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Albatross



 
 
Albatrosses, of the biological family
Family (biology)

In biological classification, family is a taxonomic rank. Exact details of formal nomenclature depend on the Nomenclature Codes which applies....
 Diomedeidae, are large seabird
Seabird

Seabirds are birds that have adaptation to life within the marine environment. While seabirds vary greatly in lifestyle, behavior and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding ecological niche have resulted in similar adaptations....
s allied to the procellariids
Procellariidae

The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the prion , and the shearwaters. This family is part of the bird order Procellariiformes , which also includes the albatrosses, the storm-petrels, and the diving petrels....
, storm-petrel
Storm-petrel

The storm-petrels are seabirds in the Family Hydrobatidae, part of the order Procellariiformes. These smallest of seabirds feed on planktonic crustaceans and small fish picked from the surface, typically while hovering....
s and diving-petrels in the order Procellariiformes
Procellariiformes

Procellariiformes is an order of seabirds that comprises four family : the albatrosses, Procellariidae, storm-petrels and diving petrels. Formerly called Tubinares and still called tubenoses in English, they are often referred to collectively as the petrels, a term that has been applied to all Procellariiformes or more commo...
 (the tubenoses). They range widely in the Southern Ocean
Southern Ocean

The Southern Ocean, also known as the Great Southern Ocean, the Antarctic Ocean and the South Polar Ocean, comprises the southernmost waters of the World Ocean south of 60th parallel south latitude....
 and the North Pacific
Pacific Ocean

The Pacific Ocean is the largest of the Earth's oceanic divisions. Its name is derived from the Latin name Mare Pacificum, "peaceful sea", bestowed upon it by the Portugal explorer Ferdinand Magellan....
. They are absent from the North Atlantic
Atlantic Ocean

The Atlantic Ocean is the second-largest of the world's oceanic divisions; with a total area of about 106.4 million square kilometres . It covers approximately one-fifth of the Earth's surface....
, although fossil
Fossil

Fossils are the preserved remains or trace fossil of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossiliferous Rock formations and sedimentary rock layers is known as the fossil record....
 remains show they once occurred there too and occasional vagrants turn up.

Albatrosses are amongst the largest of flying
Bird flight

Flight is the main mode of animal locomotion used by most of the world's bird species. Flight assists birds while feeding, breeding and avoiding predation....
 birds, and the great albatross
Great albatross

The great albatrosses are seabirds in the genus Diomedea in the albatross family . The genus Diomedea formerly included all albatrosses except the sooty albatrosses, but in 1996 the genus was split with the mollymawks and the North Pacific albatrosses both being elevated to separate genera....
es (genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 Diomedea) have the largest wingspans of any extant birds.






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Albatrosses, of the biological family
Family (biology)

In biological classification, family is a taxonomic rank. Exact details of formal nomenclature depend on the Nomenclature Codes which applies....
 Diomedeidae, are large seabird
Seabird

Seabirds are birds that have adaptation to life within the marine environment. While seabirds vary greatly in lifestyle, behavior and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding ecological niche have resulted in similar adaptations....
s allied to the procellariids
Procellariidae

The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the prion , and the shearwaters. This family is part of the bird order Procellariiformes , which also includes the albatrosses, the storm-petrels, and the diving petrels....
, storm-petrel
Storm-petrel

The storm-petrels are seabirds in the Family Hydrobatidae, part of the order Procellariiformes. These smallest of seabirds feed on planktonic crustaceans and small fish picked from the surface, typically while hovering....
s and diving-petrels in the order Procellariiformes
Procellariiformes

Procellariiformes is an order of seabirds that comprises four family : the albatrosses, Procellariidae, storm-petrels and diving petrels. Formerly called Tubinares and still called tubenoses in English, they are often referred to collectively as the petrels, a term that has been applied to all Procellariiformes or more commo...
 (the tubenoses). They range widely in the Southern Ocean
Southern Ocean

The Southern Ocean, also known as the Great Southern Ocean, the Antarctic Ocean and the South Polar Ocean, comprises the southernmost waters of the World Ocean south of 60th parallel south latitude....
 and the North Pacific
Pacific Ocean

The Pacific Ocean is the largest of the Earth's oceanic divisions. Its name is derived from the Latin name Mare Pacificum, "peaceful sea", bestowed upon it by the Portugal explorer Ferdinand Magellan....
. They are absent from the North Atlantic
Atlantic Ocean

The Atlantic Ocean is the second-largest of the world's oceanic divisions; with a total area of about 106.4 million square kilometres . It covers approximately one-fifth of the Earth's surface....
, although fossil
Fossil

Fossils are the preserved remains or trace fossil of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossiliferous Rock formations and sedimentary rock layers is known as the fossil record....
 remains show they once occurred there too and occasional vagrants turn up.

Albatrosses are amongst the largest of flying
Bird flight

Flight is the main mode of animal locomotion used by most of the world's bird species. Flight assists birds while feeding, breeding and avoiding predation....
 birds, and the great albatross
Great albatross

The great albatrosses are seabirds in the genus Diomedea in the albatross family . The genus Diomedea formerly included all albatrosses except the sooty albatrosses, but in 1996 the genus was split with the mollymawks and the North Pacific albatrosses both being elevated to separate genera....
es (genus
Genus

A genus is a low-level taxonomic rank used in the classification of living and fossil organisms. The taxonomic ranks are domain , kingdom , phylum, class , order , family , genus, and species....
 Diomedea) have the largest wingspans of any extant birds. The albatrosses are usually regarded as falling into four genera, but there is disagreement over the number of species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
.

Albatrosses are highly efficient in the air, using dynamic soaring
Dynamic soaring

Dynamic soaring is a flying technique used to gain kinetic energy by repeatedly crossing the boundary between air masses of significantly different velocity....
 and slope soaring to cover great distances with little exertion. They feed on squid
Squid

Squid are marine cephalopods of the order Teuthida, which comprises around 300 species. Like all other cephalopods, squid have a distinct head, Symmetry #Bilateral_symmetry, a mantle , and cephalopod arms....
, fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
 and krill
Krill

Krill are a type of shrimp-like marine invertebrate animal. These small crustaceans are important organisms of the zooplankton, particularly as food for baleen whales, manta rays, whale sharks, crabeater seals, and other pinniped, and a few seabird species that feed almost exclusively on them....
 by either scavenging, surface seizing or diving. Albatrosses are colonial, nesting for the most part on remote oceanic islands, often with several species nesting together. Pair bond
Pair bond

In biology, a pair bond is the strong affinity that develops in some species between the males and or females in a pair, potentially leading to breeding....
s between males and females form over several years, with the use of 'ritualised dances', and will last for the life of the pair. A breeding season
Breeding season

The breeding season is the most suitable season, usually with favorable conditions and abundant food and water, for breeding in the wild among some wild animals and birds ....
 can take over a year from laying to fledging
Fledge

Fledge is the stage in a young bird's life when the feathers and wing muscles are sufficiently developed for flight. It also describes the act of raising chicks to a fully grown state by the chick's parents....
, with a single egg
Egg (biology)

In most birds and reptiles, an egg is the zygote, resulting from fertilization of the ovum. To enable incubation the egg is usually kept within a favourable temperature range as it nourishes and protects the growing embryo....
 laid in each breeding attempt.

Of the 21 species of albatrosses recognised by the IUCN
World Conservation Union

The International Union for Conservation of Nature and Natural Resources is an international organization dedicated to natural resource Conservation ethic....
, 19 are threatened with extinction
Extinction

In biology and ecology, extinction is the death of every member of a species or group of taxon. The moment of extinction is generally considered to be the death of the last individual of that species ....
. Numbers of albatrosses have declined in the past due to harvesting for feather
Feather

Feathers are one of the epidermal growths that form the distinctive outer covering, or plumage, on birds. They are considered the most complex integumentary structures found in vertebrates....
s, but today the albatrosses are threatened by introduced species
Introduced species

A species is defined as introduced in a certain geographical area, if that area is outside the species' indigenous distributional range, and the species has arrived there by human activity....
 such as rat
Rat

Rats are various medium sized, long-tailed rodents of the Family Muroidea. "True rats" are members of the genus Rattus, the most important of which to humans are the black rat, Rattus rattus, and the brown rat, Rattus norvegicus....
s and feral cat
Feral cat

A feral cat is an unowned and untamed cat separated from domestication. Feral cats are born in the wild and may take a long time to socialize or may be abandoned or lost pets that have become Wildness....
s that attack eggs, chicks and nesting adults; by pollution
Pollution

Pollution is the introduction of contaminants into an environment that causes instability, disorder, harm or discomfort to the ecosystem i.e. physical systems or living organisms ....
; by a serious decline in fish stocks in many regions largely due to overfishing; and by long-line fishing
Long-line fishing

Longline fishing is a commercial fishing technique. It uses a long line, called the main line, with Fish bait Fish hook attached at intervals by means of branch lines called "snoods"....
. Long-line fisheries pose the greatest threat, as feeding birds are attracted to the bait
Bait (luring substance)

Bait is any substance used to attract prey, e.g. in a mousetrap....
, become hooked on the lines, and drown. Identified stakeholders
Stakeholder (general)

Sorry, no overview for this topic
 such as governments, conservation organisations and people in the fishing industry are all working toward reducing this bycatch.

Albatross biology


Taxonomy and evolution

The albatrosses comprise between 13 and 24 species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
 (the number of species is still a matter of some debate, 21 being the most commonly accepted number) in 4 genera. The four genera are the great albatross
Great albatross

The great albatrosses are seabirds in the genus Diomedea in the albatross family . The genus Diomedea formerly included all albatrosses except the sooty albatrosses, but in 1996 the genus was split with the mollymawks and the North Pacific albatrosses both being elevated to separate genera....
es (Diomedea), the mollymawk
Mollymawk

The mollymawks are a group of medium sized albatrosses that form the genus Thalassarche. The name has sometimes been used for the genus Phoebetria as well, but these are correctly called sooty albatrosses....
s (Thalassarche), the North Pacific albatross
North Pacific albatross

The North Pacific albatrosses are large seabirds from the genus Phoebastria in the albatross family . They are the most tropical of the albatrosses, with two species nesting in North Western Hawaiian island chain, one on sub-tropical islands south of Japan , and one nesting on the equator ....
es (Phoebastria), and the sooty albatross
Sooty albatross

The sooty albatrosses are small albatrosses from the genus Phoebetria. There are two species, the Sooty Albatross and the Light-mantled Albatross ....
es or sooties (Phoebetria). Of the four genera, the North Pacific albatrosses are considered to be a sister taxon to the great albatrosses, while the sooty albatrosses are considered closer to the mollymawks.

The taxonomy
Taxonomy

Taxonomy is the practice and science of classification. The word comes from the Greek language ', taxis and ', nomos .Taxonomies, or taxonomic schemes, are composed of taxonomic units known as taxa , or kinds of things that are arranged frequently in a hierarchical structure....
 of the albatross group has been a source of a great deal of debate. The Sibley-Ahlquist taxonomy
Sibley-Ahlquist taxonomy

The Sibley-Ahlquist taxonomy is a radical bird taxonomy proposed by Charles Sibley and Jon Edward Ahlquist. It is based on DNA-DNA hybridization studies conducted in the late 1970s and throughout the 1980s....
 places seabirds, birds of prey
Bird of prey

Birds of prey are birds that hunt for food primarily on the wing, using their keen senses, especially vision. Their claws and beaks tend to be relatively large, powerful and adapted for tearing and/or piercing flesh....
 and many others in a greatly enlarged order Ciconiiformes
Ciconiiformes

Traditionally, the order Ciconiiformes has included a variety of large, long-legged wading birds with large bills: storks, herons, egrets, ibises, spoonbills, and several others....
, whereas the ornithological organisations in North America, Europe, South Africa, Australia and New Zealand retain the more traditional order Procellariiformes
Procellariiformes

Procellariiformes is an order of seabirds that comprises four family : the albatrosses, Procellariidae, storm-petrels and diving petrels. Formerly called Tubinares and still called tubenoses in English, they are often referred to collectively as the petrels, a term that has been applied to all Procellariiformes or more commo...
. The albatrosses can be separated from the other Procellariiformes both genetically
Genetics

Genetics , a discipline of biology, is the science of heredity and Genetic variation in living organisms. The fact that living things inherit traits from their parents has been used since prehistoric times to improve crop plants and animals through selective breeding....
 and through morphological characteristics, size, their legs and the arrangement of their nasal tubes (see Morphology and flight).

Within the family the assignment of genera has been debated for over a hundred years. Originally placed into a single genus, Diomedea, they were rearranged by Reichenbach
Ludwig Reichenbach

Heinrich Gottlieb Ludwig Reichenbach was a Germany botanist and ornithologist.He was the son of Johann Friedrich Jakob Reichenbach, the author in 1818 of the first Greek-German dictionary....
 into four different genera in 1852, then lumped back together and split apart
Lumpers and splitters

Lumping and splitting refers to a well known problem in any discipline which has to place individual examples into rigorously defined categories....
 again several times, acquiring 12 different genus names in total (though never more than eight at one time) by 1965 (Diomedea, Phoebastria, Thalassarche, Phoebetria, Thalassageron, Diomedella, Nealbatrus, Rhothonia, Julietata, Galapagornis, Laysanornis, and Penthirenia).

By 1965, in an attempt to bring some order back to the classification of albatrosses, they were lumped into two genera, Phoebetria (the sooty albatrosses which most closely seemed to resemble the procellarids and were at the time considered "primitive" ) and Diomedea (the rest). Though there was a case for the simplification of the family (particularly the nomenclature), the classification was based on the morphological analysis of Elliott Coues
Elliott Coues

Elliott Coues was an United States army surgery, historian, ornithologist and author.Coues was born in Portsmouth, New Hampshire. He graduated at George Washington University, Washington, D.C., in 1861, and at the Medical school of that institution in 1863....
 in 1866, and paid little attention to more recent studies and even ignored some of Coues's suggestions.

Albatross Phylogeny
More recent research by Gary Nunn of the American Museum of Natural History
American Museum of Natural History

The American Museum of Natural History , located on the Upper West Side, Manhattan, New York, USA, is one of the largest and most celebrated museums in the world....
 (1996) and other researchers around the world studied the mitochondrial DNA
Mitochondrial DNA

Mitochondrial DNA is the DNA located in organelles called mitochondrion. Most other DNA present in eukaryotic organisms is found in the cell nucleus....
 of all 14 accepted species, finding that there were four, not two, monophyletic groups within the albatrosses. They proposed the resurrection of two of the old genus names, Phoebastria for the North Pacific albatrosses and Thalassarche for the mollymawks, with the great albatrosses retaining Diomedea and the sooty albatrosses staying in Phoebetria. Both the British Ornithologists' Union
British Ornithologists' Union

The British Ornithologists' Union aims to encourage the study of birds in Great Britain, Europe and elsewhere, in order to understand their biology and to aid their Conservation ecology....
 and the South African authorities split the albatrosses into four genera as Nunn suggested, and the change has been accepted by the majority of researchers.
Albatros Ceja Negra   Paso Drake   Noviembre 2005
While there is some agreement on the number of genera, there is less agreement on the number of species. Historically, up to 80 different taxa have been described by different researchers; most of these were incorrectly identified juvenile birds.

Based on the work on albatross genera, Robertson and Nunn went on in 1998 to propose a revised taxonomy with 24 different species, compared to the 14 then accepted. This interim taxonomy elevated many established subspecies
Subspecies

In biology, subspecies is the taxonomic rank immediately subordinate to a species. A subspecies is a taxonomic group which is less distinct than the Common descent or species from which it originates....
 to full species, but was criticised for not using, in every case, peer review
Peer review

Peer review is the process of subjecting an author's Scholarly method work, research, or ideas to the scrutiny of others who are experts in the same field....
ed information to justify the splits. Since then further studies have in some instances supported or disproved the splits; a 2004 paper analysing the mitochondrial DNA
Mitochondrial DNA

Mitochondrial DNA is the DNA located in organelles called mitochondrion. Most other DNA present in eukaryotic organisms is found in the cell nucleus....
 and microsatellite
Microsatellite

Microsatellites, or Simple Sequence Repeats , are Polymorphism loci present in nuclear DNA and organellar DNA DNA that consist of repeating units of 1-6 base pairs in length....
s agreed with the conclusion that the Antipodean Albatross
Antipodean Albatross

The Antipodean Albatross, Diomedea antipodensis, is a large seabird, from the albatross family . Antipodean Albatrosses are smaller than Wandering Albatrosses, and breed in predominantly brown plumage, but are otherwise difficult to distinguish from Wanderers....
 and the Tristan Albatross
Tristan Albatross

The Tristan Albatross, Diomedea dabbenena, is a large seabird from the albatross family . One of the great albatrosses of the genus Diomedea, it was only widely recognised as a full species in 1998....
 were distinct from the Wandering Albatross
Wandering Albatross

The Wandering Albatross, Snowy Albatross, or White-winged Albatross, Diomedea exulans, is a large seabird from the family Diomedeidae which has a circumpolar range in the Southern Ocean....
, per Robertson and Nunn, but found that the suggested Gibson's Albatross, Diomedea gibsoni, was not distinct from the Antipodean Albatross. For the most part, an interim taxonomy of 21 species is accepted by the IUCN and many other researchers, though by no means all — in 2004 Penhallurick and Wink called for the number of species to be reduced to 13 (including the lumping of the Amsterdam Albatross
Amsterdam Albatross

The Amsterdam Albatross or Amsterdam Island Albatross, Diomedea amsterdamensis, is a huge albatross which breeds only on Amsterdam Island in the southern Indian Ocean....
 with the Wandering Albatross
Wandering Albatross

The Wandering Albatross, Snowy Albatross, or White-winged Albatross, Diomedea exulans, is a large seabird from the family Diomedeidae which has a circumpolar range in the Southern Ocean....
), although this paper was itself controversial. On all sides, there is the widespread agreement on the need for further research to clarify the issue.

Sibley and Ahlquist's
Sibley-Ahlquist taxonomy

The Sibley-Ahlquist taxonomy is a radical bird taxonomy proposed by Charles Sibley and Jon Edward Ahlquist. It is based on DNA-DNA hybridization studies conducted in the late 1970s and throughout the 1980s....
 molecular study of the evolution
Evolution

In biology, evolution is change in the heritability trait of a population of organisms from one generation to the next. These changes are caused by a combination of three main processes: variation, reproduction, and selection....
 of the bird families has put the radiation
Adaptive radiation

An adaptive radiation is a rapid evolutionary radiation characterized by an increase in the morphological and ecological diversity of a single, rapidly diversifying lineage....
 of the Procellariiformes
Procellariiformes

Procellariiformes is an order of seabirds that comprises four family : the albatrosses, Procellariidae, storm-petrels and diving petrels. Formerly called Tubinares and still called tubenoses in English, they are often referred to collectively as the petrels, a term that has been applied to all Procellariiformes or more commo...
 in the Oligocene
Oligocene

The Oligocene is a geologic epoch of the Paleogene Geologic Timescale and extends from about 33.9 million to 23 million years before the present....
 period (35–30 million years ago), though this group probably originated earlier, with a fossil
Fossil

Fossils are the preserved remains or trace fossil of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossiliferous Rock formations and sedimentary rock layers is known as the fossil record....
 sometimes attributed to the order, a seabird known as Tytthostonyx
Tytthostonyx

Tytthostonyx is a genus of prehistoric seabird. Found in the much-debated Hornerstown Formation which straddles the Cretaceous-Paleocene boundary 65 million years ago, this animal was apparently closely related to the ancestor of some modern birds, such as Procellariiformes and/or "Pelecaniformes"....
, being found in late Cretaceous
Cretaceous

The Cretaceous , usually abbreviated K for its German translation Kreide, is a geologic period from circa to million years ago . In the geologic timescale, the Cretaceous follows on the Jurassic period and is followed by the Paleogene period....
 rocks (70 mya
Mya (unit)

In astronomy, geology, and paleontology, mya or "m.y.a." is an abbreviation for "million years ago". Like the related unit bya, mya is traditionally written in lower case....
). The molecular evidence suggests that the storm-petrels were the first to diverge from the ancestral stock, and the albatrosses next, with the procellarids and diving petrels separating later. The earliest fossil albatrosses were found in Eocene
Eocene

The Eocene Geologic time scale is a major division of the geologic timescale and the second epoch of the Palaeogene period in the Cenozoic era....
 to Oligocene rocks, although some of these are only tentatively assigned to the family and none appear to be particularly close to the living forms. They are Murunkus (Middle Eocene of Uzbekistan
Uzbekistan

Uzbekistan, officially the Republic of Uzbekistan , is a Landlocked_country#Doubly_landlocked_country country in Central Asia, formerly part of the Soviet Union....
), Manu
Manu (genus)

Manu was a genus of bird which lived during the Early Oligocene. Its fossils have been found in New Zealand, and it appears to be a procellariiform. It is quite likely that this bird was an ancestral albatross....
 (early Oligocene of New Zealand
New Zealand

New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses , and numerous Islands of New Zealand, most notably Stewart Island/Rakiura and the Chatham Islands....
), and an undescribed form from the Late Oligocene of South Carolina
South Carolina

South Carolina is a U.S. state in the Southern United States of the United States. It borders Georgia to the south and North Carolina to the north....
. Similar to the last was Plotornis, formerly often considered a petrel but now accepted as an albatross. It is from the Middle Miocene
Miocene

The Miocene is a Geologic time scale of the Neogene period and extends from about 23.03 to 5.33 million years before the present. As with other older geologic periods, the rock beds that define the start and end are well identified but the exact dates of the start and end of the period are uncertain....
 of France
France

France , officially the French Republic , is a country whose Metropolitan France is located in Western Europe and that also comprises various Overseas departments and territories of France....
, a time when the split between the four modern genera was already underway as evidenced by Phoebastria californica and Diomedea milleri, both being mid-Miocene species from Sharktooth Hill, California
California

California is a U.S. state on the West Coast of the United States of the United States, along the Pacific Ocean. It is bordered by Oregon to the north, Nevada to the east, Arizona to the southeast, and to the south the Mexico state of Baja California....
. These show that the split between the great albatrosses and the North Pacific albatrosses occurred by 15 mya. Similar fossil finds in the southern hemisphere put the split between the sooties and mollymawks at 10 mya. The fossil record of the albatrosses in the northern hemisphere is more complete than that of the southern, and many fossil forms of albatross have been found in the North Atlantic, which today has no albatrosses. The remains of a colony of Short-tailed Albatross
Short-tailed Albatross

The Short-tailed Albatross or Steller's Albatross is a large rare seabird from the North Pacific. Although related to the other North Pacific albatrosses, it also exhibits behavioural and morphological links to the albatrosses of the Southern Ocean....
es have been uncovered on the island of Bermuda
Bermuda

Bermuda is a British overseas territory in the Atlantic Ocean. Located off the east coast of the United States, it is situated around 1770 kilometres northeast of Miami, Florida, and 1350 kilometres south of Halifax Regional Municipality, Canada....
, and the majority of fossil albatrosses from the North Atlantic have been of the genus Phoebastria (the North Pacific albatrosses); one, Phoebastria anglica, has been found in deposits in both North Carolina
North Carolina

North Carolina is a U.S. state located on the Atlantic Seaboard in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north....
 and England
England

native_name =|conventional_long_name = England|common_name = England|image_flag = Flag of England.svg|image_coat = England COA.svg|symbol_type = Royal Coat of Arms...
. See the genus accounts for more data on fossil species.

Morphology and flight

Black Footed Albatross
The albatrosses are a group of large to very large bird
Bird

Birds are wing, Bipedalismal, endothermic , vertebrate animals that lay egg . There are around 10,000 living species, making them the most numerous tetrapod vertebrates....
s; they are the largest of the procellariiformes. The bill
Beak

The beak, bill or rostrum is an external anatomical structure of birds which, in addition to eating, is used for Personal grooming#In animals, manipulating objects, killing prey, probing for food, Courtship#Courtship in the animal kingdom and feeding their young....
 is large, strong and sharp-edged, the upper mandible terminating in a large hook. This bill is composed of several horny plates, and along the sides are the two "tubes", long nostrils that give the order
Order (biology)

In Biological classification used in biology, the order is a taxonomic rank between class and family . The superorder is a rank between class and order....
 its former name. The tubes of all albatrosses are along the sides of the bill, unlike the rest of the Procellariiformes
Procellariiformes

Procellariiformes is an order of seabirds that comprises four family : the albatrosses, Procellariidae, storm-petrels and diving petrels. Formerly called Tubinares and still called tubenoses in English, they are often referred to collectively as the petrels, a term that has been applied to all Procellariiformes or more commo...
 where the tubes run along the top of the bill. These tubes allow the albatrosses to have an acute sense of smell, an unusual ability for birds. Like other Procellariiformes they use this olfactory ability while foraging in order to locate potential food sources. The feet have no hind toe and the three anterior toes are completely webbed. The legs are strong for Procellariiformes, in fact, almost uniquely amongst the order in that they and the giant petrel
Giant petrel

The giant petrels are two large seabirds from the genus Macronectes. Long considered to be conspecific , the two species, the Southern Giant Petrel and Northern Giant Petrel are the largest members of the petrel family , Procellariidae, and considered, with the two fulmars to form a distinct sub-group within the pe...
s are able to walk well on land.

Albatrosses, along with all Procellariiformes
Procellariiformes

Procellariiformes is an order of seabirds that comprises four family : the albatrosses, Procellariidae, storm-petrels and diving petrels. Formerly called Tubinares and still called tubenoses in English, they are often referred to collectively as the petrels, a term that has been applied to all Procellariiformes or more commo...
 have a need to lower their salt content due to their drinking of ocean water. All birds have an enlarged nasal gland at the base of the bill, above their eyes. This gland is inactive in species that don't require it; however the Procellariiformes do require its use. Scientists are uncertain as to its exact processes, but do know in general terms that it removes salt that forms a 5% saline solution that drips out of their nose or is forcibly ejected in some birds.

The adult plumage
Plumage

Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season....
 of most of the albatrosses is usually some variation of dark upper-wing and back, white undersides, often compared to that of a gull
Gull

Gulls are Aves in the family Laridae. They are most closely related to the terns and only distantly related to auks, and skimmers, and more distantly to the waders....
. Of these, the species range from the Southern Royal Albatross
Southern Royal Albatross

The Southern Royal Albatross, Diomedea epomophora, is a large seabird from the albatross family . At an average wingspan of almost , it is the second largest albatross, behind the Wandering Albatross....
 which is almost completely white except for the ends and trailing edges of the wings in fully mature males, to the Amsterdam Albatross
Amsterdam Albatross

The Amsterdam Albatross or Amsterdam Island Albatross, Diomedea amsterdamensis, is a huge albatross which breeds only on Amsterdam Island in the southern Indian Ocean....
 which has an almost juvenile-like breeding plumage with a great deal of brown, particularly a strong brown band around the chest. Several species of mollymawk
Mollymawk

The mollymawks are a group of medium sized albatrosses that form the genus Thalassarche. The name has sometimes been used for the genus Phoebetria as well, but these are correctly called sooty albatrosses....
s and North Pacific albatross
North Pacific albatross

The North Pacific albatrosses are large seabirds from the genus Phoebastria in the albatross family . They are the most tropical of the albatrosses, with two species nesting in North Western Hawaiian island chain, one on sub-tropical islands south of Japan , and one nesting on the equator ....
es have face markings like eye patches or have grey or yellow on the head and nape. Three albatross species, the Black-footed Albatross
Black-footed Albatross

The Black-footed Albatross, Phoebastria nigripes, is a large seabird from the North Pacific. It is one of three albatross that range in the northern hemisphere, nesting on isolated tropical islands....
 and the two sooty albatross
Sooty albatross

The sooty albatrosses are small albatrosses from the genus Phoebetria. There are two species, the Sooty Albatross and the Light-mantled Albatross ....
es, vary completely from the usual patterns and are almost entirely dark brown (or dark grey in places in the case of the Light-mantled Albatross
Light-mantled Albatross

The Light-mantled Albatross , also known as the Grey-mantled Albatross or the Light-mantled Sooty Albatross, is a small albatross in the genus Phoebetria, which it shares with the Sooty Albatross....
). Albatrosses take several years to get their full adult breeding plumage.

The wingspan
Wingspan

The wingspan of an fixed-wing aircraft or a bird, is the distance from the left wingtip to the right wingtip. For example, the Boeing 777 has a wingspan of about 60 m ....
s of the largest great albatrosses (genus Diomedea) are the largest of any bird, exceeding , although the other species' wingspans are considerably smaller . The wings are stiff and cambered, with thickened streamlined leading edges. Albatrosses travel huge distances with two techniques used by many long-winged seabirds, dynamic soaring and slope soaring. Dynamic soaring enables them to minimise the effort needed by gliding across wave fronts gaining energy
Energy

In physics, energy is a scalar physical quantity that describes the amount of Work_ that can be performed by a force. Energy is an attribute of objects and systems that is subject to a conservation law....
 from the vertical wind gradient
Wind gradient

In common usage, wind gradient, more specifically wind speed gradientor wind velocity gradient,or alternatively shear wind,...
. Slope soaring is more straightforward: the albatross turns to the wind, gaining height, from where it can then glide back down to the sea. Albatross have high glide ratio
Glide ratio

Glide ratio, also called, Lift-to-drag ratio, glide number, or finesse, is an aviation term that refers to the distance an aircraft will move forward for any given amount of lost altitude ....
s, around 22:1 to 23:1, meaning that for every metre they drop, they can travel forward . They are aided in soaring by a shoulder-lock, a sheet of tendon
Tendon

A tendon is a tough band of fibrous connective tissue that usually connects muscle to bone and is capable of withstanding tension . Tendons are similar to ligaments except that ligaments join one bone to another....
 that locks the wing when fully extended, allowing the wing to be kept outstretched without any muscle expenditure, a morphological adaptation they share with the giant petrels.

Phoebastria Albatrus1
Albatrosses combine these soaring techniques with the use of predictable weather
Weather

Weather is a set of all the Phenomenon occurring in a given atmosphere at a given time. Weather phenomena lie in the hydrosphere and troposphere....
 systems; albatrosses in the southern hemisphere
Southern Hemisphere

The Southern Hemisphere is the half of a planet that is south of the equator?the word sphere literally means 'half ball'. It is also that half of the celestial sphere south of the celestial equator....
 flying north from their colonies will take a clockwise
Clockwise

A clockwise motion is one that proceeds 'like the clock's hands': from the top to the right, then down and then to the left, and back to the top....
 route, and those flying south will fly counterclockwise. Albatrosses are so well adapted to this lifestyle that their heart rate
Heart rate

Heart rate is a measure of the number of heart beats per minute . The average resting human heart rate is about 70 bpm for adult males and 75 bpm for adult females....
s while flying are close to their basal heart rate when resting. This efficiency is such that the most energetically demanding aspect of a foraging trip is not the distance covered, but the landings, take-offs and hunting they undertake having found a food source. This efficient long-distance travelling underlies the albatross's success as a long-distance forager, covering great distances and expending little energy looking for patchily distributed food sources. Their adaptation to gliding flight makes them dependent on wind and waves, however, as their long wings are ill-suited to powered flight and most species lack the muscles and energy to undertake sustained flapping flight. Albatrosses in calm seas are forced to rest on the ocean's surface until the wind picks up again. The North Pacific albatrosses can use a flight style known as flap-gliding, where the bird progresses by bursts of flapping followed by gliding. When taking off, albatrosses need to take a run up to allow enough air to move under the wing to provide lift
Lift (force)

In the context of a fluid flow relative to a body, the lift force is the Vector #Vector components of the aerodynamic force that is perpendicular to the oncoming flow direction....
.

Distribution and range at sea

Diomedeidae Distribution
Most albatrosses range in the southern hemisphere from Antarctica
Antarctica

Antarctica is Earth's southernmost continent, overlying the South Pole. It is situated in the Antarctica of the southern hemisphere, almost entirely south of the Antarctic Circle, and is surrounded by the Southern Ocean....
 to Australia
Australia

Australia, officially the Commonwealth of Australia, is a country in the southern hemisphere comprising the Australia of the world's smallest continent, the major island of Tasmania, and numerous list of islands of Australia in the Indian Ocean and Pacific Oceans....
, South Africa
South Africa

The Republic of South Africa, also known by Official names of South Africa, is a country located at the southern tip of the continent of Africa....
 and South America
South America

South America is the southern continent of the Americas, situated entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere....
. The exceptions to this are the four North Pacific albatrosses, of which three occur exclusively in the North Pacific, from Hawaii to Japan, California and Alaska; and one, the Waved Albatross
Waved Albatross

The Waved Albatross, Phoebastria irrorata - also known as Galapagos Albatross - is the only member of the Diomedeidae family located in the tropics....
, breeds in the Galapagos Islands
Galápagos Islands

Gal?pagos Islands are an archipelago of Island#Volcanic islands distributed around the equator in the Pacific Ocean, 972 km west of continental Ecuador....
 and feeds off the coast of South America. The need for wind in order to glide is the reason albatrosses are for the most part confined to higher latitudes; being unsuited to sustained flapping flight makes crossing the doldrums
Doldrums

The Doldrums is an area of the Atlantic Ocean, the Pacific Ocean and the Indian Ocean affected by the Intertropical Convergence Zone, a low-pressure area around the equator where the prevailing winds are calm....
 extremely difficult. The exception, the Waved Albatross, is able to live in the equator
Equator

The equator is the intersection of the Earth's surface with the Plane perpendicular to the Earth's rotation and containing the Earth's center of mass....
ial waters around the Galapagos Islands because of the cool waters of the Humboldt Current
Humboldt Current

The Humboldt Current is a cold, low-salinity ocean current that flows north-westward along the west coast of South America from the southern tip of Chile to northern Peru....
 and the resulting winds.

Albatross Shape
It is not known for certain why the albatrosses became extinct in the North Atlantic, although rising sea level
Sea level

Mean sea level is the average height of the sea, with reference to a suitable reference surface. Defining the reference level , however, involves complex measurement, and accurately determining MSL can prove difficult....
s due to an interglacial
Interglacial

An interglacial is a geological interval of warmer global average temperature that separates glacial periods within an ice age. The current Holocene interglacial has persisted since the Pleistocene, about 11,400 years ago....
 warming period are thought to have submerged the site of a Short-tailed Albatross colony that has been excavated in Bermuda. Some southern species have occasionally turned up as vagrants in the North Atlantic and can become exiled, remaining there for decades. One of these exiles, a Black-browed Albatross
Black-browed Albatross

The Black-browed Albatross or Black-browed Mollymawk, Thalassarche melanophrys, is a large seabird of the albatross family Diomedeidae, and it is the most widespread and common albatross....
, returned to gannet
Northern Gannet

The Northern Gannet is a seabird and is the largest member of the gannet family, Sulidae....
 colonies in Scotland
Scotland

conventional_long_name = ScotlandAlba|common_name= Scotland|image_flag = Flag of Scotland.svg|flag_width = 130px...
 for many years in a lonely attempt to breed.

The use of satellite tracking is teaching scientists a great deal about the way albatrosses forage across the ocean in order to find food. They undertake no annual migration
Bird migration

Bird migration refers to the regular seasonal journeys undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather....
, but disperse widely after breeding, in the case of southern hemisphere species, often undertaking circumpolar
Circumpolar

The term circumpolar may refer to:* circumpolar navigation: to travel the world "vertically" traversing both of the poles* the Antarctic Circumpolar Current...
 trips. There is also evidence that there is separation of the ranges of different species at sea. A comparison of the foraging niches
Ecological niche

In ecology, a niche is a term describing the relational position of a species or population in its ecosystem to each other; e.g. a dolphin will be in another ecological niche to one that travels in a different school.....
 of two related species that breed on Campbell Island
Campbell Island

Campbell Island may refer to:* Campbell Island, Torres Strait, Queensland, Australia* Campbell Island , Canada* Campbell Island, New Zealand...
, the Campbell Albatross
Campbell Albatross

The Campbell Albatross or Campbell Mollymawk, Thalassarche impavida, is a medium sized mollymawk in the albatross family . It breeds only on Campbell Island and the associated islet of Jeanette Marie, a small New Zealand island group in the South Pacific....
 and the Grey-headed Albatross
Grey-headed Albatross

The Grey-headed Albatross, Thalassarche chrysostoma, also known as the Grey-headed Mollymawk, is a large seabird from the albatross family ....
, showed the Campbell Albatross primarily fed over the Campbell Plateau
Campbell Plateau

The Campbell Plateau is a large submarine plateau to the south of New Zealand and the Chatham Rise. It originated in the Gondwanan breakup and is part of Zealandia , a largely submerged continent....
 whereas the Grey-Headed Albatross fed in more pelagic, oceanic waters. Wandering Albatross
Wandering Albatross

The Wandering Albatross, Snowy Albatross, or White-winged Albatross, Diomedea exulans, is a large seabird from the family Diomedeidae which has a circumpolar range in the Southern Ocean....
es also react strongly to bathymetry
Bathymetry

Bathymetry is the study of underwater depth, of the third dimension of lake or ocean floors. In other words, bathymetry is the underwater equivalent to hypsometry....
, feeding only in waters deeper than 1000 m (3281 ft); so rigidly did the satellite plots match this contour that one scientist remarked, "It almost appears as if the birds notice and obey a 'No Entry' sign where the water shallows to less than 1000 m". There is also evidence of different ranges for the two sexes of the same species; a study of Tristan Albatross
Tristan Albatross

The Tristan Albatross, Diomedea dabbenena, is a large seabird from the albatross family . One of the great albatrosses of the genus Diomedea, it was only widely recognised as a full species in 1998....
es breeding on Gough Island
Gough Island

Gough Island is a volcanic island rising from the South Atlantic Ocean to heights of over 900 m above sea level with an area of . It is a dependency of Tristan da Cunha, which in turn is a dependency of the British overseas territory of Saint Helena....
 showed that males foraged to the west of Gough and females to the east.

Diet

The albatross diet is predominantly cephalopod
Cephalopod

The cephalopods are the mollusc class Cephalopoda characterized by bilateral symmetry, a prominent head, and a modification of the mollusk foot, a muscular hydrostat, into the form of cephalopod arms or tentacles....
s, fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
 and crustacean
Crustacean

Crustaceans are a large group of arthropods, comprising almost 52,000 described species , and are usually treated as a subphylum . They include various familiar animals, such as crabs, lobsters, crayfish, shrimp, krill and barnacles....
s, although they will also scavenge carrion
Carrion

Carrion refers to the carcass of a dead animal. Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters, or scavengers, include Hyenas, Vultures, Virginia Opossum, Tasmanian Devils, Black Bears, Komodo Dragons, Bald Eagles, Raccoons and Blue-tongued lizards....
 and feed on other zooplankton
Zooplankton

Zooplankton are the heterotrophic type of plankton. Plankton are organisms drifting in the Pelagic zone of oceans, seas, and bodies of fresh water....
. It should be noted that for most species, a comprehensive understanding of diet is only known for the breeding season, when the albatrosses regularly return to land and study is possible. The importance of each of these food sources varies from species to species, and even from population to population; some concentrate on squid
Squid

Squid are marine cephalopods of the order Teuthida, which comprises around 300 species. Like all other cephalopods, squid have a distinct head, Symmetry #Bilateral_symmetry, a mantle , and cephalopod arms....
 alone, others take more krill
Krill

Krill are a type of shrimp-like marine invertebrate animal. These small crustaceans are important organisms of the zooplankton, particularly as food for baleen whales, manta rays, whale sharks, crabeater seals, and other pinniped, and a few seabird species that feed almost exclusively on them....
 or fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
. Of the two albatross species found in Hawaii
Hawaii

File:Pahoehoe and Aa flows at Hawaii.jpgThe State of Hawaii is a U.S. state in the United States, located on an archipelago in the central Pacific Ocean southwest of the continental United States, southeast of Japan, and northeast of Australia....
, one, the Black-footed Albatross
Black-footed Albatross

The Black-footed Albatross, Phoebastria nigripes, is a large seabird from the North Pacific. It is one of three albatross that range in the northern hemisphere, nesting on isolated tropical islands....
, takes mostly fish while the Laysan
Laysan Albatross

The Laysan Albatross, Phoebastria immutabilis, is a large seabird that ranges across the North Pacific. This small two-tone gull-like albatross is the second most common seabird in the Hawaiian Islands, with an estimated population of 2.5 million birds, and is currently expanding its range to new islands....
 feeds on squid.

Light Sooty Albatross Flying
The use of dataloggers at sea that record ingestion of water against time (providing a likely time of feeding) suggest that albatross predominantly feed during the day. Analysis of the squid beaks regurgitated by albatrosses has shown that many of the squid eaten are too large to have been caught alive, and include mid-water species likely to be beyond the reach of albatross, suggesting that, for some species (like the Wandering Albatross
Wandering Albatross

The Wandering Albatross, Snowy Albatross, or White-winged Albatross, Diomedea exulans, is a large seabird from the family Diomedeidae which has a circumpolar range in the Southern Ocean....
), scavenged
Scavenger

Scavenging, or necrophagy, is a carnivorous feeding behaviour in which a predator consumes corpses or carrion that were not killed to be eaten by the predator or others of its species....
 squid may be an important part of the diet. The source of these dead squid is a matter of debate; some certainly comes from squid fisheries, but in nature it primarily comes from the die-off that occurs after squid spawning and the vomit of squid-eating whale
Whale

Whales are marine mammals of order Cetacea which are neither dolphinsmembers, in other words, of the families Oceanic dolphin or River dolphinnor porpoises....
s (sperm whale
Sperm Whale

The Sperm Whale is the largest of all toothed whales and largest living toothed animal. The whale was named after the milky-white waxy substance, spermaceti, found in its head and originally mistaken for sperm or semen....
s, pilot whale
Pilot whale

The pilot whale is either of two species of cetacean in the genus Globicephala. The genus is part of the oceanic dolphin family although their behaviour is closer to that of the larger whales....
s and Southern Bottlenose Whales). The diet of other species, like the Black-browed Albatross
Black-browed Albatross

The Black-browed Albatross or Black-browed Mollymawk, Thalassarche melanophrys, is a large seabird of the albatross family Diomedeidae, and it is the most widespread and common albatross....
 or the Grey-headed Albatross
Grey-headed Albatross

The Grey-headed Albatross, Thalassarche chrysostoma, also known as the Grey-headed Mollymawk, is a large seabird from the albatross family ....
, is rich with smaller species of squid that tend to sink after death, and scavenging is not assumed to play a large role in their diet. Also, some species such as the Black-browed Albatross
Black-browed Albatross

The Black-browed Albatross or Black-browed Mollymawk, Thalassarche melanophrys, is a large seabird of the albatross family Diomedeidae, and it is the most widespread and common albatross....
 and the Waved Albatross
Waved Albatross

The Waved Albatross, Phoebastria irrorata - also known as Galapagos Albatross - is the only member of the Diomedeidae family located in the tropics....
 practice kleptoparisitism from boobies
Booby

The Booby, a type of seabird, is part of the Family Sulidae and the genus Sula. It is closely related to the gannets , which were often included in Sula in former times....
, Phalacrocorax or shearwater
Shearwater

Shearwaters are medium-sized long-winged seabirds. There are more than 30 species of shearwaters, a few larger ones in the genus Calonectris and many smaller species in the genus Puffinus....
s.

Until recently it was thought that albatross were predominantly surface feeders, swimming at the surface and snapping up squid and fish pushed to the surface by currents, predators or death. The deployment of capillary depth recorders, which record the maximum dive depth undertaken by a bird (between attaching it to a bird and recovering it when it returns to land), has shown that while some species, like the Wandering Albatross
Wandering Albatross

The Wandering Albatross, Snowy Albatross, or White-winged Albatross, Diomedea exulans, is a large seabird from the family Diomedeidae which has a circumpolar range in the Southern Ocean....
, do not dive deeper than a metre, some species, like the Light-mantled Albatross
Light-mantled Albatross

The Light-mantled Albatross , also known as the Grey-mantled Albatross or the Light-mantled Sooty Albatross, is a small albatross in the genus Phoebetria, which it shares with the Sooty Albatross....
, have a mean diving depth of almost 5 m and can dive as deep as 12.5 m. In addition to surface feeding and diving, they have now also been observed plunge diving from the air to snatch prey.

Breeding and dancing

Albatrosses are colonial, usually nesting on isolated islands; where colonies are on larger landmasses, they are found on exposed headlands with good approaches from the sea in several directions, like the colony on the Otago Peninsula
Otago Peninsula

The Otago Peninsula is a long, rugged indented finger of land that forms the easternmost part of Dunedin, New Zealand. Volcanic in origin, it forms one wall of the collapsed crater that now forms Otago Harbour....
 in Dunedin, New Zealand. Many Buller's Albatross
Buller's Albatross

Buller's Albatross or Buller's Mollymawk, Thalassarche bulleri, is a small mollymawk in the albatross family . It breeds on islands around New Zealand, and feeds in the seas off Australia and the South Pacific....
es and Black-footed Albatross
Black-footed Albatross

The Black-footed Albatross, Phoebastria nigripes, is a large seabird from the North Pacific. It is one of three albatross that range in the northern hemisphere, nesting on isolated tropical islands....
es nest under trees in open forest. Colonies vary from the very dense aggregations favoured by the mollymawks (Black-browed Albatross
Black-browed Albatross

The Black-browed Albatross or Black-browed Mollymawk, Thalassarche melanophrys, is a large seabird of the albatross family Diomedeidae, and it is the most widespread and common albatross....
 colonies on the Falkland Islands
Falkland Islands

The Falkland Islands are an archipelago in the South Atlantic Ocean, located from the coast of Argentina, west of the Shag Rocks , and north of the British Antarctic Territory ....
 have densities of 70 nests per 100 m˛) to the much looser groups and widely spaced individual nests favoured by the sooty and great albatrosses. All albatross colonies are on islands that historically were free of land mammal
Mammal

Mammals are a class of vertebrate animals whose name is derived from their distinctive feature, mammary glands, with which they feed their young....
s. Albatrosses are highly philopatric
Philopatry

Broadly, philopatry is the behaviour of remaining, or returning to, an individual's birthplace. More specifically, in ecology philopatry is the behaviour of elder offspring sharing the parental burden in the upbringing of their siblings, a classic example of kin selection....
, meaning they will usually return to their natal colony to breed. This tendency to return to their point of origin to breed is so strong that a study of Laysan Albatross
Laysan Albatross

The Laysan Albatross, Phoebastria immutabilis, is a large seabird that ranges across the North Pacific. This small two-tone gull-like albatross is the second most common seabird in the Hawaiian Islands, with an estimated population of 2.5 million birds, and is currently expanding its range to new islands....
 showed that the average distance between hatching site and the site where a bird established its own territory was .

Like most seabirds, albatrosses are K-selected
R/K selection theory

In ecology, r/K selection theory relates to the natural selection of Trait s which promote success in particular environments. The theory originates from work on island biogeography by the ecologists Robert MacArthur and E....
 with regard to their life history, meaning they live much longer than other birds, they delay breeding for longer, and invest more effort into fewer young. Albatrosses are very long lived; most species survive upwards of 50 years, the oldest recorded being a Northern Royal Albatross
Northern Royal Albatross

The Northern Royal Albatross or Toroa, Diomedea sanfordi, is a large seabird from the albatross family . It was split from the closely related Southern Royal Albatross as recently as 1998, though not all scientists support that conclusion and consider both of them to be subspecies of the Royal Albatross....
 that was ringed
Bird ringing

Bird ringing is an aid to studying wild birds, by attaching a small individually numbered metal or plastic ring to their legs or wings, so that various aspects of the bird's life can be studied by the ability to re-find the same individual later....
 as an adult and survived for another 51 years, giving it an estimated age of 61. Given that most albatross ringing projects are considerably younger than that, it is thought likely that other species will prove to live that long and even longer.

Laal Adult Sky Call
Albatrosses reach sexual maturity
Sexual maturity

Sexual maturity is the age or stage when an organism can sexual reproduction. It is sometimes considered synonymous with adulthood, though the two are distinct....
 slowly, after about five years, but even once they have reached maturity, they will not begin to breed for another couple of years (even up to 10 years for some species). Young non-breeders will attend a colony prior to beginning to breed, spending many years practising the elaborate breeding rituals and "dances" that the family is famous for. Birds arriving back at the colony for the first time already have the stereotyped behaviours that compose albatross language
Language

A language is a form of symbol communication in which elements are combined to represents something other than themselves. Language can also refer to the use of such systems as a general phenomenon....
, but can neither "read" that behaviour as exhibited by other birds nor respond appropriately. After a period of trial and error learning
Learning

Learning is acquiring new knowledge, behaviors, skills, Value s, preferences or understanding, and may involve synthesizing different types of information....
, the young birds learn the syntax
Syntax

In linguistics, syntax is the study of the principles and rules for constructing Sentence s in natural languages. In addition to referring to the discipline, the term syntax is also used to refer directly to the rules and principles that govern the sentence structure of any individual language, as in "the Irish syntax"....
 and perfect the dances. This language is mastered more rapidly if the younger birds are around older birds.

The repertoire of behaviour involves synchronised performances of various actions such as preening
Personal grooming

File:Cygnus atratus preening.jpgPersonal grooming is the art of cleaning, grooming, and maintaining parts of the body. In animals, it is a species-typical behavior that is controlled by Biological neural network in the brain....
, pointing, calling, bill clacking, staring, and combinations of such behaviours (like the sky-call). When a bird first returns to the colony it will dance with many partners, but after a number of years the number of birds an individual will interact with drops, until one partner is chosen and a pair is formed. They then continue to perfect an individual language that will eventually be unique to that one pair. Having established a pair bond
Pair bond

In biology, a pair bond is the strong affinity that develops in some species between the males and or females in a pair, potentially leading to breeding....
 that will last for life, however, most of that dance will never be used ever again.

Albatrosses are held to undertake these elaborate and painstaking rituals to ensure that the appropriate partner has been chosen and to perfect partner recognition, as egg laying and chick rearing is a huge investment. Even species that can complete an egg-laying cycle in under a year seldom lay eggs in consecutive years. The great albatrosses (like the Wandering Albatross
Wandering Albatross

The Wandering Albatross, Snowy Albatross, or White-winged Albatross, Diomedea exulans, is a large seabird from the family Diomedeidae which has a circumpolar range in the Southern Ocean....
) take over a year to raise a chick from laying to fledging
Fledge

Fledge is the stage in a young bird's life when the feathers and wing muscles are sufficiently developed for flight. It also describes the act of raising chicks to a fully grown state by the chick's parents....
. Albatrosses lay a single subelliptical egg
Egg (biology)

In most birds and reptiles, an egg is the zygote, resulting from fertilization of the ovum. To enable incubation the egg is usually kept within a favourable temperature range as it nourishes and protects the growing embryo....
, white with reddish brown spots, in a breeding season; if the egg is lost to predators or accidentally broken, then no further breeding attempts are made that year. The larger eggs weigh from . The "divorce" of a pair is a rare occurrence, usually only happening after several years of breeding failure.

All the southern albatrosses create large nest
Bird nest

A bird nest is the spot in which a bird lays and Avian incubation its egg and raises its young. While the term popularly refers to a specific structure made by the bird itself?such as the grassy cup nest of the American Robin or Eurasian Blackbird, or the elaborately woven hanging nest of the Montezuma Oropendola, the Village Weaver or the...
s for their egg, utilizing grass, shrubs, soil, peat, and even penguin
Penguin

Penguins are a group of Aquatic animal, flightless bird birds living almost exclusively in the Southern Hemisphere. Highly adapted for life in the water, penguins have countershading dark and white plumage, and their wings have become Flipper ....
 feathers, whereas the three species in the north Pacific make more rudimentary nests. The Waved Albatross
Waved Albatross

The Waved Albatross, Phoebastria irrorata - also known as Galapagos Albatross - is the only member of the Diomedeidae family located in the tropics....
, on the other hand, makes no nest and will even move its egg around the pair's territory, as much as , sometimes causing it to lose the egg. In all albatross species, both parents incubate the egg in stints that last between one day and three weeks. Incubation lasts around 70 to 80 days (longer for the larger albatrosses), the longest incubation period of any bird. It can be an energetically demanding process, with the adult losing as much as of body weight a day.

After hatching, the chick, which is semi-altricial, is brooded and guarded for three weeks until it is large enough to defend and thermoregulate
Thermoregulation

Thermoregulation is the ability of an organism to keep its core temperature within certain boundaries, even when the surrounding temperature is very different....
 itself. During this period the parents feed the chick small meals when they relieve each other from duty. After the brooding period is over, the chick is fed in regular intervals by both parents. The parents adopt alternative patterns of short and long foraging trips, providing meals that weigh around 12% of their body weight (around ). The meals are composed of both fresh squid
Squid

Squid are marine cephalopods of the order Teuthida, which comprises around 300 species. Like all other cephalopods, squid have a distinct head, Symmetry #Bilateral_symmetry, a mantle , and cephalopod arms....
, fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
 and krill
Krill

Krill are a type of shrimp-like marine invertebrate animal. These small crustaceans are important organisms of the zooplankton, particularly as food for baleen whales, manta rays, whale sharks, crabeater seals, and other pinniped, and a few seabird species that feed almost exclusively on them....
, as well as stomach oil
Stomach oil

Stomach oil is the light oil composed of neutral dietary lipids found in the fore-gut or proventriculus of birds in the order Procellariiformes....
, an energy
Food energy

Food energy is the amount of energy in food that is available through digestion.Like other forms of energy, food energy is expressed in calories or joules....
-rich food that is lighter to carry than undigested prey items. This oil is created in a stomach organ known as a proventriculus
Proventriculus

The proventriculus is part of the digestive system of birds, invertebrates and insects....
 from digested prey items by most tubenoses, and gives them their distinctive musty smell.

Albatross chicks take a long time to fledge. In the case of the great albatrosses, it can take up to 280 days; even for the smaller albatrosses, it takes anywhere between 140 and 170 days. Like many seabirds, albatross chicks will gain enough weight to be heavier than their parents, and prior to fledging they use these reserves to build up body condition (particularly growing all their flight feathers), usually fledging at the same weight as their parents. Between 15% and 65% of those fledged survive to breed. Albatross chicks fledge on their own and receive no further help from their parents, who return to the nest after fledging, unaware their chick has left. Studies of juveniles dispersing at sea have suggested an innate migration behaviour, a genetically coded navigation route, which helps young birds when they are first out at sea.

Albatrosses and humans


Etymology

The name albatross is derived from the Arabic
Arabic language

Arabic is a Central Semitic language, thus related to and classified alongside other Semitic languages languages such as Hebrew language and Aramaic language....
 al-câdous or al-gattas (a pelican
Pelican

A pelican is a large water bird with a distinctive pouch under the beak, belonging to the bird Family Pelecanidae.Along with the darters, cormorants, gannets, boobys, frigatebirds, and tropicbirds, pelicans make up the order Pelecaniformes....
; literally, "the diver"), which travelled to English via the Portuguese
Portuguese language

Portuguese is a Romance language that originated in what is now Galicia and Portugal. It is derived from the Latin language spoken by the Romanization Pre-Roman peoples of the Iberian Peninsula around 2000 years ago....
 form alcatraz ("gannet
Gannet

Gannets are seabirds in the family Sulidae, closely related to the Booby.The gannets are large black and white birds, with long pointed wings and long bills....
"), which is also the origin of the title of the former prison, Alcatraz. The OED
Oxford English Dictionary

The Oxford English Dictionary , published by the Oxford University Press , is a comprehensive dictionary of the English language. Two fully-bound print editions of the OED have been published under its current name, in 1928 and 1989; as of December 2008 the dictionary's current editors have completed a quarter of the third edition....
 notes that the word alcatraz was originally applied to the frigatebird
Frigatebird

The frigatebirds are a family, Fregatidae, of seabirds. There are five species in the single genus Fregata. They are also sometimes called Man of War birds or Pirate birds....
; the modification to albatross was perhaps influenced by Latin
Latin

Latin is an Italic language, historically spoken in Latium and Ancient Rome. Through the Military history of the Roman Empire, Latin spread throughout the Mediterranean and a large part of Europe....
 albus, meaning "white", in contrast to frigatebirds which are black. The Portuguese word albatroz is of English
English language

English is a West Germanic language that originated in Anglo-Saxon England and has lingua franca status in many parts of the world as a result of the military, economic, scientific, political and cultural influence of the British Empire in the 18th, 19th and early 20th centuries and that of the United States from the mid 20th century onwa...
 origin.

They were once commonly known as Goonie birds or Gooney birds, particularly those of the North Pacific. In the southern hemisphere, the name mollymawk is still well established in some areas, which is a corrupted form of malle-mugge, an old Dutch
Dutch language

Dutch is a West Germanic languages spoken by over 22 million people as a first language, and about 5 million people as a second language."1% of the EU population claims to speak Dutch well enough in order to have a conversation." Outside the European Union the number of second language speakers of Dutch is very small. Most native...
 name for the Northern Fulmar
Northern Fulmar

The Northern Fulmar , or Arctic Fulmar lives in the north Atlantic and north Pacific. These fulmars look superficially like gulls, but are unrelated, and are in fact petrels....
. The name Diomedea, assigned to the albatrosses by Linnaeus
Carolus Linnaeus

Carl Linnaeus was a Sweden botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern alpha taxonomy, and is also considered one of the fathers of modern ecology....
, references the mythical metamorphosis of the companions of the Greek warrior Diomedes
Diomedes

Diomedes or Diomed is a hero in Greek mythology, mostly known for his participation in the Trojan War. He was born to Tydeus and Deipyle and later became King of Argos, succeeding his grandfather, Adrastus....
 into birds. Finally, the family name Procellariiformes comes from the Latin
Latin

Latin is an Italic language, historically spoken in Latium and Ancient Rome. Through the Military history of the Roman Empire, Latin spread throughout the Mediterranean and a large part of Europe....
 word procella meaning a violent wind or a storm.

In culture

Albatrosses have been described as "the most legendary of all birds". An albatross is a central emblem in The Rime of the Ancient Mariner
The Rime of the Ancient Mariner

The Rime of the Ancient Mariner is the longest major poem by the England poet Samuel Taylor Coleridge written in 1797?98 and published in the first edition of Lyrical Ballads ....
 by Samuel Taylor Coleridge
Samuel Taylor Coleridge

Samuel Taylor Coleridge was an England poet, critic and Philosophy who was, along with his friend William Wordsworth, one of the founders of the Romanticism in England and one of the Lake Poets....
; a captive albatross is also a metaphor
Metaphor

Metaphor is language that directly compares seemingly unrelated subjects. It is a figure of speech that compares two or more things without using the words "like" or "as." More generally, a metaphor describes a first subject as being or equal to a second object in some way....
 for the počte maudit
Počte maudit

A po?te maudit is a poet living a life outside or against society. Abuse of drugs and alcohol, insanity, crime, violence, and in general any societal sin, often resulting in an early death are typical elements of the biography of a po?te maudit....
 in a poem of Charles Baudelaire
Charles Baudelaire

Charles Pierre Baudelaire was a nineteenth century French poetry, critic and translator. A controversial figure in his lifetime, Baudelaire's name has become a byword for literary and artistic Decadent movement....
. It is from the former poem that the usage of albatross as a metaphor
Albatross (metaphor)

The word albatross is sometimes used to mean an encumbrance, or a wearisome burden. It is an allusion to Samuel Taylor Coleridge poem The Rime of the Ancient Mariner ....
 is derived; someone with a burden or obstacle is said to have 'an albatross around their neck', the punishment given in the poem to the mariner who killed the albatross. In part due to the poem, there is a widespread myth
Urban legend

An urban legend, urban myth, or urban tale is a form of modern folklore consisting of stories thought to be factual by those circulating them....
 that sailors believe it disastrous to shoot or harm an albatross; in truth, however, sailors regularly killed and ate them, but they were often regarded as the souls of lost sailors.

Birdwatching

Albatrosses are popular birds for birdwatchers
Birdwatching

Birdwatching or birding is the observation and study of birds with the naked eye or through a visual enhancement device like binoculars....
 and their colonies popular destinations for ecotourists
Ecotourism

Ecotourism is a form of tourism, that appeals to ecologically and socially conscious individuals. Generally speaking, ecotourism focuses on volunteering, personal growth and learning new ways to live on the planet....
. Regular birdwatching trips are taken out of many coastal towns and cities, like Monterey
Monterey, New South Wales

Monterey is a suburb in Southern Sydney Sydney, in the state of New South Wales, Australia. Monterey is located 15km south of the Sydney central business district and is part of the St George, New South Wales area....
, Kaikoura
Kaikoura

Kaikoura is a town on the east coast of the South Island, New Zealand of New Zealand. It is located on New Zealand State Highway network 180 km north of Christchurch, New Zealand....
, Wollongong and Sydney
Sydney

Sydney is the List of cities in Australia by population in Australia, with a metropolitan area population of approximately 4.34 million . It is the List of Australian capital cities of New South Wales, and was the site of the first British Empire colony in Australia....
, to see pelagic seabird
Seabird

Seabirds are birds that have adaptation to life within the marine environment. While seabirds vary greatly in lifestyle, behavior and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding ecological niche have resulted in similar adaptations....
s, and albatrosses are easily attracted to these sightseeing boats by the deployment of fish oil into the sea. Visits to colonies can be very popular; the Northern Royal Albatross
Northern Royal Albatross

The Northern Royal Albatross or Toroa, Diomedea sanfordi, is a large seabird from the albatross family . It was split from the closely related Southern Royal Albatross as recently as 1998, though not all scientists support that conclusion and consider both of them to be subspecies of the Royal Albatross....
 colony at Taiaroa Head
Taiaroa Head

Taiaroa Head is a headland at the end of the Otago Peninsula in New Zealand, overlooking the mouth of the Otago Harbour. It lies within the city limits of Dunedin....
 in New Zealand attracts 40,000 visitors a year, and more isolated colonies are regular attractions on cruises to sub-Antarctic islands.

Threats and conservation

In spite of often being accorded legendary status, albatrosses have not escaped either indirect or direct pressure from humans. Early encounters with albatrosses by Polynesia
Polynesia

Polynesia is a subregion of Oceania, comprising a large grouping of over 1,000 islands scattered over the central and southern Pacific Ocean....
ns and Aleut
Aleut

The Aleuts are the Alaska Natives of the Aleutian Islands of Alaska, United States and Kamchatka Krai, Russia....
 Indians resulted in hunting and in some cases extirpation from some islands (such as Easter Island
Easter Island

Easter Island is a Polynesian island in the southeastern Pacific Ocean, at the southeastern most point of the Polynesian triangle. The island is a special territory of Chile....
). As European
European ethnic groups

The European peoples are the various nations and ethnic groups of Europe. European ethnology is the field of anthropology focusing on Europe....
s began sailing the world, they too began to hunt albatross, "fishing" for them from boats to serve at the table or blasting them for sport. This sport reached its peak on emigration lines bound for Australia
Australia

Australia, officially the Commonwealth of Australia, is a country in the southern hemisphere comprising the Australia of the world's smallest continent, the major island of Tasmania, and numerous list of islands of Australia in the Indian Ocean and Pacific Oceans....
, and only died down when ships became too fast to fish from, and regulations stopped the discharge of weapons for safety reasons. In the 19th century, albatross colonies, particularly those in the North Pacific, were harvested for the feather trade, leading to the near extinction of the Short-tailed Albatross
Short-tailed Albatross

The Short-tailed Albatross or Steller's Albatross is a large rare seabird from the North Pacific. Although related to the other North Pacific albatrosses, it also exhibits behavioural and morphological links to the albatrosses of the Southern Ocean....
.

Albatross Hook
Of the 21 albatross species recognised by IUCN on their Red List
IUCN Red List

The IUCN Red List of Threatened Species , created in 1963, is the world's most comprehensive inventory of the global Conservation movement status of plant and animal species....
, 19 are threatened, and the other two are near threatened. Two species (as recognised by the IUCN) are considered critically endangered
Endangered species

An endangered species is a population of an organism which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters....
: the Amsterdam Albatross
Amsterdam Albatross

The Amsterdam Albatross or Amsterdam Island Albatross, Diomedea amsterdamensis, is a huge albatross which breeds only on Amsterdam Island in the southern Indian Ocean....
 and the Chatham Albatross
Chatham Albatross

The Chatham Albatross, Chatham Mollymawk, or Chatham Islands Mollymawk, Thalassarche eremita, is a medium-sized black-and-white albatross which breeds only on The Pyramid, a large rock stack in the Chatham Islands, New Zealand....
. One of the main threats is commercial long-line fishing
Long-line fishing

Longline fishing is a commercial fishing technique. It uses a long line, called the main line, with Fish bait Fish hook attached at intervals by means of branch lines called "snoods"....
, as the albatrosses and other seabird
Seabird

Seabirds are birds that have adaptation to life within the marine environment. While seabirds vary greatly in lifestyle, behavior and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding ecological niche have resulted in similar adaptations....
s which will readily feed on offal
Offal

Offal is the entrails and internal organs of a butchered animal. The word does not refer to a particular list of organs, but includes most internal organs other than muscles or bones....
 are attracted to the set bait become hooked on the lines and drown. An estimated 100,000 albatross per year are killed in this fashion. Unregulated pirate fisheries exacerbate the problem.

On Midway Atoll
Midway Atoll

Midway Atoll is a 2.4 square mile atoll located in the North Pacific Ocean , about one-third of the way between Honolulu and Tokyo. Midway Atoll is an unorganized territory, unincorporated territory of the United States....
, collisions between Laysan Albatross
Laysan Albatross

The Laysan Albatross, Phoebastria immutabilis, is a large seabird that ranges across the North Pacific. This small two-tone gull-like albatross is the second most common seabird in the Hawaiian Islands, with an estimated population of 2.5 million birds, and is currently expanding its range to new islands....
 and aircraft have resulted in human and bird deaths as well as severe disruptions in military flight operations. Studies were made in the late 1950s and early 1960s that examined the results of control methods such as the killing of birds, the leveling and clearing of land to eliminate updrafts and the destruction of annual nesting sites. Tall structures such as traffic control and radio towers killed 3000 birds in flight collisions during 1964-1965 before the towers were taken down. Closure of Naval Air Facility Midway Island in 1993 eliminated the problem of collisions with military aircraft. Recent reductions in human activity on the island have helped reduce bird deaths, though lead paint pollution near military buildings continues to poison birds by ingestion. Albatross plumes were popular in the early 20th century. In 1909 alone over 300,000 albatrosses were killed on Midway Island and Laysan Island for their plumes.

Another threat to albatrosses is introduced species
Introduced species

A species is defined as introduced in a certain geographical area, if that area is outside the species' indigenous distributional range, and the species has arrived there by human activity....
, such as rats or feral cat
Feral cat

A feral cat is an unowned and untamed cat separated from domestication. Feral cats are born in the wild and may take a long time to socialize or may be abandoned or lost pets that have become Wildness....
s, which directly attack the albatross or its chicks and eggs. Albatrosses have evolved to breed on islands where land mammals are absent and have not evolved defences against them. Even species as small as mice can be detrimental; on Gough Island
Gough Island

Gough Island is a volcanic island rising from the South Atlantic Ocean to heights of over 900 m above sea level with an area of . It is a dependency of Tristan da Cunha, which in turn is a dependency of the British overseas territory of Saint Helena....
 the chicks of Tristan Albatross
Tristan Albatross

The Tristan Albatross, Diomedea dabbenena, is a large seabird from the albatross family . One of the great albatrosses of the genus Diomedea, it was only widely recognised as a full species in 1998....
es are attacked and eaten alive by introduced house mice
House mouse

The House Mouse is one of the most numerous species of the genus Mus commonly termed a mouse. It is a small mammal and a rodent. In most parts of the world, they live in close proximity to humans....
. Introduced species can have other indirect effects: cattle
Cattle

Cattle, colloquially referred to as cows, are domestication ungulates, a member of the subfamily Bovinae of the family Bovidae. They are raised as livestock for meat , dairy products , leather and as draft animals ....
 overgrazed essential cover on Amsterdam Island threatening the Amsterdam Albatross; on other islands introduced plants reduce potential nesting habitat.

Laysan Albatross Chick Remains
Ingestion of plastic
Plastic

Plastic is the general common term for a wide range of synthetic or semisynthetic organic chemistry solid materials suitable for the manufacture of industrial products....
 flotsam is another problem, one faced by many seabirds. The amount of plastic in the seas has increased dramatically since the first record in the 1960s, coming from waste discarded by ships, offshore dumping, litter on beaches and waste washed to sea by rivers. It is impossible to digest and takes up space in the stomach or gizzard
Gizzard

The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including birds, reptiles, earthworms and some fish....
 that should be used for food, or can cause an obstruction that starves the bird directly. Studies of birds in the North Pacific have shown that ingestion of plastics results in declining body weight
Body weight

Although many people prefer the less-ambiguous term body mass, the term body weight is overwhelmingly used in daily English speech and in biological and medical science contexts to describe the mass of an organism's body....
 and body condition. This plastic is sometimes regurgitated and fed to chicks; a study of Laysan Albatross
Laysan Albatross

The Laysan Albatross, Phoebastria immutabilis, is a large seabird that ranges across the North Pacific. This small two-tone gull-like albatross is the second most common seabird in the Hawaiian Islands, with an estimated population of 2.5 million birds, and is currently expanding its range to new islands....
 chicks on Midway Atoll
Midway Atoll

Midway Atoll is a 2.4 square mile atoll located in the North Pacific Ocean , about one-third of the way between Honolulu and Tokyo. Midway Atoll is an unorganized territory, unincorporated territory of the United States....
 showed large amounts of ingested plastic in naturally dead chicks compared to healthy chicks killed in accidents. While not the direct cause of death, this plastic causes physiological stress and causes the chick to feel full during feedings, reducing its food intake and the chances of survival.

Scientists and conservationists (most importantly BirdLife International
BirdLife International

BirdLife International is the international Conservation ecology organization working to bird conservation the world?s birds and their habitats....
 and their partners, who run the Save the Albatross campaign) are working with governments and fishermen to find solutions to the threats albatrosses face. Techniques such as setting long-line bait at night, dying the bait blue, setting the bait underwater, increasing the amount of weight on lines and using bird scarers can all reduce the seabird by-catch. For example, a collaborative study between scientists and fishermen in New Zealand
New Zealand

New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses , and numerous Islands of New Zealand, most notably Stewart Island/Rakiura and the Chatham Islands....
 successfully tested an underwater setting device for long-liners which set the lines below the reach of vulnerable albatross species. The use of some of these techniques in the Patagonian Toothfish
Patagonian toothfish

The Patagonian toothfish is a fish found in the cold, temperate waters of the Southern Atlantic Ocean, Southern Pacific Ocean, Indian Ocean, and Southern Oceans on seamounts and Continental shelf around most sub-Antarctic islands....
 fishery in the Falkland Islands
Falkland Islands

The Falkland Islands are an archipelago in the South Atlantic Ocean, located from the coast of Argentina, west of the Shag Rocks , and north of the British Antarctic Territory ....
 is thought to have reduced the number of Black-browed Albatross
Black-browed Albatross

The Black-browed Albatross or Black-browed Mollymawk, Thalassarche melanophrys, is a large seabird of the albatross family Diomedeidae, and it is the most widespread and common albatross....
 taken by the fleet in the last 10 years. Conservationists have also worked on the field of island restoration
Island restoration

The ecological restoration of islands, or island restoration, is the application of the principles of ecological restoration to islands and island groups....
, removing introduced species that threaten native wildlife, which protects albatrosses from introduced predators.

One important step towards protecting albatrosses and other seabird
Seabird

Seabirds are birds that have adaptation to life within the marine environment. While seabirds vary greatly in lifestyle, behavior and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding ecological niche have resulted in similar adaptations....
s is the 2001 treaty
Treaty

A Treaty is an agreement under international law entered into by actors in international law, namely states and international organizations. A Treaty may also be known as: agreement, protocol, covenant, convention, exchange of letters, etc....
 the Agreement on the Conservation of Albatrosses and Petrels
Agreement on the Conservation of Albatrosses and Petrels

The Agreement on the Conservation of Albatrosses and Petrels is a legally-binding international treaty signed in 2001.It was created in order to halt the drastic decline of seabird populations in the Southern Hemisphere, particularly albatrosses and Procellariidae....
, which came into force in 2004 and has been ratified by eight countries, Australia
Australia

Australia, officially the Commonwealth of Australia, is a country in the southern hemisphere comprising the Australia of the world's smallest continent, the major island of Tasmania, and numerous list of islands of Australia in the Indian Ocean and Pacific Oceans....
, Ecuador
Ecuador

Ecuador , officially the , literally, "Republic of the equator") is a representative democratic republic in South America, bordered by Colombia on the north, by Peru on the east and south, and by the Pacific Ocean to the west....
, New Zealand
New Zealand

New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses , and numerous Islands of New Zealand, most notably Stewart Island/Rakiura and the Chatham Islands....
, Spain
Spain

Spain or the Kingdom of Spain , is a country located in Southern Europe on the Iberian Peninsula.The Spanish constitution does not establish any official denomination of the country, even though Espa?a , Estado espa?ol and Naci?n espa?ola are used interchangeably....
, South Africa
South Africa

The Republic of South Africa, also known by Official names of South Africa, is a country located at the southern tip of the continent of Africa....
, France
France

France , officially the French Republic , is a country whose Metropolitan France is located in Western Europe and that also comprises various Overseas departments and territories of France....
, Peru
Peru

Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....
 and the United Kingdom
United Kingdom

The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom , the UK or Britain,is a sovereign state located off the northwestern coast of continental Europe....
. The treaty requires these countries to take specific actions to reduce by-catch, pollution and to remove introduced species from nesting islands. The treaty has also been signed but not ratified by another three countries, Argentina
Argentina

Argentina, officially the Argentine Republic , is a country in South America, constituted as a federation of 23 provinces and an autonomous city....
, Brazil
Brazil

Brazil , officially the Federative Republic of Brazil , is a country in South America. It is the List of countries and outlying territories by total area country by geographical area, occupying nearly half of South America, the List of countries by population country, and the fourth most populous democracy in the world....
 and Chile
Chile

Chile, officially the Republic of Chile , is a country in South America occupying a long and narrow coastal strip wedged between the Andes mountains and the Pacific Ocean....
.

Species

Current thinking divides the albatrosses into four genera. The number of species is a matter of some debate. The IUCN and BirdLife International
BirdLife International

BirdLife International is the international Conservation ecology organization working to bird conservation the world?s birds and their habitats....
 among others recognise the interim taxonomy of 22 extant species, other authorities retain the more traditional 14 species, and one recent paper proposed a reduction to 13:
  • Great albatrosses (Diomedea)
    • Wandering Albatross
      Wandering Albatross

      The Wandering Albatross, Snowy Albatross, or White-winged Albatross, Diomedea exulans, is a large seabird from the family Diomedeidae which has a circumpolar range in the Southern Ocean....
        D. exulans
    • Antipodean Albatross
      Antipodean Albatross

      The Antipodean Albatross, Diomedea antipodensis, is a large seabird, from the albatross family . Antipodean Albatrosses are smaller than Wandering Albatrosses, and breed in predominantly brown plumage, but are otherwise difficult to distinguish from Wanderers....
       D. (exulans) antipodensis
    • Amsterdam Albatross
      Amsterdam Albatross

      The Amsterdam Albatross or Amsterdam Island Albatross, Diomedea amsterdamensis, is a huge albatross which breeds only on Amsterdam Island in the southern Indian Ocean....
       D. (exulans) amsterdamensis
    • Tristan Albatross
      Tristan Albatross

      The Tristan Albatross, Diomedea dabbenena, is a large seabird from the albatross family . One of the great albatrosses of the genus Diomedea, it was only widely recognised as a full species in 1998....
       D. (exulans) dabbenena
    • Northern Royal Albatross
      Northern Royal Albatross

      The Northern Royal Albatross or Toroa, Diomedea sanfordi, is a large seabird from the albatross family . It was split from the closely related Southern Royal Albatross as recently as 1998, though not all scientists support that conclusion and consider both of them to be subspecies of the Royal Albatross....
       D. (epomorpha) sanfordi
    • Southern Royal Albatross
      Southern Royal Albatross

      The Southern Royal Albatross, Diomedea epomophora, is a large seabird from the albatross family . At an average wingspan of almost , it is the second largest albatross, behind the Wandering Albatross....
       D. epomophora
  • North Pacific albatross
    North Pacific albatross

    The North Pacific albatrosses are large seabirds from the genus Phoebastria in the albatross family . They are the most tropical of the albatrosses, with two species nesting in North Western Hawaiian island chain, one on sub-tropical islands south of Japan , and one nesting on the equator ....
    es (Phoebastria)
    • Waved Albatross
      Waved Albatross

      The Waved Albatross, Phoebastria irrorata - also known as Galapagos Albatross - is the only member of the Diomedeidae family located in the tropics....
       P. irrorata
    • Short-tailed Albatross
      Short-tailed Albatross

      The Short-tailed Albatross or Steller's Albatross is a large rare seabird from the North Pacific. Although related to the other North Pacific albatrosses, it also exhibits behavioural and morphological links to the albatrosses of the Southern Ocean....
       P. albatrus
    • Black-footed Albatross
      Black-footed Albatross

      The Black-footed Albatross, Phoebastria nigripes, is a large seabird from the North Pacific. It is one of three albatross that range in the northern hemisphere, nesting on isolated tropical islands....
       P. nigripes
    • Laysan Albatross
      Laysan Albatross

      The Laysan Albatross, Phoebastria immutabilis, is a large seabird that ranges across the North Pacific. This small two-tone gull-like albatross is the second most common seabird in the Hawaiian Islands, with an estimated population of 2.5 million birds, and is currently expanding its range to new islands....
       P. immutabilis
  • Mollymawk
    Mollymawk

    The mollymawks are a group of medium sized albatrosses that form the genus Thalassarche. The name has sometimes been used for the genus Phoebetria as well, but these are correctly called sooty albatrosses....
    s (Thalassarche)
    • Black-browed Albatross
      Black-browed Albatross

      The Black-browed Albatross or Black-browed Mollymawk, Thalassarche melanophrys, is a large seabird of the albatross family Diomedeidae, and it is the most widespread and common albatross....
       T. melanophris
    • Campbell Albatross
      Campbell Albatross

      The Campbell Albatross or Campbell Mollymawk, Thalassarche impavida, is a medium sized mollymawk in the albatross family . It breeds only on Campbell Island and the associated islet of Jeanette Marie, a small New Zealand island group in the South Pacific....
       T. (melanophris) impavida
    • Shy Albatross
      Shy Albatross

      The Shy Albatross or Shy Mollymawk, Thalassarche cauta, is a medium sized albatross that breeds off Australia and New Zealand's sub-Antarctic islands and ranges extensively across the Southern Ocean....
       T. cauta
    • White-capped Albatross
      White-capped Albatross

      The White-capped Albatross, Thalassarche steadi, is a mollymawk that breeds on the islands off of New Zealand. It is a species that not all experts agree should have been split off of Thalassarche cauta....
       T. steadi
    • Chatham Albatross
      Chatham Albatross

      The Chatham Albatross, Chatham Mollymawk, or Chatham Islands Mollymawk, Thalassarche eremita, is a medium-sized black-and-white albatross which breeds only on The Pyramid, a large rock stack in the Chatham Islands, New Zealand....
       T. (cauta) eremita
    • Salvin's Albatross
      Salvin's Albatross

      Salvin's Albatross, or Salvin's Mollymawk, Thalassarche salvini, is a large seabird that ranges across the Southern Ocean. A medium sized mollymawk in the albatross family , it was long considered to be a subspecies of the Shy Albatross....
       T. (cauta) salvini
    • Grey-headed Albatross
      Grey-headed Albatross

      The Grey-headed Albatross, Thalassarche chrysostoma, also known as the Grey-headed Mollymawk, is a large seabird from the albatross family ....
       T. chrysostoma
    • Atlantic Yellow-nosed Albatross
      Atlantic Yellow-nosed Albatross

      The Atlantic Yellow-nosed Albatross, Thalassarche chlororhynchos, is a large seabird in the albatross family . This small mollymawk was once considered conspecific with the Indian Yellow-nosed Albatross and known as the Yellow-nosed Albatross....
       T. chlororhynchos
    • Indian Yellow-nosed Albatross
      Indian Yellow-nosed Albatross

      The Indian Yellow-nosed Albatross, Thalassarche carteri, in the albatross family , and is a smallest of the mollymawks. In 2004, BirdLife International split this species from the Atlantic Yellow-nosed Albatross;however Clements has not split it yet, and the SACC has not either, but recognizes the need for a proposal....
       T. (chlororhynchos) carteri
    • Buller's Albatross
      Buller's Albatross

      Buller's Albatross or Buller's Mollymawk, Thalassarche bulleri, is a small mollymawk in the albatross family . It breeds on islands around New Zealand, and feeds in the seas off Australia and the South Pacific....
       T. bulleri
  • Sooty albatross
    Sooty albatross

    The sooty albatrosses are small albatrosses from the genus Phoebetria. There are two species, the Sooty Albatross and the Light-mantled Albatross ....
    es (Phoebetria)
    • Sooty Albatross
      Sooty albatross

      The sooty albatrosses are small albatrosses from the genus Phoebetria. There are two species, the Sooty Albatross and the Light-mantled Albatross ....
       P. fusca
    • Light-mantled Albatross
      Light-mantled Albatross

      The Light-mantled Albatross , also known as the Grey-mantled Albatross or the Light-mantled Sooty Albatross, is a small albatross in the genus Phoebetria, which it shares with the Sooty Albatross....
       P. palpebrata.


External links

  • (Handbook of Australian, New Zealand and Antarctic Birds.)
  • The global distribution of albatrosses and petrels: Results from the Global Procellariiform Tracking Workshop, 1–5 September, 2003, Gordon’s Bay, South Africa. BirdLife International
    BirdLife International

    BirdLife International is the international Conservation ecology organization working to bird conservation the world?s birds and their habitats....
  • on the Internet Bird Collection