Tooth-billed Hummingbird
Encyclopedia
The Tooth-billed Hummingbird (Androdon aequatorialis) is a species of bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

 from the family Trochilidae. It is monotypic
Monotypic
In biology, a monotypic taxon is a taxonomic group with only one biological type. The term's usage differs slightly between botany and zoology. The term monotypic has a separate use in conservation biology, monotypic habitat, regarding species habitat conversion eliminating biodiversity and...

 within the genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 Androdon. It is found in humid forests in western Colombia
Colombia
Colombia, officially the Republic of Colombia , is a unitary constitutional republic comprising thirty-two departments. The country is located in northwestern South America, bordered to the east by Venezuela and Brazil; to the south by Ecuador and Peru; to the north by the Caribbean Sea; to the...

, north-western Ecuador
Ecuador
Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

 (south to Pichincha Province
Pichincha Province
Pichincha is a province of Ecuador located in the northern sierra region; its capital and largest city is Quito. It is bordered by Imbabura & Esmeraldas to the north, Cotopaxi & Santo Domingo de los Tsáchilas to the south, Napo & Sucumbíos to the east, and Esmeraldas & Santo Domingo de los...

), and far eastern Panama
Panama
Panama , officially the Republic of Panama , is the southernmost country of Central America. Situated on the isthmus connecting North and South America, it is bordered by Costa Rica to the northwest, Colombia to the southeast, the Caribbean Sea to the north and the Pacific Ocean to the south. The...

 (Darién Province
Darién Province
Darién is a province in eastern Panama. It is also the largest province in Panama. It is hot, humid, heavily forested, and sparsely populated, having 48,378 habitants...

). While generally scarce, localized and associated with the highly threatened humid sections of the Chocó, its range remains relatively large, and it is therefore considered to be of least concern
Least Concern
Least Concern is an IUCN category assigned to extant taxon or lower taxa which have been evaluated but do not qualify for any other category. As such they do not qualify as threatened, Near Threatened, or Conservation Dependent...

 by BirdLife International
BirdLife International
BirdLife International is a global Partnership of conservation organisations that strives to conserve birds, their habitats and global biodiversity, working with people towards sustainability in the use of natural resources...

.

Appearance

The Tooth-billed Hummingbird has a total length of c. 14 cm (5½ in), which includes the long, essentially straight bill
Beak
The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

 of c. 4 cm (1½ in). Its common name
Common name
A common name of a taxon or organism is a name in general use within a community; it is often contrasted with the scientific name for the same organism...

 refers to the small tooth-like serrations on the inner part of the distal half of the bill. It is shiny green above with a copper
Copper
Copper is a chemical element with the symbol Cu and atomic number 29. It is a ductile metal with very high thermal and electrical conductivity. Pure copper is soft and malleable; an exposed surface has a reddish-orange tarnish...

y crown. The underparts are whitish streaked dusky. The rounded tail is greyish with a broad black subterminal band and white tips. There is a conspicuous white band on the rump. In males, the bill has a small hook at the tip and more prominent tooth-like spikes; their plumage
Plumage
Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season. Within species there can also be a...

 color averages brighter overall than females. While the green upper parts of this species are iridescent, its plumage is duller than that of most hummingbirds. Within its range, it is essentially unmistakable by the combination of the long, almost straight bill and the dark-streaked whitish underparts.

Habitat

The Tooth-billed Hummingbird is restricted to humid primary forests and nearby second growth
Secondary forest
A secondary forest is a forest or woodland area which has re-grown after a major disturbance such as fire, insect infestation, timber harvest or windthrow, until a long enough period has passed so that the effects of the disturbance are no longer evident...

. It occurs up to an altitude of c. 1560 m (5200 ft). In Ecuador, it is most frequent between 400–800 m (1300–2600 ft), while it mainly occurs between 600–1560 m (2000–5200 ft) in Panama, and up to 1050 m (3400 ft) in Colombia.

Behavior

Rather inconspicuous and generally poorly known. It is a swift flier. It forages from low levels to sub-canopy
Canopy (forest)
In biology, the canopy is the aboveground portion of a plant community or crop, formed by plant crowns.For forests, canopy also refers to the upper layer or habitat zone, formed by mature tree crowns and including other biological organisms .Sometimes the term canopy is used to refer to the extent...

, where it feeds on nectar from flowers of plant families such Ericaceae
Ericaceae
The Ericaceae, commonly known as the heath or heather family, is a group of mostly calcifuge flowering plants. The family is large, with roughly 4000 species spread across 126 genera, making it the 14th most speciose family of flowering plants...

 and Gesneriaceae
Gesneriaceae
Gesneriaceae is a family of flowering plants consisting of ca. 150 genera and ca. 3,200 species in the Old World and New World tropics and subtropics, with a very small number extending to temperate areas. Many species have colorful and showy flowers and are cultivated as ornamental plants.Most...

. Its main feeding strategy involves trap-lining
Trap-lining
Trap-lining or trap line refers to a feeding strategy amongst certain families of birds, notably hummingbirds and woodpeckers. These birds visit a circuit of specific plants, trees, or other feeding sites, much as trappers check their lines of traps....

. The serrated bill is also well adapted for catching small spiders and insects, while the hooked tip of the males' bill is further suited for extracting prey from within rolled leaves and cavities. It has sometimes been seen rapidly gleaning
Gleaning (birds)
Gleaning is a term for a feeding strategy by birds in which they catch invertebrate prey, mainly arthropods, by plucking them from foliage or the ground, from crevices such as rock faces and under the eaves of houses, or even, as in the case of ticks and lice, from living animals. This behavior is...

 below large leaves, while flicking its tail.

Little is known about its breeding behavior, but individuals captured in Colombia in January–March have had enlarged gonads, indicating that they were in breeding condition. In Ecuador, it has been reported that males gather in leks in February–May, which is unusual among members of the subfamily Trochilinae
Trochilinae
Trochilinae is a subfamily of the hummingbird family . Members of the subfamily Trochilinae are sometimes called typical hummingbirds. They typically display iridescent plumage in metallic reds, oranges, greens and/or blues...

, but frequent among the members of Phaethornithinae. Its calls include a high-pitches tsit-tseé-tsu and penetrating cheet notes.

Taxonomy

The Tooth-billed Hummingbird's taxonomic position has been controversial, with some arguing that it should be placed in the subfamily Phaethornithinae, and others that it, together with the two lancebills
Doryfera
Doryfera is a genus of hummingbird in the Trochilidae family.It contains the following species:* Blue-fronted Lancebill * Green-fronted Lancebill...

, should be placed in their own small subfamily, Doryferinae. Today most authorities place it at the beginning of Trochilinae.

External links

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