Throat clamp
Encyclopedia
Throat clamp is method of subduing often seen in predatory felids and occasionally canids and hyaenids. It involves the predator grasping the throat of the prey and clamping tight so that the windpipe is either crushed or blocked. Cats use this to kill prey while dogs and hyenas use this to weaken the prey before eating it, generally alive. It's more often used than the muzzle clamp
Muzzle clamp
A method of killing that large predators, usually cats such as Panthera leo, the lion, Panthera pardus, the leopard, and Uncia uncia, the snow leopard, use...

 and is generally safer, though slower. It is usually and most effective when positioned as near to the mandible
Mandible
The mandible pronunciation or inferior maxillary bone forms the lower jaw and holds the lower teeth in place...

 as the carnivore can get. Between the larynx
Larynx
The larynx , commonly called the voice box, is an organ in the neck of amphibians, reptiles and mammals involved in breathing, sound production, and protecting the trachea against food aspiration. It manipulates pitch and volume...

 and the jaw, the windpipe is surrounded with less cartilage
Cartilage
Cartilage is a flexible connective tissue found in many areas in the bodies of humans and other animals, including the joints between bones, the rib cage, the ear, the nose, the elbow, the knee, the ankle, the bronchial tubes and the intervertebral discs...

 and is more malleable, while lower down, near the chest
Chest
The chest is a part of the anatomy of humans and various other animals. It is sometimes referred to as the thorax or the bosom.-Chest anatomy - Humans and other hominids:...

, the passage way would be increasingly harder to collapse, so the throat clamp is usually positioned high up on the animal's neck.

Usage

If the prey is on the ground and pinned, most predators position themselves behind the animal to use their body's weight to help control the movements of the prey and reach around to grab the throat, usually twisting the head around, aiding in both their grasp and the blocking of the windpipe.

If the prey is standing, a solitary predator can use a throat clamp usually only if the prey is small or in shock
Acute stress reaction
Acute stress reaction is a psychological condition arising in response to a terrifying or traumatic event...

. A strong, large animal can easily kick and injure the predator from this angle. A social predator can use the standing throat clamp much more easily because there can be other individuals on the back of the prey, stopping it from kicking, which could lead to collapse, or using a throat clamp periodically to tire the animal out. This latter method usually involves large, patient carnivores such as Canis lupus or the wolf and Lyacon pictus or the African wild dog
African Wild Dog
Lycaon pictus is a large canid found only in Africa, especially in savannas and lightly wooded areas. It is variously called the African wild dog, African hunting dog, Cape hunting dog, painted dog, painted wolf, painted hunting dog, spotted dog, or ornate wolf...

. They surround the prey in relatively large numbers, individuals taking turns to jump in and grab the prey by the flanks to try and pull it down, while the prey forcing them off grows exhausted, and in the meantime, more experienced individuals secure a throat clamp on the distracted animal and hold on as long as possible to the struggling prey. The temporary throat clamp can make the prey dizzy and uncoordinated and even force it to go into shock if the chase and attack wasn't enough, aiding in the predator's attempts to pull it to the ground. Agile predators such a lions have been observed to perch on the upper back and neck of large cape buffalo and wrap around to secure a partial throat clamp.

Anatomy

Strong canines are generally required for this move: canines that are too short won't penetrate deeply enough and struggling prey can tear free, while canines that are too long or weak can be broken in the struggle with large prey. This fact created a controversy that is still generally unresolved as to the killing tactics of the extinct carnivores machairodonts, or sabre-toothed cats. Strong jaws are often needed to compress the windpipe far enough.

Most often, the canines pierce the prey behind the windpipe so that it is not torn, but compressed between premolars of upper and lower jaws. It can be compared to the bit of a horse's bridal in placement. With the mouth completely enclosing the windpipe, it is, if the predator does not relax its grip, very difficult to break free from.

Alternative forms

Occasionally, and as suggested for the killing method of machairodonts, the canines actually pierce the windpipe and the surrounding blood vessels so that the tear in arteries and adjacent windpipe lead to a flow of blood into the windpipe and down into the lungs where the animal essentially drowns in its own blood. This method has not been used regularly by any modern carnivores and is, due to the bloody nature, not suitable for an area with high competition due to the smell attracting other predators. It can be argued that this not a true throat clamp, but another specialized type of killing.
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