Harpactognathus
Encyclopedia
Harpactognathus was a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of pterosaur
Pterosaur
Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...

 found in the Late Jurassic
Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.2 ± 4.0 to 145.5 ± 4.0 million years ago , which is preserved in Upper Jurassic strata. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age...

-age Morrison Formation
Morrison Formation
The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone and limestone and is light grey, greenish...

 of Albany County, Wyoming
Wyoming
Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...

, USA. It is based on NAMAL 101, a partial skull consisting of the snout, recovered from near Bone Cabin Quarry
Bone Cabin Quarry
Bone Cabin Quarry lies approximately fifteen miles north of Laramie, Wyoming near historic Como Bluff. During the summer of 1897 Walter W. Granger, a paleontologist from the American Museum of Natural History, came upon a hillside littered with Jurassic period dinosaur bone fragments...

 in 1996. The species name honors the discoverer, Joe Gentry, a volunteer for the Western Paleontological Laboratories, in Lehi
Lehi, Utah
-Attractions:Lehi Roller MillsLehi Roller Mills was founded in 1906 by a co-op of farmers. George G. Robinson purchased the mill in 1910, and since then it has remained in the family. It is run today by grandson R. Sherman Robinson....

, Utah
Utah
Utah is a state in the Western United States. It was the 45th state to join the Union, on January 4, 1896. Approximately 80% of Utah's 2,763,885 people live along the Wasatch Front, centering on Salt Lake City. This leaves vast expanses of the state nearly uninhabited, making the population the...

.

Its describers found it to be most similar to Scaphognathus
Scaphognathus
Scaphognathus was a pterosaur that lived around Germany during the Late Jurassic. It had a wingspan of about one meter.-Naming:The first known Scaphognathus specimen was described in 1831 by August Goldfuss who mistook the tailless specimen for a new Pterodactylus species: P. crassirostris. The...

among pterosaurs, albeit substantially larger (estimated skull length of 280-300 mm (11-12 in), estimated wingspan
Wingspan
The wingspan of an airplane or a bird, is the distance from one wingtip to the other wingtip. For example, the Boeing 777 has a wingspan of about ; and a Wandering Albatross caught in 1965 had a wingspan of , the official record for a living bird.The term wingspan, more technically extent, is...

 of at least 2.5 m (8.2 ft)). Because of the similarity, Harpactognathus was assigned to the subfamily Scaphognathinae of the family Rhamphorhynchidae
Rhamphorhynchidae
Rhamphorhynchidae is a group of early "rhamphorhynchoid" pterosaurs named after Rhamphorhynchus, that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.-Classification:...

. This genus is also notable for having a low bony crest running all the way to the tip of the beak
Beak
The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

 (pterosaur bone crests usually do not reach the tip) and for being the oldest known Morrison Formation pterosaur, having been found in the Salt Wash Member (Kimmeridgian
Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age or stage in the Late or Upper Jurassic epoch or series. It spans the time between 155.7 ± 4 Ma and 150.8 ± 4 Ma . The Kimmeridgian follows the Oxfordian and precedes the Tithonian....

).

Paleobiology

As a rhamphorhynchid, Harpactognathus would have been an aerial
Flying and gliding animals
A number of animals have evolved aerial locomotion, either by powered flight or by gliding. Flying and gliding animals have evolved separately many times, without any single ancestor. Flight has evolved at least four times, in the insects, pterosaurs, birds, and bats. Gliding has evolved on many...

 predator of small prey; the authors hypothesized that scaphognathines specialized in inland freshwater habitats.
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