary spermatocyte....
 fertilizes an egg cell
Ovum

An ovum is a haploid female reproductive cell or gamete. Both animals and embryophytes have ova. The term ovule is used for the young ovum of an animal, as well as the plant structure that carries the female gametophyte and egg cell and develops into a seed after fertilization....
, the result is a cell
Cell (biology)

The cell is the structural and functional unit of all known Life organisms. It is the smallest unit of an organism that is classified as living, and is often called the building bricks of life....
 called the zygote
Zygote

A zygote is a cell that is the result of fertilization. That is, two ploidy cells—usually an ovum from a female and a sperm cell from a male—merge into a single ploidy cell called the zygote ....
 that has half of the DNA
DNA

Deoxyribonucleic acid is a nucleic acid that contains the genetics instructions used in the development and functioning of all known living organisms and some viruses....
 of each of two parents. The resulting embryo derives 50 percent of its genetic makeup from each parent. In plant
Plant

Plants are Life organisms belonging to the Kingdom Plantae. They include familiar organisms such as trees, herbs, bushes, grasses, vines, ferns, mosses, and green algae....
s, animal
Animal

Animals are a major group of multicellular, eukaryotic organisms of the Kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life....
s, and some protist
Protist

Protists ; eukaryote microorganisms. Historically, protists were treated as the kingdom Protista but this group is no longer recognized in modern taxonomy....
s, the zygote will begin to divide by mitosis
Mitosis

Mitosis is the process in which a eukaryotic cell separates the chromosomes in its cell nucleus, into two identical sets in two daughter nuclei....
 to produce a multicellular organism. The result of this process is an embryo.

In animals

In animals, the development of the zygote into an embryo proceeds through specific recognizable stages of blastula
Blastula

The blastula is an early stage of embryonic development in animals. It is also called blastosphere. It is produced by cleavage of a fertilized ovum and consists of a spherical layer of around 128 cells surrounding a central fluid-filled cavity called the blastocoel....
, gastrula, and organogenesis
Organogenesis

In embryogenesis, organogenesis is the process by which the ectoderm, endoderm, and mesoderm develop into the internal organs of the organism. Internal organs initiate development in humans within the 3rd to 8th weeks in utero.The germ layers in organogenesis differ by three processes: folds, splits, and condensation....
. The blastula stage typically features a fluid-filled cavity, the blastocoel
Blastocoele

A blastocoel or blastocele is the fluid-filled central region of a blastocyst. A blastocoele forms during embryogenesis when a zygote divides into many cells through mitosis....
, surrounded by a sphere or sheet of cells, also called blastomere
Blastomere

A blastomere is a type of Cell produced by division of the ovum after Human fertilization....
s.

During gastrulation
Gastrulation

Gastrulation is a phase early in the development of animal embryos, during which the morphology of the embryo is dramatically restructured by cell migration....
 the cells of the blastula undergo coordinated processes of cell division, invasion, and/or migration to form two (diploblastic
Diploblasty

Diploblasty is a condition of the ovum in which there are two primary germ layers: the ectoderm and endoderm.Diploblastic organisms are organisms which evolve from such an ovum, and include cnidaria and ctenophores....
) or three (triploblastic
Triploblasty

Triploblasty is a condition of the blastula in which there are three primary germ layers: the ectoderm, mesoderm, and endoderm. The germ layers form during gastrulation of the blastula....
) tissue layers. In triploblastic organisms, the three germ layer
Germ layer

A germ layer is a group of cell s, formed during animal embryogenesis. Germ layers are particularly pronounced in the vertebrates; however, all animals more complex than sea sponge produce two or three primary tissue layers ....
s are called endoderm
Endoderm

Endoderm, is one of the germ layers formed during animal embryogenesis. Cells migrating inward along the archenteron from the inner layer of the gastrula, which develops into the endoderm....
, ectoderm
Ectoderm

The ectoderm is the start of a tissue that covers the body surfaces. It emerges first and forms from the outermost of the germ layers.Generally speaking, the ectoderm differentiates to form the nervous system, Epidermis , and the outer part of integumentary system....
 and mesoderm
Mesoderm

One of the three germ layers found in the embryos of animals more complex than cnidarians, making them triploblastic. Mesoderm forms in the embryo during gastrulation when some of the cells migrating inward to form the endoderm, produce an additional layer that lies between the endoderm and the ectoderm....
. However, the position and arrangement of the germ layers are highly species-specific, depending on the type of embryo produced. In vertebrate
Vertebrate

Vertebrates are members of the subphylum Vertebrata, chordates with Vertebras or Vertebral columns. The grouping sometimes includes the hagfish, which have no vertebrae, but are genetically quite closely related to lampreys, which do have vertebrae....
s, a special population of embryonic cells called the neural crest
Neural crest

The neural crest, a transient component of the ectoderm, is located in between the neural tube and the epidermis of an embryo during neural tube formation....
 has been proposed as a "fourth germ layer", and is thought to have been an important novelty in the evolution of head structures.

During organogenesis, molecular and cellular interactions between germ layers, combined with the cells' developmental potential or competence
Competence (biology)

In microbiology, genetics, cell biology and molecular biology, competence is the ability of a cell to take up extracellular DNA from its environment....
 to respond, prompt the further differentiation of organ-specific cell types. For example, in neurogenesis
Neurogenesis

Neurogenesis is the process by which neurons are created. Most active during pre-natal development, neurogenesis is responsible for populating the growing brain....
, a subpopulation of ectoderm cells is set aside to become the brain
Brain

The brain is the center of the nervous system in all vertebrate, and most invertebrate, animals. Some primitive animals such as cnidarian and echinoderm have a decentralized nervous system without a brain, while sponges lack any nervous system at all....
, spinal cord
Spinal cord

The spinal cord is a long, thin, tubular bundle of neuron and glia that extends from the brain. The brain and spinal cord together make up the central nervous system....
 and peripheral nerves
Peripheral nervous system

The peripheral nervous system resides or extends outside the central nervous system , which consists of the brain and spinal cord. The main function of the PNS is to connect the CNS to the limbs and organs....
. Modern developmental biology is extensively probing the molecular basis for every type of organogenesis, including angiogenesis
Angiogenesis

Angiogenesis is a physiological process involving the growth of new blood vessels from pre-existing vessels. Though there has been some debate over this, vasculogenesis is the term used for spontaneous blood-vessel formation, and Intussusception is the term for new blood vessel formation by splitting off existing ones....
 (formation of new blood vessels from pre-existing ones), chondrogenesis
Chondrogenesis

Chondrogenesis is the process by which cartilage is developed....
 (cartilage), myogenesis
Myogenesis

Myogenesis is the formation of muscular tissue, in particular during embryonic development.Muscle fibers form from the fusion of myoblasts into multi-nucleated fibers ....
 (muscle), osteogenesis
Osteogenesis

Osteogenesis is the process of laying down new bone material by osteoblasts. It occurs in two different ways: Either by Intramembranous Osteogenesis , which is the direct laying down of bone into the primitive connective tissue , or by Endochondral Osteogenesis which involves a cartilage precursor....
 (bone), and many others.

Generally, if a structure pre-dates another structure in evolutionary terms, then it often appears earlier than the other in an embryo; this general observation is sometimes summarized by the phrase "ontogeny recapitulates phylogeny
Recapitulation theory

The theory of recapitulation, also called the biogenetic law or embryological parallelism, and often expressed as ontogeny recapitulates phylogeny, was put forward by ?tienne Serres in 1824?26 as what became known as the "Meckel-Serres Law" which attempted to provide a link between comparative embryology and a "pattern of un...
." For example, the backbone is a common structure among all vertebrates such as fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
, reptile
Reptile

Reptiles, or members of the class Reptilia, are air-breathing, cold-blooded vertebrates that have skin covered in scale as opposed to hair or feathers....
s and mammal
Mammal

Mammals are a class of vertebrate animals whose name is derived from their distinctive feature, mammary glands, with which they feed their young....
s, and the backbone also appears as one of the earliest structures laid out in all vertebrate embryos. The cerebrum in humans, which is the most sophisticated part of the brain
Human brain

The human brain is the center of the human nervous system and is a highly complex organ. It has the same general structure as the brains of other mammals, but is over five times as large as the "average brain" of a mammal with the same body size....
, develops last. This rule is not absolute, but it is recognized as being partly applicable to development of the human embryo.

Embryos of plants and animals

Ginkgo Embryo and Gametophyte
*Plants: In botany
Botany

Botany, plant science, phytology, or plant biology is a branch of biology and is the Scientific method of plant life and development....
, a seed plant embryo is part of a seed
Seed

A seed is a small Plant embryogenesis plant enclosed in a covering called the seed coat, usually with some Food storage. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant....
, consisting of precursor tissues for the leaves, stem (see hypocotyl
Hypocotyl

Hypocotyl is a Botany term for a part of a germination seedling of a seed plant. As the plant embryo grows at germination, it sends out a shoot called a radicle that becomes the primary root and penetrates down into the soil....
), and root (see radicle
Radicle

In botany, the radicle is the first part of a seedling to emerge from the seed during the process of germination. The radicle is the embryonic root of the plant, and grows downward in the soil....
), as well as one or more cotyledon
Cotyledon

A cotyledon is a significant part of the embryo within the seed of a plant. Upon germination, the cotyledon may become the embryonic first leaf of a seedling....
s
. Once the embryo begins to germinate
Germination

Germination is the process whereby growth emerges from a period of dormancy. The most common example of germination is the sprouting of a seedling from a seed of an flowering plant or gymnosperm....
 — grow out from the seed — it is called a seedling. Plants that do not produce seeds, but do produce an embryo, include the bryophyte
Bryophyte

Bryophytes are all embryophytes that are non-vascular plant: they have tissues and enclosed reproductive systems, but they lack vascular tissue that circulates liquids....
s and fern
Fern

A fern is any one of a group of about 20,000 species of plants classified in the phylum or division Pteridophyta, also known as Filicophyta....
s. In these plants, the embryo is a young plant that grows attached to a parental gametophyte
Gametophyte

In plants and algae that undergo alternation of generations, a gametophyte is the multicellular structure, or phase, that is haploid, containing a single set of chromosomes:...
.
  • Animals: The embryo of a placental mammal is defined as the organism between the first division
    Cell division

    Cell division is a process by which a cell , called the parent cell, divides into two or more cells, called daughter cells. Cell division is usually a small segment of a larger cell cycle....
     of the zygote
    Zygote

    A zygote is a cell that is the result of fertilization. That is, two ploidy cells—usually an ovum from a female and a sperm cell from a male—merge into a single ploidy cell called the zygote ....
    (a fertilized ovum
    Ovum

    An ovum is a haploid female reproductive cell or gamete. Both animals and embryophytes have ova. The term ovule is used for the young ovum of an animal, as well as the plant structure that carries the female gametophyte and egg cell and develops into a seed after fertilization....
    ) until it becomes a fetus
    Fetus

    A fetus is a developing mammal or other viviparous vertebrate, after the embryonic stage and before childbirth. The plural is fetuses, or sometimes feti....
    . In humans, the embryo is defined as the product of conception from implantation in the uterus
    Uterus

    The uterus is a major female hormone-responsive reproductive sex organ of most mammals, including humans. It is within the uterus that the fetus develops during gestation....
     through the eighth week of development. An embryo is called a fetus at a more advanced stage of development and up until birth
    Childbirth

    Childbirth is the culmination of a human pregnancy or gestation period with the delivery of one or more newborn infants from a woman's uterus. The process of normal human childbirth is categorized in three stages of labour: the shortening and dilation of the cervix, descent and delivery of the infant, and delivery of the placenta.....
     or hatching. In humans, this is from the eighth week of gestation
    Gestation

    Gestation is the carrying of an embryo or fetus inside a female viviparous animal. Mammals during mammalian pregnancy can have one or more gestations at the same time ....
    .

Fossil record

Fossilised embryos are known from the Precambrian, and are found in great number during the Cambrian period.

The human embryo


Growth

Week 1-3 5-7 days after fertilization, the blastula attaches to the wall of the uterus (endometrium
Endometrium

The endometrium is the inner membrane of the mammalian uterus....
). When it comes into contact with the endometrium it performs implantation
Implantation

Implantation is an event that occurs early in pregnancy in which the embryo adheres to the wall of uterus. At this stage of prenatal development, the embryo is a blastocyst....
. Implantation connections between the mother and the embryo will begin to form, including the umbilical cord
Umbilical cord

In placental mammals, the umbilical cord is the connecting cord from the developing embryo or fetus to the placenta. During prenatal development, the umbilical cord comes from the same zygote as the fetus and normally contains two arteries and one vein , buried within Wharton's jelly....
. The embryo's growth centers around an axis, which will become the spine and spinal cord. The brain, spinal cord, heart, and gastrointestinal tract begin to form.

Week 4-5 Chemicals produced by the embryo stop the woman's menstrual cycle
Menstrual cycle

The menstrual cycle is a recurring cycle of physiology changes that occurs in reproductive-age females. Overt menstruation occurs primarily in humans and close evolutionary relatives such as chimpanzees....
. Neurogenesis is underway, showing brain activity at about the 6th week. The heart will begin to beat around the same time. Limb buds appear where the arms and legs will grow later. Organogenesis begins. The head represents about one half of the embryo's axial length, and more than half of the embryo's mass
Mass

In physical science, mass refers to the degree of acceleration a body acquires when subject to a force: bodies with greater mass are accelerated less by the same force....
. The brain develops into five areas. Tissue formation occurs that develops into the vertebra and some other bones. The heart starts to beat and blood starts to flow.

Week 6-8 Myogenesis and neurogenesis have progressed to where the embryo is capable of motion, and the eyes begin to form. Organogenesis and growth continue. Hair has started to form along with all essential organs. Facial features are beginning to develop. At the end of the 8th week, the embryonic stage is over, and the fetal
Fetus

A fetus is a developing mammal or other viviparous vertebrate, after the embryonic stage and before childbirth. The plural is fetuses, or sometimes feti....
 stage begins.

Bioethical controversies

There are a number of controversial biomedical operations and phenomenae such as selective reduction
Selective reduction

Selective reduction is the practice of reducing the number of fetuses in a multifetal pregnancy . With selective reduction, one or more fetuses are "selected" for termination....
, prenatal diagnosis
Prenatal diagnosis

Prenatal testing is testing for diseases or conditions in a fetus or embryo before it is born. The aim is to detect birth defects such as neural tube defects, Down syndrome, chromosome abnormalities, genetic diseases and other conditions....
, preimplantation diagnosis
Preimplantation diagnosis

Preimplantation diagnosis is the act of checking an ovum, zygote, or blastocyst for defects or diseases by cytogenetics before implanting into a womb....
, invitro fertilization, cryopreservation?, medical abortion
Medical abortion

A medical abortion is a type of non-surgical abortion in which a approved drug is used to induce the abortion. Safe and effective medical abortion methods became an alternative for first trimester pregnancy termination in the 1970s....
, emergency contraception
Emergency contraception

Emergency contraception , or emergency postcoital contraception, refers to birth control measures that, if taken after sex, may prevent pregnancy....
, parahuman
Parahuman

A parahuman or para-human is a human-animal Hybrid . Scientists have done extensive research into the combination of genes from different species, e.g....
s, embryo transfer
Embryo transfer

Embryo transfer refers to a step in the process of in vitro fertilisation whereby one or several embryos are placed into the uterus of the female with the intent to establish a pregnancy....
, genetic engineering
Genetic engineering

Engineering There are a number of ways through which genetic engineering is accomplished. Essentially, the process has five main steps# Isolation of the genes of interest...
 and embryo donation that have been criticized as unethical by pro-life ethicists because they provoke the termination of embryonic cell reproduction. A summary of the embryonic ethical controversies, from a Roman Catholic perspective, can be found in the Vatican document Dignitas Personae, published in Rome
Rome

Rome is the capital city of Italy and Lazio, and is Italy's largest and most populous city, with 2,724,347 residents in an urban area of some ....
 in 2008.

Footnotes



See also

  • Embryogenesis
    Embryogenesis

    Embryogenesis is the process by which the embryo is formed and develops. It starts with the fertilization of the ovum, egg, which, after fertilization, is then called a zygote....
  • Embryology
    Embryology

    Embryology is the study of the development of an embryo. An embryo is defined as any organism in a stage before birth or hatching, or in plants, before germination occurs....
  • In vitro fertilization
  • Plant embryogenesis
    Plant embryogenesis

    Plant embryogenesis is the process that produces a plant embryo from a fertilised ovule by asymmetric cell division and the differentiation of undifferentiated cells into tissues and organs....
  • Embryo space colonization
    Embryo space colonization

    Embryo space colonization is a theoretical interstellar travel space colonization concept that involves sending a robotic mission to a habitable terrestrial planet transporting frozen early-stage human embryos or the technological or biological means to create human embryos....
  • Pregnancy
    Pregnancy

    Pregnancy is the carrying of one or more offspring, known as a fetus or embryo, inside the uterus of a female. In a pregnancy, there can be multiple gestations, as in the case of twins or Multiple birth....
  • Embryo adoption
    Embryo adoption

    Embryo donation is a form of third party reproduction. In vitro fertilisation, or IVF, often results in a number of frozen, unused embryo after the woman for whom they were originally created has successfully carried one or more pregnancies to term....





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