Palaeotis
Encyclopedia
Palaeotis is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of paleognath bird from the middle Eocene epoch of central Europe. One species is known, Paleotis weigelti. The holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 specimen is a fossil tarsometatarsus
Tarsometatarsus
The tarsometatarsus is a bone that is found in the lower leg of certain tetrapods, namely birds.It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsal and metatarsal bones...

 and phalanx
Phalanx bones
In anatomy, phalanx bones are those that form the fingers and toes. In primates such as humans and monkeys, the thumb and big toe have two phalanges, while the other fingers and toes consist of three. Phalanges are classified as long bones.The phalanges do not have individual names...

. Lambrect (1928) described it as an extinct bustard
Bustard
Bustards, including floricans and korhaans, are large terrestrial birds mainly associated with dry open country and steppes in the Old World...

 (genus Otis), and gave it its consequent name (Palaeotis means ancient bustard). After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that Paleotis is a palaeognath and assigned it to the same order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...

 as ostrich
Ostrich
The Ostrich is one or two species of large flightless birds native to Africa, the only living member of the genus Struthio. Some analyses indicate that the Somali Ostrich may be better considered a full species apart from the Common Ostrich, but most taxonomists consider it to be a...

es; the Struthioniformes.

In the 1930s a nearly complete fossil with catalog number GM 4362 was assigned to Palaeotis, probably by Lambrecht. Houde and Haubold found three additional specimens in the collection of the Geiseltalmuseum, Martin-Luther University, Halle/S., Germany. One of those three is the holotype specimen of Paleogrus geiseltalensis (=Ornithocnemus geiseltalensis, Lambrecht 1935). Houde and Haubold also requested permission to prepare a fossil cataloged as HLMD Me 7530 at the Hesseches Landesmuseum in Darmstadt, Germany. HLMD Me 7530 was collected from the famous Messel shales. When it was prepared, the two Ornithologists assigned it to Palaeotis as well.

Other scientists are less convinced that Palaeotis is a struthioniform, placing it instead as a more basal ratite
Ratite
A ratite is any of a diverse group of large, flightless birds of Gondwanan origin, most of them now extinct. Unlike other flightless birds, the ratites have no keel on their sternum—hence the name from the Latin ratis...

. It may be related to the mysterious Remiornis, a putative ratite known from the Eocene
Eocene
The Eocene Epoch, lasting from about 56 to 34 million years ago , is a major division of the geologic timescale and the second epoch of the Paleogene Period in the Cenozoic Era. The Eocene spans the time from the end of the Palaeocene Epoch to the beginning of the Oligocene Epoch. The start of the...

 of France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

.

If Palaeotis is, as Houde and Haubold suggested a basal or even ancestral ostrich, it would be the only ratite
Ratite
A ratite is any of a diverse group of large, flightless birds of Gondwanan origin, most of them now extinct. Unlike other flightless birds, the ratites have no keel on their sternum—hence the name from the Latin ratis...

 known from the Northern hemisphere at this early age, and this has important implications for the evolution of ratites. See Paleognathae
Paleognathae
The Palaeognathae or paleognaths are one of the two living superorders of birds. The other living superorder is Neognathae. Together these two clades form the subclass Neornithes....

.
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