Agulla (snakefly)
Encyclopedia
Agulla is a genus of modern snakeflies
Snakefly
Snakeflies are a group of insects in the order Raphidioptera, consisting of about 210 extant species. Together with the Megaloptera they were formerly placed within the Neuroptera, but now these two are generally regarded as separate orders....

 in the family Raphidiidae.

Agulla species are predatory, both as adults and larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e. In North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 occur exclusively in the Western United States
Western United States
.The Western United States, commonly referred to as the American West or simply "the West," traditionally refers to the region comprising the westernmost states of the United States. Because the U.S. expanded westward after its founding, the meaning of the West has evolved over time...

, namely in the Rocky Mountains
Rocky Mountains
The Rocky Mountains are a major mountain range in western North America. The Rocky Mountains stretch more than from the northernmost part of British Columbia, in western Canada, to New Mexico, in the southwestern United States...

 and westward, including the southwestern desert
Desert
A desert is a landscape or region that receives an extremely low amount of precipitation, less than enough to support growth of most plants. Most deserts have an average annual precipitation of less than...

s.

Systematics

At present 31 living and two extinct species of Agulla are known from North America. The species are divided into four subgenera, with the two extinct (†) species left unplaced in the genus.
  • Agulla
    • Subgenus Agulla
      • A. (A.) arazonia (Banks)
      • A. (A.) amaudi (Aspöck)
      • A. (A.) assimilis (Albarda)
      • A. (A.) astuta (Banks)
      • A. (A.) barri (Aspöck)
      • A. (A.) bicolor (Albarda)
      • A. (A.) bractea Carpenter
      • A. (A.) crotchi (Banks)
      • A. (A.) faulkneri Aspöck
      • A. (A.) flexa Carpenter
      • A. (A.) herbsti (Ebsen-Peterson
    • Subgenus Galavia
      • A. (G.) adnixa (Hagen)
      • A. (G.) modesta Carpenter
      • A. (G.) paramerica Aspöck
      • A. (G.) unicolor Carpenter
    • Subgenus Franciscoraphidia
      • A. (F.) directa Carpenter
    • Subgenus Californoraphidia
      • A. (C.) nigrinotum Woglum & McGregor
    • Subgenus incertae sedis
      • A. mineralensis
        Agulla mineralensis
        Agulla mineralensis is an extinct species of snakefly in the raphidiid genus Agulla. The species is solely known from the Middle Miocene, late Bartovian stage, Pacific Union Site in the Stewart Valley Group, Mineral County, Nevada....

        (Miocene
        Miocene
        The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene...

        , Stewart Valley, Nevada, USA)
      • A. protomaculata
        Agulla protomaculata
        Agulla protomaculata is an extinct species of snakefly in the raphidiid genus Agulla. The species is solely known from the Middle Eocene, Lutetian stage, Parachute Member, part of the Green River Formation, in the Piceance Creek Basin and Uinta Basin, Garfield County, northwestern Colorado,...

        (Lutetian
        Lutetian
        The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between and . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene subepoch...

        , Green River Formation
        Green River Formation
        The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes. The sediments are deposited in very fine layers, a dark layer during the growing season and a light-hue inorganic layer in winter. Each pair of layers is called a varve and...

        , Colorado, USA)
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