Tetrameridae
Encyclopedia
Tetrameridae is a family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of spiruria
Spiruria
Subclass Spiruria comprises mostly parasitic secernentean nematodes. In an alternate classification, they are treated as suborder Spirurina, with the orders listed here being ranked as infraorders....

n nematode
Nematode
The nematodes or roundworms are the most diverse phylum of pseudocoelomates, and one of the most diverse of all animals. Nematode species are very difficult to distinguish; over 28,000 have been described, of which over 16,000 are parasitic. It has been estimated that the total number of nematode...

s. It is the smallest of the large genera making up the bulk of the superfamily
Taxonomic rank
In biological classification, rank is the level in a taxonomic hierarchy. Examples of taxonomic ranks are species, genus, family, and class. Each rank subsumes under it a number of less general categories...

 Habronematoidea
Habronematoidea
Habronematoidea is a superfamily of spirurian nematodes in the large order Spirurida. Like all nematodes, they have neither a circulatory nor a respiratory system....

. Like all nematodes, they have neither a circulatory
Circulatory system
The circulatory system is an organ system that passes nutrients , gases, hormones, blood cells, etc...

 nor a respiratory system
Respiratory system
The respiratory system is the anatomical system of an organism that introduces respiratory gases to the interior and performs gas exchange. In humans and other mammals, the anatomical features of the respiratory system include airways, lungs, and the respiratory muscles...

. They are parasites, chiefly of bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s and cetacea
Cetacea
The order Cetacea includes the marine mammals commonly known as whales, dolphins, and porpoises. Cetus is Latin and is used in biological names to mean "whale"; its original meaning, "large sea animal", was more general. It comes from Ancient Greek , meaning "whale" or "any huge fish or sea...

ns..

This family contains the largest known nematode: Placentonema gigantissima is several meters long and has been found in the placenta
Placenta
The placenta is an organ that connects the developing fetus to the uterine wall to allow nutrient uptake, waste elimination, and gas exchange via the mother's blood supply. "True" placentas are a defining characteristic of eutherian or "placental" mammals, but are also found in some snakes and...

 of the Sperm Whale
Sperm Whale
The sperm whale, Physeter macrocephalus, is a marine mammal species, order Cetacea, a toothed whale having the largest brain of any animal. The name comes from the milky-white waxy substance, spermaceti, found in the animal's head. The sperm whale is the only living member of genus Physeter...

 (Physeter catodon).

Systematics

The Tetrameridae number about half a dozen genera
Genera
Genera is a commercial operating system and development environment for Lisp machines developed by Symbolics. It is essentially a fork of an earlier operating system originating on the MIT AI Lab's Lisp machines which Symbolics had used in common with LMI and Texas Instruments...

 only, but some are rather speciose; the type genus
Type genus
In biological classification, a type genus is a representative genus, as with regard to a biological family. The term and concept is used much more often and much more formally in zoology than it is in botany, and the definition is dependent on the nomenclatural Code that applies:* In zoological...

 Tetrameres contains a lot of species even by nematode standards. While it might be overlumped
Lumpers and splitters
Lumping and splitting refers to a well-known problem in any discipline which has to place individual examples into rigorously defined categories. The lumper/splitter problem occurs when there is the need to create classifications and assign examples to them, for example schools of literature,...

, as presently delimited about half of the more than 100 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of Tetrameridae are placed in it.

Subfamily Crassicaudinae Yorke & Maplestone, 1926
  • Crassicauda Leiper & Atkinson, 1914
  • Placentonema Gubanov, 1951

Subfamily Geopetitiinae Chabaud, 1951
  • Geopetitia Chabaud, 1951 [Diagnostic; French, Todd, Zachary & Meehan 1992

Subfamily Tetramerinae Travassos, 1914
  • Microhadjelia Jogis, 1965
  • Microtetrameres Travassos, 1915
  • Tetrameres Creplin, 1846
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