Keuppia
Encyclopedia
Keuppia is an extinct genus of octopus
Octopus
The octopus is a cephalopod mollusc of the order Octopoda. Octopuses have two eyes and four pairs of arms, and like other cephalopods they are bilaterally symmetric. An octopus has a hard beak, with its mouth at the center point of the arms...

. It consists of two species, Keuppia hyperbolaris and Keuppia levante, both of which lived approximately 95 million years ago. Both species were found in fossilized form, which is very uncommon for extinct octopuses, as the soft tissue of dead octopuses almost always disintegrates before it has a chance to fossilize. These fossils, along with those of the genus Styletoctopus
Styletoctopus
Styletoctopus is an extinct genus of octopus. The genus consists of the single species Styletoctopus annae, which lived approximately 95 million years ago. Very few octopus species appear in the fossil record, as octopuses consist of soft tissue that usually decomposes before it has time to...

, were found from the Cretaceous
Cretaceous
The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...

-age Hâqel and Hâdjoula localities in Lebanon
Lebanon
Lebanon , officially the Republic of LebanonRepublic of Lebanon is the most common term used by Lebanese government agencies. The term Lebanese Republic, a literal translation of the official Arabic and French names that is not used in today's world. Arabic is the most common language spoken among...

. The presence of a gladius vestige in this genus shows a transition from squid to octopus in which the inner shell has divided in two in early forms to eventually be reduced to lateralized stylets, as can be seen in Styletoctopus.
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