Caloptilia betulicola
Encyclopedia
Caloptilia betulicola is a moth
Moth
A moth is an insect closely related to the butterfly, both being of the order Lepidoptera. Moths form the majority of this order; there are thought to be 150,000 to 250,000 different species of moth , with thousands of species yet to be described...

 of the Gracillariidae
Gracillariidae
Gracillariidae is an important family of insects in the order Lepidoptera and the principal family of leaf miners that includes several economic, horticultural or recently invasive pest species such as the horse-chestnut leaf miner, Cameraria ohridella....

 family. It is found from Scandinavia
Scandinavia
Scandinavia is a cultural, historical and ethno-linguistic region in northern Europe that includes the three kingdoms of Denmark, Norway and Sweden, characterized by their common ethno-cultural heritage and language. Modern Norway and Sweden proper are situated on the Scandinavian Peninsula,...

 and the north of European Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

 to the Pyrenees
Pyrenees
The Pyrenees is a range of mountains in southwest Europe that forms a natural border between France and Spain...

 and Alps
Alps
The Alps is one of the great mountain range systems of Europe, stretching from Austria and Slovenia in the east through Italy, Switzerland, Liechtenstein and Germany to France in the west....

 and from Ireland
Ireland
Ireland is an island to the northwest of continental Europe. It is the third-largest island in Europe and the twentieth-largest island on Earth...

 to Poland
Poland
Poland , officially the Republic of Poland , is a country in Central Europe bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian exclave, to the north...

 and Slovakia
Slovakia
The Slovak Republic is a landlocked state in Central Europe. It has a population of over five million and an area of about . Slovakia is bordered by the Czech Republic and Austria to the west, Poland to the north, Ukraine to the east and Hungary to the south...

. In the east it is found up to China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

, Japan
Japan
Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

 and the Russian Far East
Russian Far East
Russian Far East is a term that refers to the Russian part of the Far East, i.e., extreme east parts of Russia, between Lake Baikal in Eastern Siberia and the Pacific Ocean...

.

The wingspan
Wingspan
The wingspan of an airplane or a bird, is the distance from one wingtip to the other wingtip. For example, the Boeing 777 has a wingspan of about ; and a Wandering Albatross caught in 1965 had a wingspan of , the official record for a living bird.The term wingspan, more technically extent, is...

 is 14–16 mm. There are two generations per year, with adults on wing in June and July and again in September and October.

The larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e feed on Betula pendula and Betula pubecens. They mine
Leaf miner
Leaf miner is a term used to describe the larvae of many different species of insect which live in and eat the leaf tissue of plants. The vast majority of leaf-mining insects are moths , sawflies and flies , though some beetles and wasps also exhibit this behavior.Like Woodboring beetles, leaf...

 the leaves of their host plant. The mine starts with an inconspicuous epidermal corridor. Later, a blotch is formed, that quickly develops into a tentiform mine. Generally, the mine is lower-surface, but upper-surface mines are not rare. The frass
Frass
Frass is the fine powdery material phytophagous insects pass as waste after digesting plant parts. It causes plants to excrete chitinase due to high chitin levels, it is a natural bloom stimulant, and has high nutrient levels. Frass is known to have abundant amoeba, beneficial bacteria, and fungi...

is deposited in a mass of grains in a corner of the mine. After leaving the mine, the larva moves twice. First it lives in a rolled or folded leaf margin and later in a leaf that is rolled downwards, starting from the leaf tip. Pupation takes place in a white, shining, parchment-like cocoon, that is attached to the leaf margin with silk.
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