Protobactrites
Encyclopedia
Protobactrites is an extinct nautiloid cephalapod belonging to the Orthoceratoidea
Orthoceratoidea
The Orthoceratoidea is a superorder , sometimes considered an infraclass or even a subclass, that comprizes nautiloid orders that have orthoconic to slightly cyrtoconic shells and central to subcentral siphuncles in which there may be internal deposits...

 that lived in what would be Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 and Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 during the Ordovician
Ordovician
The Ordovician is a geologic period and system, the second of six of the Paleozoic Era, and covers the time between 488.3±1.7 to 443.7±1.5 million years ago . It follows the Cambrian Period and is followed by the Silurian Period...

 and Silurian
Silurian
The Silurian is a geologic period and system that extends from the end of the Ordovician Period, about 443.7 ± 1.5 Mya , to the beginning of the Devonian Period, about 416.0 ± 2.8 Mya . As with other geologic periods, the rock beds that define the period's start and end are well identified, but the...

 from 466–421.3 mya, existing for approximately .

Taxonomy

Protobactrites, named by Hyatt in Zittel (1900), is an orthocerid
Orthocerida
Orthocerida is an order of extinct nautiloid cephalopods also known as the Michelinocerda that lived from the Early Ordovician possibly to the Late Triassic . A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous...

 of unknown familial affiliation where it was retained by Sepkoski (2002). In spite of the name, Protobactrites has no known relationship to the Bactritida
Bactritida
The Bactritida form a small order of more or less straight-shelled cephalopods that first appeared during the Emsian Stage of the Devonian Period and persisted until the Carnian Stage of the Triassic Period...


Morphology

Protobactrites is characterized by a long slender orthoconic
Orthocone
An orthocone is a usually long straight shell of a nautiloid cephalopod. During the 18th and 19th centuries, all shells of this type were named Orthoceras, but it is now known that many groups of nautiloids developed or retained this type of shell....

 or faintly curved longiconic shell with a circular or subcircular cross section, transverse sutures, long body chamber and oblique aperture. The siphuncle is eccentric; exact structure unknown. The surface has transverse and in some species longitudinal striae. Adult shells may be naturally truncated.

Nothing is known of the living animal which may have had eight or ten arms, and tentacles, like modern coleoids
Coleoidea
Subclass Coleoidea, or Dibranchiata, is the grouping of cephalopods containing all the primarily soft-bodied creatures. Unlike its sister group Nautiloidea, whose members have a rigid outer shell for protection, the coleoids have at most an internal bone or shell that is used for buoyancy or support...

 such as modern squid
Squid
Squid are cephalopods of the order Teuthida, which comprises around 300 species. Like all other cephalopods, squid have a distinct head, bilateral symmetry, a mantle, and arms. Squid, like cuttlefish, have eight arms arranged in pairs and two, usually longer, tentacles...

 and octopus
Octopus
The octopus is a cephalopod mollusc of the order Octopoda. Octopuses have two eyes and four pairs of arms, and like other cephalopods they are bilaterally symmetric. An octopus has a hard beak, with its mouth at the center point of the arms...

.
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