Palaeospondylus
Encyclopedia
Palaeospondylus gunni is a mysterious, fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

-like fossil vertebrate
Vertebrate
Vertebrates are animals that are members of the subphylum Vertebrata . Vertebrates are the largest group of chordates, with currently about 58,000 species described. Vertebrates include the jawless fishes, bony fishes, sharks and rays, amphibians, reptiles, mammals, and birds...

. Its fossils are described from Achannaras slate quarry in Caithness, Scotland
Scotland
Scotland is a country that is part of the United Kingdom. Occupying the northern third of the island of Great Britain, it shares a border with England to the south and is bounded by the North Sea to the east, the Atlantic Ocean to the north and west, and the North Channel and Irish Sea to the...

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The fossil as preserved is carbonized, and indicates an eel
Eel
Eels are an order of fish, which consists of four suborders, 20 families, 111 genera and approximately 800 species. Most eels are predators...

-shaped animal of up to 6 centimetres (2.4 in) in length. The skull, which must have consisted of hardened cartilage, exhibits pairs of nasal and auditory capsules, with a gill-apparatus below its hinder part, but no indications of ordinary jaws.

The phylogeny of this bizarre fossil has puzzled scientists since its discovery in 1890, and many taxonomies have been suggested. In 2004, researchers proposed that Palaeospondylus was a larval lungfish. Previously it had been classified as a larval tetrapod
Tetrapod
Tetrapods are vertebrate animals having four limbs. Amphibians, reptiles, birds and mammals are all tetrapods; even snakes and other limbless reptiles and amphibians are tetrapods by descent. The earliest tetrapods evolved from the lobe-finned fishes in the Devonian...

, unarmored placoderm, an agnatha
Agnatha
Agnatha is a superclass of jawless fish in the phylum Chordata, subphylum Vertebrata. The group excludes all vertebrates with jaws, known as gnathostomes....

n, and a chimaera
Chimaera
Chimaeras are cartilaginous fish in the order Chimaeriformes, known informally as ghost sharks, ratfish , spookfish , or rabbitfishes...

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