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Epigeal

Epigeal

Overview
Epigeal, epigean, epigeic and epigeous are biological terms describing an organism
Organism
In biology, an organism is any living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homeostasis as a stable whole...

's activity above the soil
Soil
Soil is a natural body consisting of layers of mineral constituents of variable thicknesses, which differ from the parent materials in their morphological, physical, chemical, and mineralogical characteristics. It is composed of particles of broken rock that have been altered by chemical and...

 surface.

In botany
Botany
Botany, plant science, phytology, or plant biology is a branch of biology and is the scientific study of plant life and development...

, a seed
Seed
A seed , referred to as a kernel in some plants, is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

 is described as epigeal when the cotyledon
Cotyledon
A cotyledon is a significant part of the embryo within the seed of a plant. Upon germination, the cotyledon may become the embryonic first leaves of a seedling. The number of cotyledons present is one characteristic used by botanists to classify the flowering plants...

s of the germinating seed expand, throw off the seed shell and become photosynthetic
Photosynthesis
Photosynthesis is a process that converts carbon dioxide into organic compounds, especially sugars, using the energy from sunlight. Photosynthesis occurs in plants, algae, and many species of Bacteria, but not in Archaea...

 above the ground. The opposite kind, where the cotyledons remain non-photosynthetic, inside the seed shell, and below ground, is hypogeal
Hypogeal
Hypogeal means "underground".* In botany, a seed is described as hypogeal when the cotyledons of the germinating seed remain non-photosynthetic, inside the seed shell, and below ground...

.

An organism is epigean, epigeic or epigeous if it crawls (epigean), creeps like a vine
Vine
The term vine may refer to a climbing or trailing plant. The word, derived from Latin vīnea, in the original sense referred to the grapevines . The modern extended sense is mostly restricted to North American English, which uses "grapevine" to refer to the grape-bearing Vitis species...

 (epigeal), or grows (epigeous) on the soil surface, or more generally in animal
Animal
Animals are a major group of mostly multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously...

s, neither burrows nor swims nor flies.
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Encyclopedia
Epigeal, epigean, epigeic and epigeous are biological terms describing an organism
Organism
In biology, an organism is any living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homeostasis as a stable whole...

's activity above the soil
Soil
Soil is a natural body consisting of layers of mineral constituents of variable thicknesses, which differ from the parent materials in their morphological, physical, chemical, and mineralogical characteristics. It is composed of particles of broken rock that have been altered by chemical and...

 surface.

In botany
Botany
Botany, plant science, phytology, or plant biology is a branch of biology and is the scientific study of plant life and development...

, a seed
Seed
A seed , referred to as a kernel in some plants, is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

 is described as epigeal when the cotyledon
Cotyledon
A cotyledon is a significant part of the embryo within the seed of a plant. Upon germination, the cotyledon may become the embryonic first leaves of a seedling. The number of cotyledons present is one characteristic used by botanists to classify the flowering plants...

s of the germinating seed expand, throw off the seed shell and become photosynthetic
Photosynthesis
Photosynthesis is a process that converts carbon dioxide into organic compounds, especially sugars, using the energy from sunlight. Photosynthesis occurs in plants, algae, and many species of Bacteria, but not in Archaea...

 above the ground. The opposite kind, where the cotyledons remain non-photosynthetic, inside the seed shell, and below ground, is hypogeal
Hypogeal
Hypogeal means "underground".* In botany, a seed is described as hypogeal when the cotyledons of the germinating seed remain non-photosynthetic, inside the seed shell, and below ground...

.

An organism is epigean, epigeic or epigeous if it crawls (epigean), creeps like a vine
Vine
The term vine may refer to a climbing or trailing plant. The word, derived from Latin vīnea, in the original sense referred to the grapevines . The modern extended sense is mostly restricted to North American English, which uses "grapevine" to refer to the grape-bearing Vitis species...

 (epigeal), or grows (epigeous) on the soil surface, or more generally in animal
Animal
Animals are a major group of mostly multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously...

s, neither burrows nor swims nor flies. Consequently, the opposite term depends on the circumstances. It can be fossorial
Fossorial
A fossorial organism is one that is adapted to digging and life underground such as the badger, the naked mole rat, and the mole salamanders Ambystomatidae...

 (burrow
Burrow
A burrow is a hole or tunnel dug into the ground by an animal to create a space suitable for habitation, temporary refuge, or as a byproduct of locomotion. Burrows provide a form of shelter against predation and exposure to the elements, so the burrowing way of life is quite popular among the...

ing), troglobitic
Troglobite
A troglobite is an animal that lives entirely in the dark parts of caves. Such creatures have become specifically adapted for life in total darkness and over time they have evolved to develop improved senses of smell, taste and vibration detection, while losing anatomical features that are...

 - or stygobitic
Stygofauna
Stygofauna are any fauna that live within groundwater systems, such as caves and aquifers, or more specifically small, aquatic groundwater invertebrates, though terrestrial air-breathing subterranean animals are also sometimes included...

, hypogean etc. - (for cave
Cave
A cave or cavern is a natural underground void large enough for a human to enter. Some people suggest that the term cave should only apply to cavities that have some part that does not receive daylight; however, in popular usage, the term includes smaller spaces like sea caves, rock shelters, and...

-living organisms), or hypogeic and hypogeous (for plants and fungi that grow underground).