Corneous
Encyclopedia
Corneous is a biological and medical term meaning horny, in other words made out of a substance similar to that of horn
Horn (anatomy)
A horn is a pointed projection of the skin on the head of various animals, consisting of a covering of horn surrounding a core of living bone. True horns are found mainly among the ruminant artiodactyls, in the families Antilocapridae and Bovidae...

s and hooves in some mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...

s.

The word is generally used to describe natural or pathological anatomical structures made out of a hard layer of protein
Protein
Proteins are biochemical compounds consisting of one or more polypeptides typically folded into a globular or fibrous form, facilitating a biological function. A polypeptide is a single linear polymer chain of amino acids bonded together by peptide bonds between the carboxyl and amino groups of...

. In mammals this protein would usually be keratin
Keratin
Keratin refers to a family of fibrous structural proteins. Keratin is the key of structural material making up the outer layer of human skin. It is also the key structural component of hair and nails...

.

The word corneous is also often used to describe the operculum
Operculum (gastropod)
The operculum, meaning little lid, is a corneous or calcareous anatomical structure which exists in many groups of sea snails and freshwater snails, and also in a few groups of land snails...

 of a snail
Snail
Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

, a gastropod mollusc. Not all gastropods have opercula, but in the great majority of those that do have one, the operculum is corneous. (However in several genera within a few families including the marine Naticidae
Naticidae
Naticidae, common name the moon snails, is a family of minute to large-sized predatory sea snails, marine gastropod molluscs in the clade Littorinimorpha.Naticidae is the only family in the superfamily Naticoidea....

 and the terrestrial Pomatiidae
Pomatiidae
The family Pomatiidae is a taxonomic family of small operculate land snails, terrestrial gastropod mollusks that can be found over the warmer parts of the Old World. In the older literature, this family is designated as Pomatiasidae....

, the operculum is primarily calcareous
Calcareous
Calcareous is an adjective meaning mostly or partly composed of calcium carbonate, in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines.-In zoology:...

, in other words mostly made of calcium carbonate
Calcium carbonate
Calcium carbonate is a chemical compound with the formula CaCO3. It is a common substance found in rocks in all parts of the world, and is the main component of shells of marine organisms, snails, coal balls, pearls, and eggshells. Calcium carbonate is the active ingredient in agricultural lime,...

.) Corneous opercula are made out of the protein conchiolin
Conchiolin
Conchiolin and perlucin are complex proteins which are secreted by a mollusc's outer epithelium ....

.
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