Tichosteus
Encyclopedia
Tichosteus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of herbivorous
Herbivore
Herbivores are organisms that are anatomically and physiologically adapted to eat plant-based foods. Herbivory is a form of consumption in which an organism principally eats autotrophs such as plants, algae and photosynthesizing bacteria. More generally, organisms that feed on autotrophs in...

 dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

 from the Late Jurassic
Jurassic
The Jurassic is a geologic period and system that extends from about Mya to  Mya, that is, from the end of the Triassic to the beginning of the Cretaceous. The Jurassic constitutes the middle period of the Mesozoic era, also known as the age of reptiles. The start of the period is marked by...

. It is known only from vertebrae recovered from Kimmeridgian
Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age or stage in the Late or Upper Jurassic epoch or series. It spans the time between 155.7 ± 4 Ma and 150.8 ± 4 Ma . The Kimmeridgian follows the Oxfordian and precedes the Tithonian....

 rocks in the Morrison Formation
Morrison Formation
The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone and limestone and is light grey, greenish...

, Colorado
Colorado
Colorado is a U.S. state that encompasses much of the Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains...

.

In 1877 Edward Drinker Cope
Edward Drinker Cope
Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

 named the type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 Tichosteus lucasanus. The generic name is derived from Greek teichos, "wall", and osteon, "bone", referring to the fact that the vertebrae, though hollow inside, had no lateral pneumatic foramina, openings in the side wall for air sacks to invade the bone. The specific name honours superintendent of public schools Oramel W. Lucas, who collected two vertebrae for Cope, near Arkansas River
Arkansas River
The Arkansas River is a major tributary of the Mississippi River. The Arkansas generally flows to the east and southeast as it traverses the U.S. states of Colorado, Kansas, Oklahoma, and Arkansas. The river's initial basin starts in the Western United States in Colorado, specifically the Arkansas...

. The dorsal vertebrae, together forming the holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 AMNH 5770, are about 23 millimetre (0.905511811023622 in) long. Cope estimated the size of the animal as that of an alligator
Alligator
An alligator is a crocodilian in the genus Alligator of the family Alligatoridae. There are two extant alligator species: the American alligator and the Chinese alligator ....

; Charles Craig Mook later compared it to a wolf. The specimen represents a subadult individual, as can be seen from the suture between the vertebral centrum
Centrum
Centrum means center in Latin.Centrum may refer to:* The central portion of a vertebra*Centrum , a Washington state performing arts organization* Centrum , metro station in Warsaw, Poland...

 and the neural arch.

In 1878 Cope named a second species: Tichosteus aequifacies. The specific name is derived from Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 aequus, "equal", and facies, "face", referring to the more symmetric extremities of the vertebrae as compared to T. lucasanus. The type specimen, AMNH 5771, again consists of two vertebrae.

Cope himself was rather puzzled by the affinities of Tichosteus, providing no classification apart from the fact that it was named in an article dedicated to reptiles. Later authors often saw Tichosteus as a theropod. At the end of the twentieth century it was suggested that it may have been a basal iguanodont
Iguanodont
Iguanodonts were herbivorous dinosaurs that lived from the mid-Jurassic to Late Cretaceous. Some members include Camptosaurus, Callovosaurus, Iguanodon, Ouranosaurus, and the hadrosaurids or "duck-billed dinosaurs". Iguanodonts were one of the first groups of dinosaurs to be found...

; the species are nomina dubia
Nomen dubium
In zoological nomenclature, a nomen dubium is a scientific name that is of unknown or doubtful application...

and are incertae sedis
Incertae sedis
, is a term used to define a taxonomic group where its broader relationships are unknown or undefined. Uncertainty at specific taxonomic levels is attributed by , , and similar terms.-Examples:*The fossil plant Paradinandra suecica could not be assigned to any...

within Ornithopoda.
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