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Coral Reef

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Coral reef



 
 
Coral reefs are aragonite
Aragonite

Aragonite is a carbonate mineral, one of the two common, naturally occurring polymorphism of calcium carbonate, calciumcarbonoxygen3....
 structures produced by living organisms. In most reefs the predominant organisms are colonial cnidarians that secrete an exoskeleton
Exoskeleton

An exoskeleton is an external skeleton that supports and protects an animal's body, in contrast to the internal endoskeleton of, for example, a human skeleton....
 of calcium carbonate
Calcium carbonate

Calcium carbonate is a chemical compound with the chemical formula CalciumCarbonOxygen3. It is a common substance found as Rock in all parts of the world, and is the main component of seashells, snails, and eggshells....
. The accumulation of this skeletal material, broken and piled up by wave action and bioeroders
Bioerosion

Bioerosion describes the erosion of hard Substrate s – and less often terrestrial substrates – by living organisms by a number of mechanisms....
, produces massive calcareous formations that make ideal habitats for living corals and a great variety of other animal and plant life.

ral reefs can take a variety of forms, defined in following:

BVI salts ponds were all shallow and mostly hypersaline and their waters were generally well mixed.






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Nwhi   French Frigate Shoals Reef   Many Fish
Coral reefs are aragonite
Aragonite

Aragonite is a carbonate mineral, one of the two common, naturally occurring polymorphism of calcium carbonate, calciumcarbonoxygen3....
 structures produced by living organisms. In most reefs the predominant organisms are colonial cnidarians that secrete an exoskeleton
Exoskeleton

An exoskeleton is an external skeleton that supports and protects an animal's body, in contrast to the internal endoskeleton of, for example, a human skeleton....
 of calcium carbonate
Calcium carbonate

Calcium carbonate is a chemical compound with the chemical formula CalciumCarbonOxygen3. It is a common substance found as Rock in all parts of the world, and is the main component of seashells, snails, and eggshells....
. The accumulation of this skeletal material, broken and piled up by wave action and bioeroders
Bioerosion

Bioerosion describes the erosion of hard Substrate s – and less often terrestrial substrates – by living organisms by a number of mechanisms....
, produces massive calcareous formations that make ideal habitats for living corals and a great variety of other animal and plant life.

Formations

Coral Reef Diagram
Coral reefs can take a variety of forms, defined in following:
  • Fringing reef
    Fringing reef

    A Fringing reef is a kind of coral reef, that is located in the tropics generally immediately near the shoreline. This type of coral reef is the most common type of reef that is found....
     – a reef that is directly attached to a shore or borders it with an intervening shallow channel or lagoon.
  • Barrier reef
    Barrier reef

    Barrier reef can refer to:*a coral reef,*the Great Barrier Reef in Australia,*the Belize Barrier Reef,*the New Caledonia Barrier Reef,*Barrier Reef , an Australian television series....
     – a reef separated from a mainland or island shore by a deep lagoon
    Lagoon

    A lagoon is a body of comparatively shallow sea water or brackish water separated from the deeper sea by a shallow or exposed Bar , reef, or similar feature....
     (see Great Barrier Reef
    Great Barrier Reef

    The Great Barrier Reef is the largest coral reef system in the world, composed of over 2,900 individual reefs and 900 islands stretching for 2,600 kilometres over an area of approximately ....
    ).
  • Patch reef – an isolated, often circular reef, usually within a lagoon
    Lagoon

    A lagoon is a body of comparatively shallow sea water or brackish water separated from the deeper sea by a shallow or exposed Bar , reef, or similar feature....
     or embayment.
  • Apron reef – a short reef resembling a fringing reef, but more sloped; extending out and downward from a point or peninsular shore.
  • Bank reef – a linear or semi-circular shaped-outline, larger than a patch reef.
  • Ribbon reef – a long, narrow, somewhat winding reef, usually associated with an atoll lagoon.
  • Atoll
    Atoll

    An atoll is an island of coral that encircles a lagoon partially or completely....
     reef
    – a more or less circular or continuous barrier reef extending all the way around a lagoon without a central island.
  • Table reef – an isolated reef, approaching an atoll type, but without a lagoon.


BVI salts ponds were all shallow and mostly hypersaline and their waters were generally well mixed. They exhibited large variations in salinity, both temporally and spatially among ponds. -http://www.biology-online.org/

Distribution

Coral Reef Locations
Coral reefs are estimated to cover 284,300 square kilometres, with the Indo-Pacific
Indo-Pacific

The Indo-Pacific is a biogeography region of the earth's seas, comprising the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the seas connecting the two in the general area of Indonesia....
 region (including the Red Sea
Red Sea

The Red Sea is a salt water inlet of the Indian Ocean between Africa and Asia. The connection to the ocean is in the south through the Bab el Mandeb sound and the Gulf of Aden....
, Indian Ocean
Indian Ocean

The Indian Ocean is the third largest of the world's oceanic divisions, covering about 20% of the water on the Earth's surface. It is bounded on the north by Asia ; on the west by Africa; on the east by Indochina, the Sunda Islands, and Australia; and on the south by the Southern Ocean ....
, Southeast Asia
Southeast Asia

Southeast Asia or Southeastern Asia is a subregion of Asia, consisting of the countries that are geographically south of China, east of India and north of Australia....
 and the Pacific) accounting for 91.9% of the total. Southeast Asia accounts for 32.3% of that figure, while the Pacific including Australia
Australia

Australia, officially the Commonwealth of Australia, is a country in the southern hemisphere comprising the Australia of the world's smallest continent, the major island of Tasmania, and numerous list of islands of Australia in the Indian Ocean and Pacific Oceans....
 accounts for 40.8%. Atlantic and Caribbean
Caribbean

The Caribbean is a region consisting of the Caribbean Sea, its islands , and the surrounding coasts. The region is located southeast of the Gulf of Mexico and Northern America, east of Central America, and to the north of South America....
 coral reefs only account for 7.6% of the world total.

Coral reefs are either restricted or absent from the west coast of the Americas
Americas

The Americas are the region of the Western hemisphere that consists of the continents of North America and South America with their associated islands and regions....
, as well as the west coast of Africa
Africa

Africa is the world's second-largest and second most-populous continent, after Asia. At about 30.2 million km? including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area....
. This is due primarily to upwelling
Upwelling

An Upwelling is an physical oceanography phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water towards the ocean surface, replacing the warmer, usually nutrient-depleted surface water....
 and strong cold coastal currents that reduce water temperatures in these areas. Corals are also restricted from off the coastline of South Asia
South Asia

South Asia, also known as Southern Asia, is the southern region of the Asian continent, which comprises the sub-Himalayan countries and, for some authorities , also includes the adjoining countries on the west and the east....
 from Pakistan
Pakistan

Pakistan , officially the Islamic Republic of Pakistan, is a country located in South Asia and borders Central Asia and the Middle East. It has a 1,046 kilometre coastline along the Arabian Sea and Gulf of Oman in the south, and is bordered by Afghanistan and Iran in the west, India in the east and People's Republic of China in th...
 to Bangladesh
Bangladesh

, officially the People's Republic of Bangladesh is a country in South Asia. It is bordered by India on all sides except for a small border with Burma to the far southeast and by the Bay of Bengal to the south....
. They are also restricted along the coast around north-eastern South America
South America

South America is the southern continent of the Americas, situated entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere....
 and Bangladesh
Bangladesh

, officially the People's Republic of Bangladesh is a country in South Asia. It is bordered by India on all sides except for a small border with Burma to the far southeast and by the Bay of Bengal to the south....
 due to the release of vast quantities of freshwater from the Amazon
Amazon River

The Amazon River of South America is the list of rivers by length in the world by volume, with a total river flow greater than the next top eight largest rivers combined....
 and Ganges Rivers respectively.

Although corals are found in temperate and tropical waters, shallow-water reefs are formed only in a zone extending at most from 30°N to 30°S of the equator. This zone is very important to whales because many types of plankton
Plankton

Plankton consist of any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. Plankton are defined by their ecological niche rather than their Phylogenetics or taxonomy classification....
 live there. Tropical corals do not grow at depths of over 50 m (165 ft). Temperature has less of an effect on the distribution of tropical coral, but it is generally accepted that they do not exist in waters below 18 °C
Celsius

Celsius is a temperature scale that is named after the Swedish astronomer Anders Celsius , who developed a similar temperature scale two years before his death....
., and that the optimum temperature is 26-27 °C for most coral reefs. The reefs in the Persian gulf however have coral adapted to changing temperatures of 13 °C in winter and 38 °C in summer, thus having significantly colder and hotter ambient environments respectively than most coral reefs. Also, deep water coral
Deep water coral

Deep Water Coral, also known as cold water coral, are found worldwide. They live in deeper, darker parts of the oceans than coral. Deep Water Corals belong to the Cnidaria and are most often Scleractinia....
 is more exceptional still as it can exist at greater depths and colder temperatures. Although deep water corals also form reefs, very little is known about them.

Famous coral reefs and reef areas of the world include:
  • The Great Barrier Reef
    Great Barrier Reef

    The Great Barrier Reef is the largest coral reef system in the world, composed of over 2,900 individual reefs and 900 islands stretching for 2,600 kilometres over an area of approximately ....
     - largest coral reef system in the world, Queensland, Australia
    Queensland

    Queensland is a States and territories of Australia of Australia, occupying the north-eastern section of the mainland continent. It is bordered by the Northern Territory to the west, South Australia to the south-west and New South Wales to the south....
    ;
  • The Belize Barrier Reef
    Belize Barrier Reef

    The Belize Barrier Reef is a series of coral reefs straddling the coast of Belize, roughly offshore in the north and in the south within the country limits....
     - second largest in the world, stretching from southern Quintana Roo, Mexico and all along the coast of Belize down to the Bay Islands of Honduras.
  • The New Caledonia Barrier Reef
    New Caledonia Barrier Reef

    The New Caledonia Barrier Reef is located in New Caledonia in the Oceania, and is the second-longest coral reef in the world, after Australia's Great Barrier Reef....
     - second longest double barrier reef in the world, with a length of about 1500km.
  • The Andros, Bahamas
    Andros, Bahamas

    Andros Island is the largest island of the Bahamas and the fifth largest island in the West Indies at roughly 2300 square miles in area and 104 miles long and 40 miles wide at its widest point....
     Barrier Reef - third largest in the world, following along the east coast of Andros Island, Bahamas between Andros and Nassau.
  • The Red Sea
    Red Sea

    The Red Sea is a salt water inlet of the Indian Ocean between Africa and Asia. The connection to the ocean is in the south through the Bab el Mandeb sound and the Gulf of Aden....
     Coral Reef - located off the coast of Israel, Egypt and Saudi Arabia.
  • Pulley Ridge
    Pulley Ridge

    Pulley Ridge is a coral reef off the coast of southwestern Florida, United States . The reef lies 100 miles west of the Tortugas Ecological Reserve and stretches north about 60 miles....
     - deepest photosynthetic coral reef, Florida
    Florida

    Florida is a U.S. state located in the Southeastern United States of the United States, bordering Alabama to the northwest and Georgia to the northeast....
  • Many of the numerous reefs found scattered over the Maldives
    Maldives

    The Maldives , or Maldive Islands, officially the Republic of Maldives, is an island nation consisting of a Atolls of the Maldivess stretching south of India's Lakshadweep islands between Minicoy Island and the Chagos Archipelago, and about seven hundred kilometres south-west of Sri Lanka in the Laccadive Sea of Indian Ocean....


Biology

Coral Polyp
The building blocks of coral reefs are the generation of reef-building, and other organisms that are composed of calcium carbonate
Calcium carbonate

Calcium carbonate is a chemical compound with the chemical formula CalciumCarbonOxygen3. It is a common substance found as Rock in all parts of the world, and is the main component of seashells, snails, and eggshells....
. For example, as a coral head grows, it lays down a skeletal structure encasing each new polyp
Polyp

In zoology, a polyp is one of two forms of individuals found in many species of cnidarians. The two are the polyp or hydroid and the medusa . Polyps are approximately cylindrical, elongated on the axis of the body....
. Waves, grazing fish (such as parrotfish
Parrotfish

Parrotfish are mostly tropical, perciform marine fish of the family Scaridae. Abundant on shallow reefs of the Red Sea, Atlantic Ocean, Indian Ocean and Pacific Oceans, the parrotfish family contains ten genera and about 90 species....
), sea urchin
Sea urchin

Sea urchins are small, spiny, globular creatures that compose most of class Echinoidea. They are found in oceans all over the world. Their shell, or "test", is round and spiny, typically from 3 to 10 cm across....
s, sponges
Sea sponge

The sponges or poriferans are animals of the phylum Porifera . Their bodies consist of an outer thin layer of cells, the pinacoderm and an inner mass of cells and skeletal elements, the choanoderm....
, and other forces and organisms break down the coral skeletons into fragments that settle into spaces in the reef structure. Many other organisms living in the reef community contribute their skeletal calcium carbonate
Calcium carbonate

Calcium carbonate is a chemical compound with the chemical formula CalciumCarbonOxygen3. It is a common substance found as Rock in all parts of the world, and is the main component of seashells, snails, and eggshells....
 in the same manner. Coralline algae
Coralline algae

Coralline algae are red algae in the Family Corallinaceae of the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls....
 are important contributors to the structure of the reef in those parts of the reef subjected to the greatest forces by waves (such as the reef front facing the open ocean). These algae contribute to reef-building by depositing limestone in sheets over the surface of the reef and thereby contributing also to the structural integrity of the reef.

Reef-building or hermatypic corals are only found in the photic zone
Photic zone

The photic zone or euphotic zone is the depth of the water in a lake or ocean, that is exposed to sufficient sunlight for photosynthesis to occur....
 (above 50 m depth), the depth to which sufficient sunlight penetrates the water for photosynthesis
Photosynthesis

File:Seawifs global biosphere.jpgPhotosynthesis is a metabolic pathway that converts carbon dioxide into organic compounds, especially sugars, using the energy from sunlight....
 to occur. The coral polyps do not photosynthesize, but have a symbiotic relationship with single-celled algae called zooxanthellae; these algal cells within the tissues of the coral polyps carry out photosynthesis and produce excess organic nutrients that are then used by the coral polyps. Because of this relationship, coral reefs grow much faster in clear water, which admits more sunlight. Indeed, the relationship is responsible for coral reefs in the sense that without their symbionts, coral growth would be too slow for the corals to form impressive reef structures. Corals can get up to 90% of their nutrients from their zooxanthellae symbionts.

Corals can reproduce both sexually and asexually. An individual polyp may use both reproductive modes within its lifetime. Corals reproduce sexually by either internal or external fertilization. The reproductive cells are found on the mesentery membranes that radiate inward from the layer of tissue that lines the stomach cavity. Some mature adult corals are hermaphroditic; others are exclusively male or female. A few even change sex as they grow.

Internally fertilized eggs are brooded in the polyp for a period ranging from days to weeks. Subsequent development produces a tiny larva, known as a planula. Externally fertilized eggs develop during a synchronized spawning. Polyps release eggs and sperm into the water simultaneously. This spawning method disperses eggs over a larger area. Synchronous spawning depends on four factors: time of year, water temperature, and tidal and lunar cycles. Spawning is most successful when there is little variation between high and low tides. The less water movement there is over the reef, the better the chance that an egg will be fertilized. Ideal timing occurs in the spring, release of eggs or planula larvae usually occurs at night and is sometimes in phase with the lunar cycle (3-6 days after a full moon). The period from release to settlement lasts only a few days, but some planulae can survive afloat for several weeks (7, 14). They are vulnerable at this time to heavy predation and adverse environmental conditions. For the lucky few which survive to attach to substrate, the challenge comes from competition for food and space.

Ecology and biodiversity

Blue Linckia Starfish
Elephant Ear Sponge
Coral reefs support an extraordinary biodiversity
Biodiversity

Biodiversity is the variation of life forms within a given ecosystem, biome, or for the entire Earth. Biodiversity is often used as a measure of the health of biological systems....
; although they are located in nutrient-poor tropical waters. The process of nutrient cycling
Biogeochemical cycle

In ecology and Earth science, a biogeochemical cycle or nutrient cycle is a pathway by which a chemical element or molecule moves through both biotic and abiotic compartments of Earth....
 between corals, zooxanthellae, and other reef organisms provides an explanation for why coral reefs flourish in these waters: recycling ensures that fewer nutrients are needed overall to support the community.

Cyanobacteria
Cyanobacteria

Cyanobacteria, also known as blue-green algae, blue-green bacteria or Cyanophyta, is a phylum of bacteria that obtain their energy through photosynthesis....
 also provide soluble nitrate
Nitrate

In inorganic chemistry, a nitrate is a salt of nitric acid with an ion composed of one nitrogen and three oxygen atoms . In organic chemistry the esters of nitric acid and various alcohols are called nitrates....
s for the coral reef through the process of nitrogen fixation
Nitrogen fixation

Nitrogen fixation is the process by which nitrogen is taken from its relatively inert molecular form in the Earth's atmosphere and converted into nitrogen compounds ....
. Corals absorb nutrients, including inorganic nitrogen and phosphorus
Phosphorus

Phosphorus is the chemical element that has the symbol P and atomic number 15. The name comes from the and . A Valency nonmetal of the nitrogen group, phosphorus is commonly found in inorganic phosphate minerals....
, directly from the water, and they feed upon zooplankton
Zooplankton

Zooplankton are the heterotrophic type of plankton. Plankton are organisms drifting in the Pelagic zone of oceans, seas, and bodies of fresh water....
 that are carried past the polyps by water motion. Thus, primary productivity on a coral reef is very high, which results in the highest biomass
Biomass (ecology)

Biomass, in ecology, is the mass of living biological organisms in a given area or ecosystem at a given time. Biomass can refer to species biomass, which is the mass of one or more species, or to community biomass, which is the mass of all species in the community....
 per square meter, at 5-10g C m-2 day-1. Producers in coral reef communities include the symbiotic zooxanthellae, sponges, marine worm
Marine worm

Any worm that lives in a ocean environment is considered a marine worm. Marine worms are found in several different phylum , including the Platyhelminthes, Nematoda, Annelida, Chaetognatha, Hemichordata, etc....
s, seaweed, coralline algae
Coralline algae

Coralline algae are red algae in the Family Corallinaceae of the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls....
 (especially small types called turf algae), ... although scientists disagree about the importance of these particular organisms.

Coral reefs often also depend on other habitats as seagrass meadows and mangrove forests in the surrounding area for the supply of nutrients. Seagrass and mangroves supply dead plants and animals, which are rich in nitrogen and also serve to feed fish and animals from the reef by supplying wood and vegetation to eat. Reefs in turn protect mangroves and seagrass from fierce waves and produce sediment
Sediment

Sediment is any particulate matter that can be sediment transport by fluid dynamics, and which eventually is deposited.Sediments are most often transported by water transported by wind and glaciers....
 for the mangroves and seagrass to root in.

Coral reefs are home to a variety of tropical or reef fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
 which can be distinguished. These include:
  • fish that swim right adjust the coral (such as Labridae and parrotfish
    Parrotfish

    Parrotfish are mostly tropical, perciform marine fish of the family Scaridae. Abundant on shallow reefs of the Red Sea, Atlantic Ocean, Indian Ocean and Pacific Oceans, the parrotfish family contains ten genera and about 90 species....
    ) These types of fish feed either on small animals living near the coral, seaweed, or on the coral itself. Fish that feeds on small animals include cleaner fish
    Cleaner fish

    Cleaner fish are fishes that provide a service to other fish species by removing dead skin and parasites. This is an example of mutualism, an ecological interaction that benefits both parties involved....
     (these fish eat them from between the jaws of larger predatory fish), bullet fish and Balistidae (these eat sea urchins) while fish eating seaweed include the Serranidae
    Serranidae

    Serranidae is a large family of fishes, belonging to the order Perciformes. The family contains about 450 species of serranids in 64 genera, including the bass and the groupers ....
    . Serranidae even tend to cultivate the weed by removing creatures feeding on it (as sea urchins), and they even remove unedible seaweeds. Fish that eats coral includes the parrotfish and butterflyfish
    Butterflyfish

    The butterflyfish are a group of conspicuous tropical marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group....
    .
  • fish that swim above and in the surrounding area of the coral reef. these include predatory fish as eg pompano
    Pompano

    Pompanos are marine fishes in the Trachinotus genus of the Carangidae family . Pompano may also refer to various other, similarly shaped members of Carangidae, or the Order Perciformes....
    s, grouper
    Grouper

    For other meanings, see Grouper .Groupers are fish of any of a number of genus in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes....
    s, Horse mackerel
    Horse mackerel

    mackerel is the popular name of various large fishes, such as the tuna, and the scad or saurel of the Pacific coast:*Australian bonito *various Jack mackerels...
    s, certain types of shark, Epinephelus marginatus
    Epinephelus marginatus

    The Dusky Grouper is the best known grouper of the Mediterranean Sea and North Africa coast. It is said to have the best taste of all Mediterranean fishes....
    , barracuda
    Barracuda

    The barracuda is a ray-finned fish known for its large size and fearsome appearance. Its body is long, fairly compressed, and covered with small, smooth scale ....
    s, snappers, ...) They also include herbivorous and plankton-eating fish. Fish eating seagrass include Horse mackerel, snapper, Pagellus, Conodon, ... Fish eating plankton includes Caesio, manta ray, chromis, Holocentridae, pterapogon kauderni, ...


Generally, fish that swim in coral reefs are just as colourful as the reef itself. Examples are the beautiful parrotfish
Parrotfish

Parrotfish are mostly tropical, perciform marine fish of the family Scaridae. Abundant on shallow reefs of the Red Sea, Atlantic Ocean, Indian Ocean and Pacific Oceans, the parrotfish family contains ten genera and about 90 species....
, angelfish, damselfish
Damselfish

Damselfish refers to members of the family Pomacentridae, except those of the two genera Amphiprion and Premnas. Other species within the family have common names that include the word 'damselfish', but in almost all cases this is qualified with an adjective or other descriptor....
, Pomacanthus paru, Clinidae and butterflyfish
Butterflyfish

The butterflyfish are a group of conspicuous tropical marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group....
. At night however, some change colour to a more less catchy colour. Also, it should be noted that besides colourful fish swapping their colour to that of the environment, other fish (eg predatory and certain herbivorous fish as Lampanyctodes hectoris, Holocentridae, pterapogon kauderni, ...) as well as aquatic animals (Comatulida, Crinoidea, Ophiuroidea, ...) emerge and become active and certain go to rest.

Other fish groups found on coral reefs include grouper
Grouper

For other meanings, see Grouper .Groupers are fish of any of a number of genus in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes....
s, grunts and wrasse
Wrasse

The wrasses are a family , Labridae, of marine fish, many of which are brightly colored. The family is large and diverse, with about 500 species in 60 genera....
s. Over 4,000 species of fish inhabit coral reefs. It has been suggested that the high number of fish species that inhabit coral reefs are able to coexist in such high numbers because any free living space is rapidly inhabited by the first planktonic fish larvae that occupy it. These fish then inhabit the space for the rest of their life. The species that inhabit the free space are random and have therefore been termed "a lottery for living space".

Reefs are also home to a large variety of other organisms, including sponges
Sea sponge

The sponges or poriferans are animals of the phylum Porifera . Their bodies consist of an outer thin layer of cells, the pinacoderm and an inner mass of cells and skeletal elements, the choanoderm....
, Cnidaria
Cnidaria

Cnidaria Cnidarians were for a long time grouped with Ctenophores in the phylum Coelenterata, but increasing awareness of their differences caused them to be placed in separate phyla....
ns (which includes some types of corals and jellyfish
Jellyfish

Jellyfish are free-swimming members of the phylum Cnidaria. They have several different morphologies that represent several different cnidarian classes including the Scyphozoa , Staurozoa , Cubozoa , and Hydrozoa ....
), worm
Worm

A worm is a common name given to a diverse group of invertebrate animals that have a long, soft body and no legs. There are hundreds of thousands of species of worms, 2,700 of these are earthworms....
s, crustacean
Crustacean

Crustaceans are a large group of arthropods, comprising almost 52,000 described species , and are usually treated as a subphylum . They include various familiar animals, such as crabs, lobsters, crayfish, shrimp, krill and barnacles....
s (including shrimp
Shrimp

Shrimp are swimming, Decapoda crustaceans classified in the infraorder Caridea, found widely around the world in both fresh water and seawater. Adult shrimp are Filter feeder benthic animals living close to the bottom....
, cleaner shrimp
Cleaner shrimp

Cleaner shrimp is a generic term for any natantia Decapoda crustacean that cleans other organisms of parasites. This is a widely-cited example of symbiosis: a relationship in which both parties benefit....
s, spiny lobster
Spiny lobster

Spiny lobsters, also known as langouste or rock lobsters are a family of about 45 species of Achelata crustaceans, in the Decapoda Reptantia....
s and crab
Crab

Crabs are Decapoda crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" , or where the reduced abdomen is entirely hidden under the thorax....
s), molluscs (including cephalopod
Cephalopod

The cephalopods are the mollusc class Cephalopoda characterized by bilateral symmetry, a prominent head, and a modification of the mollusk foot, a muscular hydrostat, into the form of cephalopod arms or tentacles....
s), echinoderm
Echinoderm

Echinoderms are a Phylum of Marine animals . Echinoderms are found at every ocean depth, from the intertidal zone to the abyssal zone.Aside from the problematic Arkarua, the first definitive members of the phylum appeared near the start of the Cambrian period....
s (including starfish, sea urchins and sea cucumber
Sea cucumber

Holothuroidea is a class of marine animals with an elongated body and leathery skin, which is found on the sea floor worldwide. Many holothurian species and genera, informally known as sea cucumbers, are targeted for human consumption....
s), sea squirts, turtles such as the sea turtle
Sea turtle

Sea turtles are turtles found in all the world's oceans except the Arctic Ocean. There are seven living species of sea turtles: Flatback Sea Turtle, Green Sea Turtle, Hawksbill turtle, Kemp's Ridley, leatherback sea turtle, Loggerhead Sea Turtle and Olive Ridley Sea Turtle....
, green turtle and hawksbill turtle and sea snake
Sea snake

Sea snakes, or seasnakes, are venomous snake Elapidae snakes that inhabit marine environments for most or all of their lives. Though they evolved from terrestrial ancestors, most are extensively adapted to a fully aquatic life and are unable to even move on land, except for the genus Laticauda, which retain ancestral characteristics...
s. Aside from humans, mammal
Mammal

Mammals are a class of vertebrate animals whose name is derived from their distinctive feature, mammary glands, with which they feed their young....
s are rare on coral reefs, with visiting cetaceans such as dolphin
Dolphin

File:Bottlenose_Dolphin_KSC04pd0178.jpgDolphins are marine mammals that are closely related to whales and porpoises. There are almost forty species of dolphin in seventeen genus....
s being the main group. A few of these varied species feed directly on corals, while others graze on algae on the reef and participate in complex food webs.

These other organisms also have their part in the food-chain of the reef. Sea urchins for example eat seaweed, while the Hawksbill turtle
Hawksbill turtle

The hawksbill turtle is a critically endangered sea turtle belonging to the family Cheloniidae. It is the only species in its genus. The species has a worldwide distribution, with Atlantic Ocean and Pacific Ocean subspecies....
 eats sponges. Nudibranchia eat sponges too, as well as sea anemones. Dotidae and sea slugs eat seaweed.

A number of invertebrates, collectively called cryptofauna, inhabit the coral skeletal substrate itself, either boring into the skeletons (through the process of bioerosion
Bioerosion

Bioerosion describes the erosion of hard Substrate s – and less often terrestrial substrates – by living organisms by a number of mechanisms....
) or living in pre-existing voids and crevices. Those animals boring into the rock include sponges, bivalve molluscs, and sipuncula
Sipuncula

The Sipuncula or Sipunculida, sipunculid worms or peanut worms, are a Phylum containing 144-320 species of bilateral symmetry, segmentation sea worms....
ns. Those settling on the reef include many other species, particularly crustaceans and polychaete
Polychaete

The Polychaeta or polychaetes are a class of annelid worms, generally marine. Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin....
 worms.

Researchers have found evidence of algae dominance in locations of healthy coral reefs. In surveys done around largely uninhabited US Pacific islands, algae consists of a large percentage of the surveyed coral locations. The algae population consists of turf algae, coralline algae
Coralline algae

Coralline algae are red algae in the Family Corallinaceae of the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls....
, and macroalgae.

Threats

Coral Reef Bioerosion
Human activity may represent the greatest threat to coral reefs living in Earth's oceans. In particular, global warming
Global warming

Global warming is the increase in the Instrumental temperature record of the Earth's near-surface air and the oceans since the mid-twentieth century and its projected continuation....
, coral mining
Mining

Mining is the extraction of value minerals or other geology materials from the earth, usually from an ore body, vein or seam. Materials recovered by mining include base metals, precious metals, iron, uranium, coal, diamonds, limestone, oil shale, Sodium chloride and potash....
, pollution
Pollution

Pollution is the introduction of contaminants into an environment that causes instability, disorder, harm or discomfort to the ecosystem i.e. physical systems or living organisms ....
 (organic and non-organic/chemical), over-fishing, blast fishing
Blast fishing

Blast fishing or dynamite fishing is the practice of using explosives to stun or kill schools of fish for easy collection. This often illegal practice can be extremely destructive to the surrounding ecosystem, as the explosion often destroys the underlying habitat that supports the fish....
 and the digging of canal
Canal

Canals are artificial channels for water. There are two types of canals: Aqueduct canals, which are used for the conveyance and delivery of water, and waterways, which are navigable transportation canals used for passage of goods and people, often connected to existing lakes, rivers, or oceans....
s and access ways into islands and bays are the most serious threats to these ecosystems. Runoff from agricultural areas may also threaten reefs by encouraging the growth of harmful algae
Algae

Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms. The largest and most complex marine forms are called seaweeds....
.

In order to find answers for these problems, researchers are currently working to determine the degree various factors impact the reef systems. The list of factors is long but includes the oceans acting as a carbon dioxide sink
Carbon dioxide sink

A carbon sink is a natural or manmade reservoir that accumulates and stores some carbon-containing chemical compound for an indefinite period....
, changes in Earth's atmosphere
Earth's atmosphere

The Earth's atmosphere is a layer of gases surrounding the planet Earth that is retained by the Earth's gravity. Dry air contains roughly 78.08% nitrogen, 20.95% oxygen, 0.93% argon, 0.038% Carbon dioxide in the Earth's atmosphere, and trace amounts of other gases....
, ultraviolet light, ocean acidification
Ocean acidification

Ocean acidification is the name given to the ongoing decrease in the pH of the Earth's oceans, caused by their uptake of anthropogenic carbon dioxide from the Earth's atmosphere....
, biological virus
Virus

A virus is a Optical microscope#Limitations of light microscopes infectious agent that is unable to grow or reproduce outside a host cell . Viruses infect all cellular life....
, impacts of dust storms carrying agents to far flung reef systems, various pollutants, impacts of algal bloom
Algal bloom

An algal bloom is a rapid increase in the population of algae in an aquatic system. Algal blooms may occur in freshwater as well as marine environments....
s and others. Reefs are threatened well beyond coastal areas and so the problem is broader than factors from land development and pollution though those are too causing considerable damage.

Overfishing

The live food fish trade
Live food fish trade

The live food fish trade is a global system that links fishing communities with markets, primarily in Hong Kong and mainland China. Many of the fish are captured on coral reefs in Southeast Asia or the Pacific Island nations....
 has been implicated as a driver of decline due to the use of cyanide
Cyanide

A cyanide is any chemical compound that contains the nitrile , which consists of a carbon atom chemical bond to a nitrogen atom. Inorganic cyanides are hydrogen cyanide salts in which cyanide is generally the anion CN-....
 and disaster for people living in the tropics. Hughes, et al, (2003), writes that "with increased human population
World population

The world population is the total number of living humans on Earth at a given time. As of March 2009, the world's population is estimated to be about 6.76 1,000,000,000 ....
 and improved storage and transport systems, the scale of human impacts on reefs has grown exponentially. For example, markets for fishes and other natural resource
Natural resource

Renewable resources Renewable resources are sometimes living resources,, which can restock themselves if used sustainably and not over- harvested....
s have become global, supplying demand for reef resources far removed from their tropical sources."

Overfishing (and particularly selective overfishing) also creates another problem. They promote the abundant growth of certain fish and organisms that can be damaging to the reef if they appear in great numbers. For example the fishing of bullet fish, Balistidae and other natural predators as lobsters promote the population growth of sea urchins. Also, acanthaster planci, Drupella, tapiro, Terpios, and Rhodactis have been known to destroy reefs when their population became too big.

Aquarium fish
The hobby of keeping saltwater aquaria
Aquaria

Aquaria may refer to:*The plural of Aquarium*Aquaria , a 2D sidescrolling computer game*Aquaria , a symphonic power metal band from Rio de Janeiro, Brazil...
 has experienced an increase in world popularity since the 1990s. Beyond sales of aquaria, air pumps, food, medications and other supplies, the primary product of the aquarium industry is fish. However, the world market is limited in the diversity of collected species. For example, among 4000 coral reef fish species, only 200–300 are exploited. Selection of species results from a demand for fish being highly colorful and being able to be maintained and fed in aquaria. The last point is very important in the choice of imported species.

Although a few fish species (e.g. Pomacentridae) can be reproduced in aquaria, 95% of exploited fish are directly collected in the coral environment. Intense sampling of coral reef fish, especially in South-East Asia (including Indonesia and the Philippines), has caused great damage to the environment. A major catalyst of cyanide fishing
Cyanide fishing

Cyanide fishing is an illegal form of fishing common in South East Asia, which usually uses the chemical compound sodium cyanide. Since 2000, increasing restrictions on illegal dynamite fishing have led to an increasing growth in this indiscriminate method ? particularly as it can be used without generating noise....
 is poverty within fishing communities. In areas like the Philippines where cyanide is regularly used to catch live aquarium fish, the percentage of the population below the poverty line is 40%. In such developing countries
Developing country

A developing country is a country that has often low standards of democracy, industrialisation, Social work, and Human rights for its citizens....
, a fisherman might resort to such unethical practices in order to prevent his or her family from starving.

Most, 80–90%, of aquarium fish exported from the Philippines are captured with sodium cyanide
Sodium cyanide

Sodium cyanide is the inorganic compound with the chemical formula NaCN. This highly toxic colourless salt is used mainly in gold mining but has other niche applications....
.. This toxic chemical is dissolved in sea water and released into fish shelters. It has a rapid narcotic effect on fish, which are then easily captured. However, most fish collected with cyanide die a few months after capture from extensive liver damage. Moreover, other fish species that are not interesting for the aquarium market also die in the field.

Pollution

Coral reefs are biological assemblages adapted to waters with low nutrient content, and the addition of nutrients (called eutrofication) favors species (as algae, seaweed, ...) that disrupt the balance of the reef communities. Some algae are toxic, and both plants reduce the levels of sunlight and oxygen, killing of other marine organisms as fish and coral. Especially the addition of nutrients such as phosphate
Phosphate

A phosphate, an inorganic chemical, is a Salt of phosphoric acid. Inorganic phosphates are mining to obtain phosphorus for use in agriculture and industry....
s and nitrate
Nitrate

In inorganic chemistry, a nitrate is a salt of nitric acid with an ion composed of one nitrogen and three oxygen atoms . In organic chemistry the esters of nitric acid and various alcohols are called nitrates....
s are very damaging to reefs. High nitrate levels are toxic to corals, while phosphates slow down the growth of coral skeleton.

Poor water quality has also been shown to encourage the spread of infectious disease
Infectious disease

An infectious disease is a clinically evident disease resulting from the presence of pathogenic microbial agents, including pathogenic viruses, pathogenic bacteria, Mycosis, protozoa, multicellular parasites, and aberrant proteins known as prions....
s among corals.

Organic pollutants

Soil runoff
Extensive and poorly managed land development can threaten the survival of coral reefs. Runoff caused by farming and construction of roads, buildings, ports, channels, and harbors, can carry soil laden with carbon, nitrogen, phosphorus, and minerals. This nutrient-rich water can cause fleshy algae and phytoplankton to thrive in coastal areas, known as algal blooms, which have the potential to create hypoxic
Hypoxia (environmental)

Hypoxia or oxygen depletion is a phenomenon that occurs in aquatic environments as oxygen becomes reduced in concentration to a point detrimental to aquatic organisms living in the system....
 conditions by using all available oxygen.

Windborne
In addition to local soil runoff, additional soil (sand) is blown in from other regions. Dust from the Sahara
Sahara

The Sahara is the world's largest hot desert. At over 9,000,000 square kilometers , it covers most of Northern Africa, making it almost as large as the United States or the continent of Europe....
 moving around the southern periphery of the subtropical ridge
Subtropical ridge

The subtropical ridge is a large belt of High pressure area situated around the latitudes of 30th parallel north in the Northern Hemisphere and 30th parallel south in the Southern Hemisphere....
 moves into the Caribbean
Caribbean

The Caribbean is a region consisting of the Caribbean Sea, its islands , and the surrounding coasts. The region is located southeast of the Gulf of Mexico and Northern America, east of Central America, and to the north of South America....
 and Florida
Florida

Florida is a U.S. state located in the Southeastern United States of the United States, bordering Alabama to the northwest and Georgia to the northeast....
 during the warm season as the ridge builds and moves northward through the subtropical Atlantic. Dust can also be attributed to a global transport from the Gobi and Taklamakan
Taklamakan

For the novelette by Bruce Sterling see Taklamakan .The Taklamakan Desert , also known as Taklimakan, is a desert in Central Asia, in the Xinjiang Uyghur people Autonomous Region of the People's Republic of China....
 deserts across Korea
Korea

Korea is a geographic area composed of two sovereign countries, a civilization, and a former state situated on the Korean Peninsula in East Asia....
, Japan
Japan

Japan is an island country in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, People's Republic of China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south....
, and the Northern Pacific to the Hawaiian Islands
Hawaiian Islands

The Hawaiian Islands are an archipelago of 19 islands and atolls, numerous smaller islets, and undersea seamounts in the North Pacific Ocean, extending some 1,500 miles from the Hawaii in the south to northernmost Kure Atoll....
. Since 1970, dust outbreaks have worsened due to periods of drought in Africa. There is a large variability in the dust transport to the Caribbean and Florida from year to year; however, the flux of dust is greater during positive phases of the North Atlantic Oscillation
North Atlantic oscillation

The North Atlantic oscillation is a climate phenomenon in the North Atlantic Ocean of fluctuations in the difference of atmospheric Atmospheric_pressure#Mean_sea_level_pressure between the Icelandic Low and the Azores high....
. Dust events have been linked to a decline in the health of coral reefs across the Caribbean and Florida, primarily since the 1970s. Studies have shown that corals can incorporate dust into their skeletons as identified from dust from the 1883 eruption of Krakatoa
Krakatoa

Krakatoa , also spelled Krakatao, is a Island#Oceanic islands in the Sunda Strait between the islands of Java and Sumatra in Indonesia. The name is used for the island group, the main island , and the volcano as a whole....
 in Indonesia
Indonesia

The Republic of Indonesia , is a transcontinental country in Southeast Asia and Oceania. Comprising Islands of Indonesia, it is the world's largest Archipelago state....
 in the annular bands of the reef-building coral Montastraea annularis
Montastraea annularis

Montastraea annularis, commonly known as the boulder star coral is a species of coral that lives in the western Atlantic ocean and is the most thoroughly studied and most abundant species of reef-building coral in the Caribbean to date....
 from the Florida
Florida

Florida is a U.S. state located in the Southeastern United States of the United States, bordering Alabama to the northwest and Georgia to the northeast....
 reef tract. The relative abundance of chemical elements, particularly metals, has been used to distinguish soil derived from volcanic dust from mineral dust
Mineral dust

Mineral dust is a term used to indicate particulate originated from thesuspension of minerals constituting the soil, being composed of various oxides and carbonates....
.

Sewage
Another major pollutant is those generated by the people themselves. Most islanders use traditional sewage which often goes unfiltered into the sea. Filtering the sewage is something which is normally done in the first world, but in developing countries, this very important step is often skipped. Also, most experts now agree that composting toilet
Composting toilet

A composting toilet is a closed unit, not connected to a sewage system or septic tank, used to receive, contain, and Composting human waste via aerobic biodegradation....
 alongside a ecological sanitation
Ecological sanitation

Ecological sanitation, also known as ecosan or eco-san, is a new paradigm in sanitation that recognises human excreta and household wastewater not as waste but as resources that can and are recovered, treated , and reused....
 approach is best followed in small island nations, yet these countries already implemented such system and for the moment prefer to keep using it.

Mines
Copper, gold,and other mines inland where minerals are collected also form a major center of pollution. Most of the pollution is simply soil, which ends up in rivers flowing to the sea and ultimately covers the coral, but small mineral fractions may also introduce trouble. Copper
Copper

Copper is a chemical element with the symbol Cu and atomic number 29.It is a ductile metal with very high thermal and electrical conductivity....
, a common industrial pollutant, has been shown to interfere with the life history
Life history

The term life history has been given many meanings in several scientific fields. It can refer to a variety of methods and techniques that are used for conducting qualitative research interviews, especially in the fields of sociology and anthropology....
 and development of coral polyps.

Non-organic
Leaked oil and chemicals (e.g. from detergents, paints, ...) flowing into the sea from factory outlets are another key threat.

Chemical fertilizers (based on ammonium nitrate) are another pollutant.

Radioactive waste is often dumped by the USA near its military installations (Mororua, Fangataufa, Johnston Atoll, ... Also, nuclear tests (eg at Kwajalein, Bikini, Enewetak) may also caused some nuclear pollution, yet compared to the other forms of pollution noted, they remain only small.

Global warming

Global warming introduces sea level rise, effectively asking the coral to grow faster to keep up. Also, the sea temperature increases, which is very disturbing to the coral. This was seen during the 1998 and 2004 El Niño weather phenomena, in which sea surface temperature
Sea surface temperature

Sea surface temperature is the water temperature close to the surface.In practical terms, the exact meaning of surface varies according to the measurement method used....
s rose well above normal, many tropical coral reefs were bleached
Coral bleaching

Coral bleaching is the loss of color of corals, due to stress-induced expulsion of symbiotic unicellular algae or due to the loss of pigmentation within the algae....
 or killed. Some recovery has been noted in more remote locations, but global warming
Global warming

Global warming is the increase in the Instrumental temperature record of the Earth's near-surface air and the oceans since the mid-twentieth century and its projected continuation....
 could negate some of this recovery in the future. High seas surface temperature (SSTs) coupled with high irradiance (light intensity), triggers the loss of zooxanthellae, a symbiotic algae, and its dinoflagellate pigmentation in corals causing coral bleaching. Zooxanthellae provide 95% of the energy to the coral host. Reefs can often recover from bleaching if they are healthy to begin with and water temperatures cool. However, recovery will not be possible if CO2 levels rise to 500 ppm because there will not be enough carbonate ions lefct in the oceans for corals to grow.Refer to Hoegh-Guldberg 1999 for more information.

Global warming is also the basis of a new emerging problem: increasing coral diseases. Due to global warming (which is the main cause of coral bleaching), corals have been weakened. In their weakened state, the coral is much more prone to disease. As such, coral diseases have been beginning to spread more rapidly. These include Black band disease
Black band disease

Black band disease is characterized by complete coral tissue degradation due to a pathogenic microbial consortium that appears as a dark red or black migrating microbial mat....
 and White band disease
White band disease

'White band disease' is characterized by complete coral tissue degradation of Caribbean acroporid corals. Two species are affected, Acropora palmata and A....
. With the projected 2°C temperature increase, it is likely that coral will not be able to adapt enough physiologically or genetically to keep up with climate change

Ocean acidification

A related problem to global warming is ocean acidification, which is caused by one and the same problem; namely increasing CO2 emissions.

The decreasing ocean surface pH is of increasing long-term concern for coral reefs. Increased atmospheric CO2 increases the amount of CO2 dissolved in the oceans. Carbon dioxide gas dissolved in the ocean reacts with water to form carbonic acid
Carbonic acid

Carbonic acid has the Molecular formula H2CO3. It is also a name sometimes given to solutions of carbon dioxide in water , which contain small amounts of H2CO3....
, resulting in ocean acidification
Ocean acidification

Ocean acidification is the name given to the ongoing decrease in the pH of the Earth's oceans, caused by their uptake of anthropogenic carbon dioxide from the Earth's atmosphere....
. Ocean surface pH is estimated to have decreased from approximately 8.25 to 8.14 since the beginning of the industrial era, and it is estimated that it will drop by a further 0.3 - 0.4 units by 2100 as the ocean absorbs more anthropogenic CO2. Under normal conditions, the conditions for calcium carbonate production are stable in surface waters since the carbonate ion is at supersaturating
Supersaturation

The term supersaturation refers to a solution that contains more of the dissolved material than could be dissolved by the solvent under normal circumstances....
 concentrations. However, as ocean pH falls, so does the concentration of this ion, and when carbonate becomes under-saturated, structures made of calcium carbonate are vulnerable to dissolution. Research has already found that corals experience reduced calcification or enhanced dissolution when exposed to elevated CO2.

Mangroves and seagrassbeds

Within the last 20 years, once prolific seagrass
Seagrass

Seagrasses are flowering plants from one of four plant families , which grow in marine , fully-saline water environments....
beds and mangrove forests, which absorb massive amounts of nutrients and sediment
Sediment

Sediment is any particulate matter that can be sediment transport by fluid dynamics, and which eventually is deposited.Sediments are most often transported by water transported by wind and glaciers....
 have been destroyed. Both the loss of wetlands, mangrove habitats and seagrassbeds are considered to be significant factors affecting water quality
Water quality

Water quality is the physical, chemical and biological characteristics of water. It is most frequently used by reference to a set of standards against which compliance can be assessed....
 on inshore reefs.

Coral mining

Coral mining is another threat, conducted by the villagers themselves, as well as large-scale companies. Mining is often done for construction purposes, and is of particular value as these rocks can be obtained up to 50% cheaper than other rocks (eg from quarries
Quarries

Quarries - The "Royal Quarries" ? not found in Scripture ? is the namegiven to the vast caverns stretching far underneath the northern hill, Bezetha, on which Jerusalem is built....
). The coral rocks are ground and mixed with other materials as cement in order to make concrete. Ancient coral reef material used for construction is known as "coral rag
Coral rag

Coral rag is a rubbly limestone composed of ancient coral reef material. The term also refers to the building blocks quarried from these strata which are an important local building material in areas such as the east African coast and Florida....
".

Other physical destruction

Dynamite fishing
Blast fishing

Blast fishing or dynamite fishing is the practice of using explosives to stun or kill schools of fish for easy collection. This often illegal practice can be extremely destructive to the surrounding ecosystem, as the explosion often destroys the underlying habitat that supports the fish....
 is extremely destructive method. Sticks of dynamite, grenades, or home-made explosives are activated and thrown in the water. This method of fishing not only kills the fish within the main blast area, but also takes the lives of many reef animals that are not edible or wanted. The blast also kills the corals in the area, eliminating the very structure of the reef, destroying the habitat for fish and other animals important for the maintenance of a healthy reef.

Other types of fishing like muro-ami and kayakas kills all fish in certain areas, causing havoc on the ecosystem of the reef.

Boats and ships require an access point into bays and islands to load/unload cargo/people. For this, often parts of the reef are chopped away to clear a path. Alhough this seems but minor destruction of the reef, it can have additional negative consequences. These are altered water circulation and altered tidal patterns, which cause a turnaround in the reefs supply of nutrients; hereby sometimes destroying a great part of the reef and its ecosystem.

Fishing boats and other large vessels cause a form of destruction in shallow waters known as grounding. Two types of damage can occur. Collision damage occurs when a coral reef is crushed and split by a vessel's hull into multiple fragments. Scarring occurs when boat propellers tear off the live coral and expose the skeleton. The physical damage can be noticed as striations in the reefs.

Mooring
Mooring

Mooring may refer to:* Mooring , any device used to hold secure an object by means of cables, anchors, or lines* Mooring spoken in Germany* Arthur Mooring, British Resident to the Sultan of Zanzibar...
 also causes considerable damage. To reduce the amount of (considerable) devastation which may occure, mooring buoys have been invented. They are already placed in most major wetparks and marine sanctuaries. Most buoys are of the Halas Mooring Buoy System-type.

Construction
Construction

In the fields of architecture and civil engineering, construction is a process that consists of the building or assembling of infrastructure. Far from being a single activity, large scale construction is a feat of multitasking....
 also takes its toll. Besides the previous discussed problem of using corals as construction material, the buildings themselves are often also placed right unto the reef, which causes even more destruction. This as it hinders the aquatic life and may alter water circulation and tides (which bring the nutrients for the reef). Buildings often placed include tourist resorts, airports, docks, and dwellings. The main reason why people build on top of reefs (despite possible problems of moist) is simply because of the lack of space.

Destruction worldwide

Southeast Asian coral reefs
Southeast Asia coral reefs

Southeast Asian coral reefs have the highest levels of biodiversity for the world's marine ecosystems. They serve many functions, such as forming the of livelihood for subsistence fishermen and even function as jewellery and construction materials....
 are at risk from damaging fishing
Fishing

Fishing is the activity of catching fish. Fishing techniques include Fish net, Fish trap, Spearfishing, angling and Gathering seafood by hand. The term fishing may be applied to catching other aquatic animals such as different types of shellfish, squid, octopus, turtles, Edible frog and some edible marine invertebrates....
 practices (such as cyanide
Cyanide fishing

Cyanide fishing is an illegal form of fishing common in South East Asia, which usually uses the chemical compound sodium cyanide. Since 2000, increasing restrictions on illegal dynamite fishing have led to an increasing growth in this indiscriminate method ? particularly as it can be used without generating noise....
 and blast fishing
Blast fishing

Blast fishing or dynamite fishing is the practice of using explosives to stun or kill schools of fish for easy collection. This often illegal practice can be extremely destructive to the surrounding ecosystem, as the explosion often destroys the underlying habitat that supports the fish....
), overfishing
Overfishing

Overfishing occurs when fishing activities reduce fish stocks below an acceptable level. This can occur in any body of water from a pond to the oceans....
, sedimentation, pollution and bleaching. A variety of activities, including education, regulation, and the establishment of marine protected areas are under way to protect these reefs. Indonesia
Indonesia

The Republic of Indonesia , is a transcontinental country in Southeast Asia and Oceania. Comprising Islands of Indonesia, it is the world's largest Archipelago state....
, for example has nearly of coral reefs. Its waters are home to a third of the world's total corals and a quarter of its fish species. Indonesia's coral reefs are located in the heart of the Coral Triangle
Coral Triangle

The Coral Triangle is a geographical term referring to the waters of Indonesia, Malaysia, Papua New Guinea, Philippines, Solomon Islands and Timor-Leste with an abundance of coral reef....
 and have been victim to destructive fishing, unregulated tourism, and bleaching due to climatic changes. Data from 414 reef monitoring stations throughout Indonesia in 2000 found that only 6% of Indonesia's coral reefs are in excellent condition, while 24% are in good condition, and approximately 70% are in poor to fair condition (2003 The Johns Hopkins University
Johns Hopkins University

The Johns Hopkins University, commonly referred to as Hopkins or JHU, is a private university research university located in Baltimore, Maryland, Maryland, United States....
).

On September 24, 2007, Reef Check
Reef Check

Reef Check is an international non-governmental organization dedicated to the conservation of two reef ecosystems: tropical coral reefs and Californian rocky reefs....
 (the world’s largest reef conservation
Habitat conservation

To conserve habitat areas for wild conservation reliant species and prevent their extinction or reduction in range is a priority of a great many groups that cannot be easily characterized in terms of any one ideology....
 organization
Organization

An organization is a social arrangement which pursues collective goals, which controls its own performance, and which has a boundary separating it from its environment....
) stated that only 5% of Philippines
Philippines

The Philippines, officially known as the Republic of the Philippines, is a country in Southeast Asia with Manila as its capital city. It comprises 7,107 islands in the western Pacific Ocean....
 27,000 square-kilometers of coral reef are in "excellent condition": Tubbataha Reef
Tubbataha Reef

Tubbataha Reef is an atoll coral reef located in the Sulu Sea of the Philippines. It is a marine sanctuary protected as Tubbataha Reef National Marine Park....
, Marine Park
Marine park

A marine park is a park consisting of an area of sea sometimes protected for recreational use, but more often set aside to preserve a specific habitat and ensure the ecosystem is sustained for the organisms that exist there....
 in Palawan
Palawan

Palawan is an island province of the Philippines located in the MIMAROPA Regions of the Philippines. Its capital is Puerto Princesa City, and it is the largest province in the country in terms of total area of jurisdiction....
, Apo Island
Apo Island

Apo Island is a small volcanic island, 7km off the southeastern tip of Negros Island in the Philippines. It is under the jurisdiction of the municipality of Dauin, Negros Oriental....
 in Negros Oriental
Negros Oriental

Negros Oriental is a Provinces of the Philippines of the Philippines located in the Central Visayas Regions of the Philippines. It occupies the south-eastern half of the island of Negros, with Negros Occidental comprising the north-western half....
, Apo Reef in Puerto Galera, Mindoro
Mindoro

Mindoro is the seventh-largest island in the Philippines. It is located southwest of Luzon, and northeast of Palawan. In past times, it has been called Ma-i or Mait by ancient China traders and, by Spain, as Mina de Oro from where the island got its current name....
, and Verde Island
Verde Island

This article refers to an island, for the body of water see Isla Verde Passage.Verde Island is located along the bodies of the Isla Verde Passage found between the provinces of Batangas and Oriental Mindoro, Luzon, Philippines....
 Passage off Batangas
Batangas

Batangas is a Provinces of the Philippines of the Philippines located on the southwestern part of Luzon in the CALABARZON Regions of the Philippines....
. Philippine coral reefs is second largest in Asia
Asia

Asia is the world's largest and most populous continent. It covers 8.6% of the Earth's total surface area and, with over 4 billion people, it contains more than 60% of the world's current human population....
.

General estimates show approximately 10% of the coral reefs around the world are already dead.Problems range from environmental effects of fishing
Environmental effects of fishing

File:Fishing down the food web.jpgThe environmental effects of fishing can be divided into issues that involve the availability of fish to be caught, such as overfishing, sustainable fisheries, and fisheries management; and issues that involve the impact of fishing on the environment, such as by-catch....
 techniques, described above, to ocean acidification
Ocean acidification

Ocean acidification is the name given to the ongoing decrease in the pH of the Earth's oceans, caused by their uptake of anthropogenic carbon dioxide from the Earth's atmosphere....
. Coral bleaching
Coral bleaching

Coral bleaching is the loss of color of corals, due to stress-induced expulsion of symbiotic unicellular algae or due to the loss of pigmentation within the algae....
 is another manifestation of the problem and is showing up in reefs across the planet.

Threatened species

The global standard for recording threatened
Threatened species

Threatened species are any species which are vulnerable to extinction in the near future.World Conservation Union is the foremost authority on threatened species, and treats threatened species not as a single category, but as a group of three categories: Vulnerable species, endangered species, and Critically endangered species, depending...
 marine species is the IUCN Red List of Threatened Species. This list is the foundation for marine conservation priorities worldwide. A species is listed in the threatened category if it is considered to be critically endangered
Critically endangered

---- Organisms with a conservation status of critically endangered have an extremely high risk of becoming extinct....
, endangered, or vulnerable
Vulnerable

Vulnerable may refer to:*Vulnerability*Vulnerable species* Vulnerable , by Tricky* Vulnerable * Vulnerable ...
. Other categories are near threatened
Near Threatened

Near Threatened is a conservation status assigned to species or lower taxa that may be considered threatened species with extinction in the near future, although it does not currently qualify for the threatened status....
 and data deficient
Data Deficient

Data Deficient is a category applied by the IUCN to a species when the available information is not sufficient for a proper assessment of conservation status to be made....
. By 2008, the IUCN had assessed all known reef-building corals species as follows

Group Species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
Threatened
Threatened species

Threatened species are any species which are vulnerable to extinction in the near future.World Conservation Union is the foremost authority on threatened species, and treats threatened species not as a single category, but as a group of three categories: Vulnerable species, endangered species, and Critically endangered species, depending...
Near threatened
Near Threatened

Near Threatened is a conservation status assigned to species or lower taxa that may be considered threatened species with extinction in the near future, although it does not currently qualify for the threatened status....
Data deficient
Data Deficient

Data Deficient is a category applied by the IUCN to a species when the available information is not sufficient for a proper assessment of conservation status to be made....
Reef-building corals
Coral

Corals are marine organisms from the class Anthozoa and exist as small sea anemone?like polyps, typically in colonies of many identical individuals....
845 27% 20% 17%


According to the IUCN, the primary threats to coral reefs are bleaching and disease which has been linked to an increase in sea temperatures. Other threats include coastal development, coral extraction, sedimentation and pollution. The coral triangle
Coral Triangle

The Coral Triangle is a geographical term referring to the waters of Indonesia, Malaysia, Papua New Guinea, Philippines, Solomon Islands and Timor-Leste with an abundance of coral reef....
 (Indo-Malay-Philippine archipelago) region has the highest number of reef-building coral species in threatened category as well as the highest coral species diversity. The loss of coral reef ecosystems will have devastating effects on many marine species, as well as on people that depend on reef resources for their livelihoods. Various chemicals are dropped into the sea this kills off the reefs.

Protection and restoration

Manusreefs L7 2000feb20
Inhabitants of Ahus Island, Manus Province
Manus Province

Manus Province is the smallest province in Papua New Guinea with a land area of 2,100 km?, but with more than 220,000 km? of water. The capital of the province is Lorengau and the total population is 43,387 ....
, Papua New Guinea
Papua New Guinea

Papua New Guinea , officially the Independent State of Papua New Guinea, is a country in Oceania, occupying the eastern half of the island of New Guinea and numerous offshore islands ....
, have followed a generations-old practice of restricting fishing in six areas of their reef lagoon. While line fishing is permitted, net and spear fishing are restricted based on cultural traditions. The result is that both the biomass
Biomass

Biomass, as a renewable energy source, refers to living and recently dead biological material that can be used as fuel or for industrial production....
 and individual fish sizes are significantly larger in these areas than in places where fishing is completely unrestricted. It is estimated that about 60% of the world's reefs are at risk due to destructive, human-related activities. The threat to the health of reefs is particularly strong in Southeast Asia
Southeast Asia

Southeast Asia or Southeastern Asia is a subregion of Asia, consisting of the countries that are geographically south of China, east of India and north of Australia....
, where an enormous 80% of reefs are considered endangered
Endangered species

An endangered species is a population of an organism which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters....
.

Protected areas
Many governments worldwide take measures to protect their coral reefs.

One method of coastal reef management that has become increasingly prominent is the implementation of Marine Protected Areas (MPAs)
Marine Protected Area

Marine Protected Area is a protected area whose boundaries include some area of ocean. MPA is often used as an umbrella term covering a wide range of marine areas with some level of restriction to protect living, non-living, cultural, and/or historic resources....
. MPAs have been introduced in Southeast Asia and elsewhere around the world to attempt to promote responsible fishery management
Fisheries management

Fisheries management is today often referred to as a governmental system of management rules based on defined objectives and a mix of management means to implement the rules, which is put in place by a system of Monitoring control and surveillance....
 and habitat protection
Habitat (ecology)

A habitat is an ecological or Natural_environment area that is inhabited by a particular animal or plant species. It is the natural environment in which an organism lives, or the physical environment that surrounds a species population....
. Much like the designation of national park
National park

A national park is a reserve of land, usually declared and owned by a national government, protected from most human development and pollution....
s and wild life refuges, potentially damaging extraction activities are prohibited. The objectives of MPAs are both social and biological, including restoration of coral reefs, aesthetic maintenance, increased and protected biodiversity, and economic benefits. Conflicts surrounding MPAs involve lack of participation, clashing views and perceptions of effectiveness, and funding.

Biosphere reserve
Biosphere reserve

A biosphere reserve is an international conservation designation given by UNESCO under its Programme on Man and the Biosphere . The World Network of Biosphere Reserves is the collection of all 531 biosphere Nature reserve in 105 countries ....
s are other protected areas that may protect reefs.

Also, Marine park
Marine park

A marine park is a park consisting of an area of sea sometimes protected for recreational use, but more often set aside to preserve a specific habitat and ensure the ecosystem is sustained for the organisms that exist there....
s, as well as world heritage sites can provide protection for coral reefs. World heritage designation is something that is not immediately thought of for the protection of coral reefs, yet also play a vital role. For example the Chagos archipelago
Chagos Archipelago

The Chagos Archipelago is a group of seven atolls comprising more than 60 individual tropical islands roughly in the centre of the Indian Ocean....
, Sian Ka'an
Sian Ka'an

Sian Ka'an is a non profit and non governmental organization Biosphere Reserve formed by a group of conservasionists in the state of Quintana Roo, Mexico whose mission is the coservation of the biodiversity....
, the Great Barrier Reef
Great Barrier Reef

The Great Barrier Reef is the largest coral reef system in the world, composed of over 2,900 individual reefs and 900 islands stretching for 2,600 kilometres over an area of approximately ....
, Henderson Island
Henderson Island

Henderson Island may refer to:* Henderson Island , in the southern Pacific Ocean* Henderson Island , in Antarctica* Henderson Island , in the Ohio River...
, the Galapagos islands, Belize's Barrier reef and Palau
Palau

Palau , officially the Republic of Palau , is an borderless country in the Pacific Ocean, some 500 miles east of the Philippines and 2,000 miles south of Tokyo....
 have been designated as protected by nominating it a world heritage site.

In Australia, the Great Barrier Reef is protected by the Great Barrier Reef Marine Park Authority, and is the subject of much legislation, including a Biodiversity Action Plan
Biodiversity Action Plan

This article is about a conservation biology topic. For other uses of BAP, see BAP .A 'Biodiversity Action Plan' is an internationally recognized program addressing threatened species and habitats and is designed to protect and restore biological systems....
.

Restoration technologies
Low voltage electrical currents applied through seawater crystallizes dissolved minerals onto steel structures. The resultant white carbonate (aragonite
Aragonite

Aragonite is a carbonate mineral, one of the two common, naturally occurring polymorphism of calcium carbonate, calciumcarbonoxygen3....
) is the same mineral that makes up natural coral reefs. Corals rapidly colonize and grow at faster than normal rates onto these coated structures. The change in the environment produced by electrical currents also accelerates formation and growth of both chemical limestone rock and the skeletons of corals and other shell-bearing organisms. Within the vicinity of the anode and cathode is a high pH environment which inhibits the growth of filamentous and fleshy algae, which compete with coral for space. This, and the increased growth rates cease when the mineral accretion process stops.

The effects of mineral accretion is, however, only temporary. During the process the settled corals have an increased growth rate, and size, and density, but after the process is complete the corallites are comparable to naturally growing corallites in growth rate and density, and are about the same size or slightly smaller.

In large restoration projects, depending on the type of coral, placement of propagated hermatype coral
Coral

Corals are marine organisms from the class Anthozoa and exist as small sea anemone?like polyps, typically in colonies of many identical individuals....
 coral unto substrate is often done through the use of metal pins, superglue or milliput
Milliput

Milliput is a UK-based brand of epoxy putty popular among Model building, and which is also useful in countless household and Building restoration applications....
 . A-hermatype coral
Coral

Corals are marine organisms from the class Anthozoa and exist as small sea anemone?like polyps, typically in colonies of many identical individuals....
 can also be placed unto substrate using needle and thread . Also, to restore large sections of broken coral reef, concrete has also been used. Finally, special structures as reef balls can be placed to provide corals an ground base to grow on.

Organizations
Organizations which currently undertake coral reef/atoll restoration projects using simple methods of plant propagation
Plant propagation

'Plant propagation' is the process of artificially or naturally distributing plants....
:
  • U.S. Coral Reef Task Force (CRTF)
  • National Coral Reef Institute (NCRI)
  • US Department of Commerce’s National Oceanic and Atmospheric Administration (NOAA): Coral Reef Conservation Program
  • National Center for Coral Reef Research (NCORE)
  • Reef Ball
  • Southeast Florida Coral Reef Initiative (SEFCRI)
  • WorldFishCenter: promotes sustainable mariculture
    Mariculture

    Mariculture is a specialized branch of aquaculture involving the cultivation of marine organisms for food and other products in the open ocean, an enclosed section of the ocean, or in tanks, ponds or Raceway which are filled with seawater....
     techniques to grow reef organisms as tridacnidae
    Tridacnidae

    The Tridacninae are a family of Bivalves colloquially known as giant clams. The family contains the biggest bivalve species, including Tridacna gigas, the giant clam....


Organizations which carry out research to better understand the workings of coral reefs:
  • Tropical Marine Research Unit


Organizations which promote interest, provide knowledge bases about coral reef survival, and promote activities to protect and restore coral reefs:
  • Australian Coral Reef Society
  • Chagos Conservation Trust
  • Conservation Society of Pohnpei
  • Coral Reef Care
  • Cousteau Society
  • Crusoe Reef Society
  • CEDAM International
    CEDAM International

    CEDAM International is an American non-profit group for divers, dedicated to marine research and preservation. CEDAM stands for Conservation, Education, Diving, Awareness and Marine research....
  • Earthwatch
    Earthwatch

    The Earthwatch Institute is an international non-profit organization that was founded in 1971 near Boston to promote action and understanding necessary to sustain the Natural environment....
  • Environmental Defense Fund
  • Environmental Solutions International
  • Friends of Saba Marine Park
  • Global Coral Reef Alliance
  • Great Barrier Reef Marine Park Authority
  • International Marinelife Alliance
    International Marinelife Alliance

    The International Marinelife Alliance is a non-profit organization for Marine conservation. It is based in Hawaii, United States....
  • International Society for Reef Studies
  • Intercoast Network
  • Kosrae Conservation and Safety Organization
  • Marine Conservation Group
  • Marine Conservation Society
    Marine Conservation Society

    The Marine Conservation Society is a United Kingdom-based Non-governmental organization dedicated to the protection of the marine environment and its wildlife....
  • Nature Conservancy
  • Ocean Voice International
  • PADI
    Padi

    Padi or PADI can refer to:* Professional Association of Diving Instructors, diver training organization* Paddy field * Padi, Chennai, satellite township of Chennai, India...
  • Practical Action
    Practical Action

    Practical Action is a developing country charitable trust registered in the United Kingdom which works directly in four regions of the developing world ? Latin America, East Africa, Southern Africa and South Asia, with particular concentration on Peru, Kenya, Sudan, Zimbabwe, Sri Lanka and Nepal....
  • Project Reefkeeper
  • Reef Check
    Reef Check

    Reef Check is an international non-governmental organization dedicated to the conservation of two reef ecosystems: tropical coral reefs and Californian rocky reefs....
  • Reef Relief
  • Reefwatch
  • Seacology
    Seacology

    Seacology is a non-profit, non-governmental organization with the sole mission of preserving the ecosystems and cultures of islands throughout the globe....
  • Singapore Underwater Federation
  • Society for Andaman and Nicobar Ecology
    Society for Andaman and Nicobar Ecology

    The Society for Andaman and Nicobar Ecology, also known as SANE, is an Indian environmental organization based in Port Blair in the Andaman and Nicobar Islands union territory of India....
  • Tubbataha Foundation
  • Wildlife Conservation International
    Wildlife Conservation International

    Wildlife Conservation International is a field division of the Wildlife Conservation Society. It is based in Bronx, New York.The organization is mainly present in Africa and Asia as well as North America and South America....
  • WWF
    World Wide Fund for Nature

    The World Wide Fund for Nature is an Internationalism non-governmental organization for the Conservation biology, Environmental science and Restoration ecology of the environment , formerly named the World Wildlife Fund, which remains its official name in the United States and Canada....
There are separate layers of the ocean otherwise called as “zones”. The first zone is the Erysipelas Zone , this is where most of the visible light exists. Second is the Mesosphere Zone the light that penetrates to this depth is extremely faint. Then the Bathysphere Zone, here the only visible light is that produced by the creatures themselves. Fourth, the Abyssinia Zone, the water temperature is near freezing, and there is no light at all, very few creatures can be found at these very low depths. Lastly, the Hadalpelagic Zone, these areas are mostly found in deep water trenches and canyons.

Reefs in the past

Throughout the Earth
Earth

Earth is the third planet from the Sun. Earth is the largest of the terrestrial planets in the Solar System in diameter, mass and density. It is also referred to as the World and Wiktionary:Terra.Note that by International Astronomical Union convention, the term "Terra" is used for naming extensive land masses, rather...
 history, from a few thousand years after hard skeletons were developed by marine organisms, there were almost always reefs formed by reef-building organisms in the ancient seas. The times of maximum development were in the Middle Cambrian (513-501 Ma
Annum

Annum is one form of the Latin noun meaning year, not a form normally used for derivatives in modern languages: the accusative case Grammatical number of the second declension grammatical gender noun annus , anni ....
), Devonian
Devonian

The Devonian is a geologic period of the Paleozoic era spanning from . It is named after Devon, England, where rocks from this period were first studied....
 (416-359 Ma) and Carboniferous
Carboniferous

The Carboniferous is a geologic period that extends from the end of the Devonian period, about 359.2 ? 2.5 annum , to the beginning of the Permian period, about 299.0 ? 0.8 Ma ...
 (359-299 Ma), due to Order Rugosa
Rugosa

The Rugosa, also called the Tetracoralla, are an extinct order of coral that were abundant in Middle Ordovician to Late Permian seas....
 extinct
Extinction

In biology and ecology, extinction is the death of every member of a species or group of taxon. The moment of extinction is generally considered to be the death of the last individual of that species ....
 corals, and Late Cretaceous
Late Cretaceous

Late Cretaceous refers to the second half of the Cretaceous Period , named after the famous white chalk cliffs of southern England, which date from this time....
 (100-65 Ma) and all Neogene
Neogene

The Neogene is a Geologic time scale#Terminology starting 23.03 ? 0.05 million years ago and lasting either until today or ending 2.588 million years ago with the beginning of the Quaternary....
 (23 Ma - present), due to Order Scleractinia
Scleractinia

Scleractinia, also called Stony corals, are exclusively marine animals; they are very similar to sea anemones but generate a hard skeleton....
 corals.

Not all reefs in the past were formed by corals: in the Early Cambrian (542-513 Ma) resulted from calcareous algae
Algae

Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms. The largest and most complex marine forms are called seaweeds....
 and archaeocyathids
Archaeocyatha

The Archaeocyatha or archaeocyathids were Sessility , reef-building Marine organisms of warm tropical and subtropical waters that lived during the early Cambrian period....
 (small animals with conical shape, probably related to sponges) and in the Late Cretaceous
Late Cretaceous

Late Cretaceous refers to the second half of the Cretaceous Period , named after the famous white chalk cliffs of southern England, which date from this time....
 (100 - 65 Ma), when there also existed reefs formed by a group of bivalves
Bivalvia

Bivalves are molluscs belonging to the class Bivalvia. They have two-part animal shells, and typically both valves are symmetry along the hinge line....
 called rudists
Rudists

Rudists are a group of bizarrely shaped Marine heterodont bivalves that arose during the Jurassic, and became so diverse during the Cretaceous that they were major reef-building organisms in the Tethys Ocean....
; one of the valves formed the main conical structure and the other, much smaller valve acted as a cap.

General references

External links

  • Non-profit effort to benefit coral reef conservation globally, starting in 2008, the International Year of the Reef
  • Nova Southeastern University
  • (GCRMN)
  • University of Miami
  • (CORAL)
  • : &