The orange-footed pimpleback mussel or orangefoot pimpleback, scientific name Plethobasus cooperianus, is a rare species

Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of freshwater mussel in the family Unionidae

Unionidae

Unionidae is a family of freshwater mussels, the largest in the order Unionoida, the bivalve mollusks sometimes known as river mussels, naiads, or simply as unionids.The range of distribution for this family is world-wide...

, the river mussels. This aquatic

Aquatic animal

An aquatic animal is an animal, either vertebrate or invertebrate, which lives in water for most or all of its life. It may breathe air or extract its oxygen from that dissolved in water through specialised organs called gills, or directly through its skin. Natural environments and the animals that...

 bivalve mollusk is native to the Tennessee

Tennessee River

The Tennessee River is the largest tributary of the Ohio River. It is approximately 652 miles long and is located in the southeastern United States in the Tennessee Valley. The river was once popularly known as the Cherokee River, among other names...

, Cumberland

Cumberland River

The Cumberland River is a waterway in the Southern United States. It is long. It starts in Harlan County in far southeastern Kentucky between Pine and Cumberland mountains, flows through southern Kentucky, crosses into northern Tennessee, and then curves back up into western Kentucky before...

, and lower Ohio River

Ohio River

The Ohio River is the largest tributary, by volume, of the Mississippi River. At the confluence, the Ohio is even bigger than the Mississippi and, thus, is hydrologically the main stream of the whole river system, including the Allegheny River further upstream...

s in the United States, where its distribution has declined over 70%.

This rounded mussel grows up to 9.5 centimeters long by 7.8 wide by 4.6 high. It has bumps on the posterior part of the shell and the foot is orange in live specimens. The mussel mainly lives in deep, fast-flowing rivers, especially with gravel or sand on the bottoms. It buries itself in the substrate under up to 29 feet of water.

This species was once common in some areas. It is now restricted for several reasons. Rivers throughout much of its range have been impounded

Dam

A dam is a barrier that impounds water or underground streams. Dams generally serve the primary purpose of retaining water, while other structures such as floodgates or levees are used to manage or prevent water flow into specific land regions. Hydropower and pumped-storage hydroelectricity are...

. There has been an increase in silt

Silt

Silt is granular material of a size somewhere between sand and clay whose mineral origin is quartz and feldspar. Silt may occur as a soil or as suspended sediment in a surface water body...

, which the mussel cannot tolerate. Pollution

Pollution

Pollution is the introduction of contaminants into a natural environment that causes instability, disorder, harm or discomfort to the ecosystem i.e. physical systems or living organisms. Pollution can take the form of chemical substances or energy, such as noise, heat or light...

 of the waterways has led to the decline of this and other mussels. The zebra mussel (Dreissena polymorpha) is a widespread threat.